澳大利亚泥盆纪皱纹珊瑚生物地层及生物地理研究展望

通过对澳大利亚泥盆纪皱纹珊瑚属级分布的分析,认为泥盆纪珊瑚在该区经历了两大繁盛期,尤以Ｌｏｃｈｋｏｖｉａｎ－Ｅｍｓｉａｎ最盛,有大量新属产生,内板珊瑚和菲力甫珊瑚科分子要Ｐｒａｇｉａｎ期开始出现,绳珊瑚科的分子也在Ｅｍｓｉａｎ大量出现,这些分子继而通过洋流逐渐扩散到华南及老世界域其他省区。     

Devonian conodont biogeography: Quantitative analysis of provinciality

Cluster analysis of conodont faunas from each of 17 Lower‐lower Upper Devonian zones and subzones (data as reported by Klapper and Johnson, 1980) reveal changing patterns of provinciality.

Provinciality, expressed by a differentiation into western Laurussian and proto‐Tethyan biogeographic regions, is moderate in the lower Lochkovian but is low or absent in the upper Lochkovian‐lower Pragian. Provincialism returns in the Pragian and reaches its maximum development during the Emsian. Most Australian faunas are distinct from those of western Laurussia and the proto‐Tethys. Conodont faunas from suspect terranes of western North America display no unusual biogeographic affinities. Provincialism declines during the Eifelian and is only weakly developed in Givetian‐lower Frasnian faunas.

Changing global sea level during the Devonian may explain the development of Devonian conodont provinciality. As proposed by Klapper and Johnson (1980), low provinciality is associated with low stands of sea level. Endemic faunas develop in isolated epeiric seas during intermediate stages of sea level rise. High stands of sea level ultimately drown barriers to faunal exchange and prompt a return to low provinciality conditions.     

On the origin of bactritoids (Cephalopoda)

There is a high probability that bactritoids represent a paraphylum or polyphylum. The initial chambers or protoconchs of the Early-Middle DevonianBactrites Sandberger,Devonobactrites Shimansky, andLobobactrites Schindewolf are elongated spheres with a diameter of 0.3–1.0 mm. The initial chambers are larger in diameter than the slender, smooth shaft located adorally to the initial chamber. Similar apices occur in a number of Late Silurian sphaerorthoceridans with central siphuncles. Sphaerorthoceridans with a bactritoid-like apex and an eccentric siphuncle are known from the Early Devonian. The earliest questionableBactrites occurs in the Pragian (middle Early Devonian). By Emsian time bactritoids are common elements of cephalopod faunas.Bactrites-like orthocones of the Middle Ordovician and Late Silurian are homeomorphs with clearly different early growth stages. Thus, the time interval between the first appearance ofBactrites and the origin of ammonoids can be narrowed down to the Pragian to Early Emsian. The placement of the siphuncle in a ventral marginal position has been used as one of the critical morphologic features in defining the bactritoids. However, the displacement of the siphuncle from subcentral or eccentric positions toward the conch margin occurred at least three times during the Ordovician — Early Devonian evolution of the Orthocerida. Thus, there is a high probability that a marginal shift of the orthocerid siphuncle occurred in post-Emsian times, too.     

Considérations paléobiogéographiques sur les Streptelasmatina (Coelenterata,Rugosa) solitaires du Dévonien des monts Cantabriques (NW de l'Espagne)

The study of numerous fossil solitary forms ofStreptelasmatina (Coelenterata, Rugosa) and some ones belonging to different suborders from the Devonian of the Cantabrian Mountains (NW Spain), shows the importance of such organisms concerning the relations among the major faunal provinces during the Devonian time.The analysis of the Devonian Rugose coral fauna(from Emsian to Frasnian) lets to establish the paleobiogeographical relations of the Cantabrian genera. Particularly, during the Lower Devonian (Emsian) the relationships between the Eastern of North America and Spain would be closer than it was supposed until now.The evidence on the world distribution of thestudied forms suggest possible migration roads.     

Body size of orthoconic cephalopods from the late Silurian and Devonian of the Anti‐Atlas (Morocco)

Fluctuations in body size of orthoconic cephalopods are reported from late Silurian to Late Devonian sediments at several locations in the Tafilalt and in the Dra‐Valley (Anti‐Atlas, Morocco). The combination of measurements of diameters and apical angles allows the reconstruction of their total conch size (length and volume), which revealed a strongly right‐skewed size distribution with an average length of 278 mm, while the largest Devonian actinocerids exceeded 2 m. Within the examined groups (Actinocerida, Orthocerida, Pseudorthocerida), there is no uniform trend, but rather frequent fluctuations with maximum sizes in the late Lochkovian and early Emsian. Body size decreased in times of extinction events, while stable periods are mostly associated with a size increase. Additionally, conch size correlates well with gamma diversity and global δ¹³C values. Furthermore, the apical angle and septal diameter of orthocones appear to correlate, but only when their mean values are compared across beds.     

Integrated study of fauna and microflora from the Early Devonian (Pragian–Emsian) of northwestern Argentina

The Devonian System in northern Argentina has been broadly analysed, but details of its lithologies, biostratigraphy and fossil content have not been presented in a comprehensive study. We performed the first integrative analysis of the palynological and macrofossil content from the Pescado Formation at the Zenta Range, Argentina. We define a new species of cryptospore and extend the stratigraphic record of the ichnogenus Psammichnites isp. for South America. The stratigraphic ranges of the palynomorphs suggest a time span from the ?late Lochkovian to Pragian–earliest Emsian, but the co-occurrences of key invertebrates narrow the age of the beds to the late Pragian and early Emsian. Moreover, sedimentary analysis indicates a proximal shoreface–foreshore depocenter during this time range for the Zenta region. The contraction phase of the basin during the middle Pragian and Emsian is evidenced by the presence of sand bodies at the top of the column and the higher supply of terrigenous components. During this regression event, a low diversity Malvinokaffric Realm brachiopod assemblage occurs, with dominance of Australospirifer hawkinsi. The predominance of the latter species during this event is coeval with the first decline of the Malvinokaffric Realm in the neighbouring Paraná basin.     

Evolution of Devonian biodiversity of radiolarians with two porous spheres and one main spine

Features and major distinctions in the appearance and distribution of radiolarians with two porous spheres and one main spine in the Devonian basins are considered. Four solitary population waves and expansion scenario of radiolarians of this morphotype have been recognized in the Devonian. A new species, Radiobisphaera rozanovi sp. nov., from the Upper Emsian, Upper Eifelian, and Middle Frasnian of the southern Ural Mountains, from the Upper Eifelian of the Prague Basin (Barrandian), from the Middle-Upper Frasnian of the Rudnyi Altai, and from the Lower Famennian of the Timan-Pechora Basin and the Polar Ural Mountains is described.     

Tentaculitids in palynological preparations: New evidence from the Famennian (Late Devonian) of Southern Timan

This is the first report of organic tentaculitid remains (zoomorphs) in palynological preparations of Famennian Age (Southern Timan, borehole Sosnovka-1). Earlier two similar finds have been described from the Frasnian of Poland as similar to embryonic and juvenile forms of dacryoconarids belonging to the orders Nowakiida and Stylionida.     

Sublethal injuries in non-marine microconchid tubeworms from the Lower Carboniferous of West Virginia,USA

More than 10,000 non-marine microconchid tubeworms (order Tentaculita) from the Lower Carboniferous (Mauch Chunk Group) of West Virginian, USA, have been inspected with respect to the occurrence of specimens having repaired injuries. The inspection showed that only nine specimens bear distinct regenerations of their tubes which constitute only 0.08% of the total specimens. This value, although much lower than that obtained from other encrusting tentaculitoid tubeworms, is very similar to that calculated before for the thousands of Early Devonian planktonic tentaculitids. This clearly indicates that sample size (specimen number) is critical for obtaining the reliable data about the proportion of regenerated vs. unaffected specimens. The sublethal injuries present in these few individuals probably resulted from failed predation, most probably by grazing fishes. However, the repaired injuries only indicate that some individuals survived but do not provide any information about the predation intensity on the group in the given paleoenvironment. Microconchids encrusted a given substrate in large quantities, so grazing fishes may have easily removed the majority of individuals, leaving no trace of predation activity in the Early Carboniferous, non-marine paleoenvironments of what is now West Virginia.     

Life-cycles of some Devonian ammonoids

Whorl expansion rates of six representative ammonoid genera from late Emsian and Eifelian strata of Morocco were calculated for each whorl. The corresponding body chamber lengths and the orientations of the apertures were computed based on these values. The resulting body chamber length and orientation of the aperture graphs were compared with other conch features, ecology of Recent cephalopods, and sedimentological data of the host rocks in the Tafilalt (eastern Anti-Atlas, Morocco). A subdivision of the ontogeny of these ammonoids was achieved comprising the early and late embryonic periods, the juvenile period, the preadult, and the adult growth period. All growth periods are defined by specific changes in growth, conch morphology, and mode of life. According to this reconstruction, hatchlings were probably already capable of active movements. Differentiation in two main modes of life of the examined taxa occurred in the late juvenile or early preadult period. As preadult animals, most of the Mimagoniatitoidea and Agoniatitoidea became active swimmers (Nektonic), whereas the representatives of the Anarcestoidea were capable of slow movements only (Planktonic). As adults, most representatives of the three superfamilies had an approximately horizontally oriented aperture, allowing active swimming and possibly active choice of spawning sites. Additionally, the new ammonoid taxon Rherisites tuba gen. nov., sp. nov. from the late Emsian is introduced.     

Description of the early ontogenic part of the Tentaculitids, with implications for classification

Ontogenic development and classification of tentaculitids at high systematic levels are reevaluated in the light of new findings on shell structure and morphology of larval parts, and these features are here regarded as being of primary importance for taxonomy. Class Tentaculita Bouček, 1964 is subdivided into two subclasses, of which subclass Chionioconarida Farsan, 1994 is distinguished by a tubular larval process closed at the apex and covered with microrings. The process is differentiated into a prolarval, metalarval and epilarval part, of which the latter coincides with metamorphosis. Morphology of the larval parts suggests that metamorphosis proceeds in two different manners, giving rise to superorders within this subclass. Within superorder Trompetoconarida Farsan, 1994 a bilaterally symmetrical larval cone develops with an aperture oblique to the long axis of the conch; following metamorphosis the conch becomes radially symmetrical and the aperture perpendicular to the axis; secondary shell, septa and pseudopunctae develop in the adult phase, and the structure of the shell is lamellar. In contrast, within the second superorder, Lirioconarida Farsan, 1994 the epilarval tube develops into a larval bulb with no changes in symmetry and position of the aperture; secondary shell, septa and pseudopuncta are absent. The microstructure of the shell is lamellar in the larval part whereas in postlarval parts it is either sigmoidal or lamellar. The subclass Dacryoconarida Fisher, 1962 possesses a subspherical, tear- or drop-like embryonic chamber which may have a caudal process. The microstructure of the embryonic chamber is variable within this group, being lamellar in some taxa whereas in others, a single layer of shell is present. The postembryonic parts of the lamellar forms possess nacreous or sigmoidal structures.     

Late Emsian Rutoceratoidea (Nautiloidea) from the Prague Basin,Czech Republic: morphology,diversity and palaeoecology

Abstract: Nautiloids of the superfamily Rutoceratoidea from the late Emsian (late Early Devonian) of the Prague Basin (Czech Republic) are commented upon. Species recognized include the hercoceratids Hercoceras mirum, H.? transiens, Ptenoceras proximum, P. nudum, P. minusculum and Anomaloceras anomalum, as well as the rutoceratids Adelphoceras bohemicum, Homoadelphoceras devonicans, Pseudorutoceras bolli and Goldringia? devonicans. In addition, four new species are described: Parauloceras regulare sp. nov., Roussanoffoceras chlupaci sp. nov., Otomaroceras sp. nov. and Goldringia sp. nov. Morphology and distribution patterns of Pragian and late Emsian rutoceratoid faunas from the Prague Basin are compared. They show that an increased diversity was accompanied by a higher level of specialization of rutoceratoids, which manifested itself in low abundance, increased facies dependence and greater variation in shell size during the Early Devonian. The evolution of sculpture and a contracted aperture, both regarded as protective adaptive features, was also examined, but no adaptive trend towards more pronounced sculpture and constriction of the aperture was found to have occurred in the Early Devonian. A more distinctive sculpture was, however, observed in shallow‐water assemblages of P. proximum in comparison with deeper‐water faunules, and two additional cephalopod species were examined in order to obtain comparative data. The presence of distinct sculpture patterns in coeval shallow‐ and deeper‐water assemblages suggests limited migration between them and consequently reflects some degree of territoriality in Devonian nautiloids. New data on early shell development in P. proximum are presented. During the Chote? Event, rutoceratoid generic diversity dropped dramatically, one family became extinct and the Early Devonian diversification of the group came to an end. The recovery of nautiloid faunas was slower than that of other cephalopods and associated, unrelated invertebrates. The absence of change in abundance patterns between Pragian and late Emsian rutoceratoid faunas, i.e. prior to and subsequent to ammonoid radiation, suggests that the appearance and radiation of the latter group in the early Emsian did not affect the structure of nautiloid assemblages, i.e. these two clades did not occupy the same niches.     

桂东、桂东南晚泥盆世一个弗拉斯期放射虫动物群

本文系统描述了广西玉林、贺县等地晚泥盆世榴江组硅质岩中的放射虫化石4属14种,其中有1新属2新种。这个放射虫动物群的多数分子曾见于苏联南乌拉尔地区和澳大利亚的弗拉斯阶。     

Diversity and richness of the Devonian terrestrial plants in the Southeastern Mountainous Altay (Southern Siberia): Regional versus global patterns

Devonian deposits of the Southeastern Mountainous Altay, a large region in Southern Siberia, contain abundant remains of terrestrial plants. A semi-quantitative analysis of this regional palaeobotanical record reveals how both the diversity (number of taxa) and richness (number of taxa with an account of their abundances) of floristic assemblages changed during the Emsian–early Famennian time interval. A total of 60 species, representing 42 genera, are known from 8 regional assemblages. Changes in diversity of species and genera occurred simultaneously. The number of taxa was high in the early Emsian, declined in the late Emsian, rose again in the Eifelian–middle Givetian, dropped in the late Givetian, reached the highest values in the early Frasnian, and experienced the greatest decline taking place in the late Frasnian–early Famennian. The standing diversity (number of taxa crossing the time boundaries) was maximal in the Middle Devonian. The dynamics of terrestrial plant richness was similar to that of diversity with an exception of middle Givetian decline in richness despite growth of diversity. The floras were dominated by pteridophytes. Propteridophytes were less abundant, and pinophytes were very rare. Propteridophyte extinctions were high in the middle Givetian, concurrent with a brief decline in pteridophytes. Some regional and global patterns of floral dynamics were similar. However, propteridophyte decline was not abrupt globally. The main abiotic driving factor influencing species richness and diversity appears to have been regional shoreline shifts. It does not appear that climate changes was important for regional changes in phytodiversity, although both regional and global phytodiversity was at its high during the Middle Devonian cooling phase. A comparison of palaeobotanical data from the Southeastern Mountainous Altay and Kazakhstan suggests palaeogeographic proximity and proves an idea of Altay-Mongolian terrane wandered between Gondwana, Kazakhstan, and Siberia. The early Emsian and the early Frasnian, when floras of the study region were diverse and rich, are characterized by the very high degree of similarity. Thus, interregional floral exchanges would facilitate plant radiations.     

Devonian Vertebrates From Colombia

Vertebrate remains are reported from the Emsian–Eifelian Floresta Formation and the Late Devonian (?Frasnian) Cuche Formation of north‐eastern Colombia. The material from the Floresta Formation is associated with a marine invertebrate fauna and includes an arthrodire and probably a rhenanid. Several vertebrate‐bearing localities are recorded from the Cuche Formation; vertebrates occur with plant remains and lingulid fragments. They include an acanthodian (Cheiracanthoides? sp.), a chondrichthyan (Antarctilamna? sp.), placoderms (Bothriolepis sp., Asterolepis? sp. and an undetermined groenlandaspidid or primitive brachythoracid arthrodire), a stegotrachelid actinopterygianand three sarcopterygians (a cosmine‐covered form tentatively referred to an osteolepidid, the porolepiform Holoptychiussp., and the rhizodontid Strepsodus? sp.). This assemblage suggests a Late Frasnian age and is surprisingly similar to Late Devonian vertebrate assemblages found in similar facies of Europe and North America, notwithstanding the presence of the Gondwanan chondrichthyan Antarctilamna?. key words: Vertebrata, Devonian, Colombia, South America, biostratigraphy, palaeobiogeography.     

Diversity dynamics of Zosterophyllopsida

The Zosterophyllopsida were major contributors to the diversification of early land plants. We present the first detailed analysis of the diversity dynamics of these plants from an updated database of all currently recognized zosterophyllopsid species. A set of quantitative methods classically used in palaeodiversity studies was applied to two data sets. The first one, ‘Zosterophyllopsida sensu stricto’, corresponds to the clade identified by Hao & Xue (The Early Devonian Posongchong Flora of Yunnan. (2013), Science Press). In the second, called ‘Zosterophyllopsida sensu lato’, barinophytalean‐type plants and taxa for which zosterophyllopsid affinities are suspected are added. The number of localities is used to explore sampling bias. Results show that sampling effect is minimal for the Early Devonian. For this time interval, both data sets record consistent patterns of changes suggesting that, whatever their affinities, all taxa included in the Zosterophyllopsida sensu lato show similar evolutionary trends. The diversity dynamics of zosterophyllopsids are characterized by a radiation during the Lochkovian, maximal values in the Pragian and a decline starting in the Emsian. The proportion of zosterophyllalean taxa with terminal sporangia is high until the Late Lochkovian when gosslingialean taxa without terminal sporangia evolved. During the Middle and Late Devonian, when diversity patterns are strongly affected by sampling, zosterophyllopsid diversity is low and characterized by a high proportion of barinophytacean and gosslingialean taxa, the latter becoming extinct in the Early Frasnian.     

Soft‐tissue imprints in fossil and Recent cephalopod septa and septum formation

Several soft‐tissue imprints and attachment sites have been discovered on the inside of the shell wall and on the apertural side of the septum of various fossil and Recent ectocochleate cephalopods. In addition to the scars of the cephalic retractors, steinkerns of the body chambers of bactritoids and some ammonoids from the Moroccan and the German Emsian (Early Devonian) display various kinds of striations; some of these striations are restricted to the mural part of the septum, some start at the suture and terminate at the anterior limit of the annular elevation. Several of these features were also discovered in specimens of Mesozoic and Recent nautilids. These structures are here interpreted as imprints of muscle fibre bundles of the posterior and especially the septal mantle, blood vessels as well as the septal furrow. Most of these structures were not found in ammonoids younger than Middle Devonian. We suggest that newly formed, not yet mineralized (or only slightly), septa were more tightly stayed between the more numerous lobes and saddles in more strongly folded septa of more derived ammonoids and that the higher tension in these septa did not permit soft‐parts to leave imprints on the organic preseptum. It is conceivable that this permitted more derived ammonoids to replace the chamber liquid faster by gas and consequently, new chambers could be used earlier than in other ectocochleate cephalopods, perhaps this process began even prior to mineralization. This would have allowed faster growth rates in derived ammonoids.     

Teruelia diezii gen. et sp. nov.: an early polysporangiophyte from the Lower Devonian of the Iberian Peninsula

A new basal land plant, Teruelia diezii gen. et sp. nov., is described from the shallow‐water marine deposits of the Lower Devonian (Lochkovian–Pragian) Nogueras Formation of the Iberian Peninsula (north Gondwana palaeocontinent). Teruelia is preserved as a compression fossil and consists of isotomously branched, robust stems terminated in large, fusiform, twisted sporangia. This morphology suggests that Teruelia is very probably equivalent to Aglaophyton, a permineralized early polysporangiophyte known up to now only from the Lower Devonian (early Pragian to ?earliest Emsian) Rhynie Chert in Scotland (Laurussia palaeocontinent), which represents an early terrestrial hot‐spring ecosystem. Accepted phylogenies identify Aglaophyton as sister to vascular plants. Our phylogeny‐based results identify the Aglaophyton/Teruelia biological entity (i.e. Aglaophyton anatomical characters plus Teruelia external morphology) as the most direct vascular plant precursor. It shows that at least one Rhynie Chert type plant had a much wider distribution than previously known and suggests that Aglaophyton was not restricted to hydrothermal environments, unlike other Rhynie Chert plants.     

Palaeoecology and palaeophytogeography of the rhynie chert plants: evidence from integrated analysis of in situ and dispersed spores

The Rhynie cherts yield exceptionally preserved early land plants, and provide a unique insight into the nature of Lower Devonian vegetation. Hitherto they have been poorly age constrained, and the palaeoecology and palaeophytogeography of the flora are poorly understood. Well-preserved dispersed-spore assemblages have been recovered from a number of borehole cores through the stratigraphical sequence of the Rhynie outlier. They are all similar and belong with the polygonalis-emsiensis (PE) spore zone, indicating an Early Devonian age (Early (but not earliest) Pragian to earliest Emsian). Comparisons with PE spore-zone assemblages from elsewhere suggest that the flora of the Rhynie drainage basin was slightly impoverished, with certain plant taxa that occurred at other locations not represented. This probably reflects differences between the flora of an inland intermontaine basin (Rhynie) and that of the lowland flood-plains. In situ spores have been characterized for all seven Rhynie chert plants. Analysis of the distribution of these spore types in the Rhynie sequence, in addition to those of coeval deposits from elsewhere, enables interpretation of the palaeoecology and palaeophytogeography of the Rhynie chert plants. It is concluded that at least some of the plants were not highly specialized or adapted to the peculiar hot-springs environment in which they are preserved. Rather, they were components of a diverse and widespread flora, but were the only elements able to tolerate the inhospitable hot-springs environment (i.e. they were preadapted).