Variabilité de l’industrie lithique entre les structures de l’habitation no 1 de Mezhyrich,site du Paléolithique supérieur d’Ukraine

The investigation of the lithic assemblages from three archaeological structures of the Upper Palaeolithic Mezhyrich site, mammoth bone dwelling no. 1, nearby located pit no. 7 and workshop with a portion of cultural layer situated on the border of the pit, exhibits a sufficient diversity of the main indexes of flint assemblages. According to the latter, the main flint resources, cores and large pieces of flint, as well as blades and several tools (scrapers), were kept just in the dwelling. At the same time, the remains of “debitage” and the modification products of several tools (burin spalls) are more numerous in the pit and the workshop. The higher percentages of backed microliths, included the forms with diagnostic projectile impact fracture, and burins with several working edges are also observed in the assemblages of the last two archaeological structures.     

Understanding Fossil Phytolith Preservation: The Role of Partial Dissolution in Paleoecology and Archaeology

Opaline phytoliths are important microfossils used for paleoecological and archaeological reconstructions that are primarily based on relative ratios of specific morphotypes. Recent studies have shown that phytolith assemblages are prone to post-depositional alteration involving partial dissolution, however, the manner in which partial dissolution affects morphotype composition is poorly understood. Here we show that morphotype assemblages from four different plant species subjected to controlled partial dissolution are significantly different from the original assemblages, indicating that the stability of various morphotypes differs, mainly depending on their surface area to bulk ratios. This underlying mechanism produces distorted morphotype compositions in partially dissolved phytolith assemblages, bearing vast implications for morphotype-based paleoecological and archaeological interpretation. Together with analyses of phytolith assemblages from a variety of archaeological sites, our results establish criteria by which well-preserved phytolith assemblages can be selected for accurate paleoecological and archaeological reconstructions.     

First GIS Analysis of Modern Stone Tools Used by Wild Chimpanzees (Pan troglodytes verus) in Bossou,Guinea, West Africa

Stone tool use by wild chimpanzees of West Africa offers a unique opportunity to explore the evolutionary roots of technology during human evolution. However, detailed analyses of chimpanzee stone artifacts are still lacking, thus precluding a comparison with the earliest archaeological record. This paper presents the first systematic study of stone tools used by wild chimpanzees to crack open nuts in Bossou (Guinea-Conakry), and applies pioneering analytical techniques to such artifacts. Automatic morphometric GIS classification enabled to create maps of use wear over the stone tools (anvils, hammers, and hammers/ anvils), which were blind tested with GIS spatial analysis of damage patterns identified visually. Our analysis shows that chimpanzee stone tool use wear can be systematized and specific damage patterns discerned, allowing to discriminate between active and passive pounders in lithic assemblages. In summary, our results demonstrate the heuristic potential of combined suites of GIS techniques for the analysis of battered artifacts, and have enabled creating a referential framework of analysis in which wild chimpanzee battered tools can for the first time be directly compared to the early archaeological record.     

Quantifying Traces of Tool Use: A Novel Morphometric Analysis of Damage Patterns on Percussive Tools

Percussive technology continues to play an increasingly important role in understanding the evolution of tool use. Comparing the archaeological record with extractive foraging behaviors in nonhuman primates has focused on percussive implements as a key to investigating the origins of lithic technology. Despite this, archaeological approaches towards percussive tools have been obscured by a lack of standardized methodologies. Central to this issue have been the use of qualitative, non-diagnostic techniques to identify percussive tools from archaeological contexts. Here we describe a new morphometric method for distinguishing anthropogenically-generated damage patterns on percussive tools from naturally damaged river cobbles. We employ a geomatic approach through the use of three-dimensional scanning and geographical information systems software to statistically quantify the identification process in percussive technology research. This will strengthen current technological analyses of percussive tools in archaeological frameworks and open new avenues for translating behavioral inferences of early hominins from percussive damage patterns.     

Hominin Dispersal into the Nefud Desert and Middle Palaeolithic Settlement along the Jubbah Palaeolake,Northern Arabia

The Arabian Peninsula is a key region for understanding hominin dispersals and the effect of climate change on prehistoric demography, although little information on these topics is presently available owing to the poor preservation of archaeological sites in this desert environment. Here, we describe the discovery of three stratified and buried archaeological sites in the Nefud Desert, which includes the oldest dated occupation for the region. The stone tool assemblages are identified as a Middle Palaeolithic industry that includes Levallois manufacturing methods and the production of tools on flakes. Hominin occupations correspond with humid periods, particularly Marine Isotope Stages 7 and 5 of the Late Pleistocene. The Middle Palaeolithic occupations were situated along the Jubbah palaeolake-shores, in a grassland setting with some trees. Populations procured different raw materials across the lake region to manufacture stone tools, using the implements to process plants and animals. To reach the Jubbah palaeolake, Middle Palaeolithic populations travelled into the ameliorated Nefud Desert interior, possibly gaining access from multiple directions, either using routes from the north and west (the Levant and the Sinai), the north (the Mesopotamian plains and the Euphrates basin), or the east (the Persian Gulf). The Jubbah stone tool assemblages have their own suite of technological characters, but have types reminiscent of both African Middle Stone Age and Levantine Middle Palaeolithic industries. Comparative inter-regional analysis of core technology indicates morphological similarities with the Levantine Tabun C assemblage, associated with human fossils controversially identified as either Neanderthals or Homo sapiens.     

Experimental investigations into the preservation of pollen grains: A pilot study of four pollen types

Preservation condition of pollen grains from different sedimentary environments can be a useful tool for interpreting pollen assemblages, but current understanding of the controls on pollen preservation is limited. One constraint is the complexity of studying these processes in the natural system. This paper presents the results of a pilot study of laboratory assessments of damage processes. A standard mixture of dried pollen grains (4 taxa: Alnus, Secale, Dactylis and Ambrosia) was rehydrated and suspended in glycerine, then subjected to two types of treatment, chemical oxidation or agitation in water with added sand or pebbles. The treatments generated distinctive damage patterns, and the pattern of damage observed varies with duration of treatment, combinations of treatments and pollen taxon. The results suggest that laboratory methods, which can generate damaged pollen assemblages rapidly and under controlled conditions, are a useful route for investigating the processes controlling pollen degradation and therefore interpreting the damage spectra recorded from past assemblages in terms of environmental processes.     

The fragmented character of Middle Palaeolithic stone tool technology

The importance of the transport of stone artefacts in structuring Neandertal lithic assemblages has often been addressed, but the degree to which this led to fragmentation of lithic reduction over Middle Palaeolithic landscapes has not been explicitly studied thus far. Large-scale excavations of Middle Palaeolithic open-air sites and refitting studies of the retrieved assemblages have yielded new, high-resolution data on the mobile aspects of Neandertal stone tool technology. In this paper, we integrate lithic technology and raw material data from recent studies of Middle Palaeolithic open-air and rock shelter sites in Western Europe. We demonstrate that the results of a variety of typological, technological (especially refitting), and lithological studies have important consequences for our knowledge of the acquisition of raw materials and subsequent production, usage and discard of stone artefacts in the Middle Palaeolithic. Neandertal production and use of stone tools was fragmented in three domains: the spatial, the temporal and the social domain. We show that this versatile segmentation of stone artefact handling strategies is a main determinant of the character of the Neandertal archaeological record. Our data testify to ubiquitous and continuous transport of stone artefacts of a wide variety of forms, picked by Neandertals using selection criteria that were sometimes far removed from what archaeologists have traditionally considered, and to some degree still consider, to be desired end products of knapping activities. The data presented here testify to the variability and versatility of Middle Palaeolithic stone tool technology, whose fragmented character created very heterogeneous archaeological assemblages, usually the product of a wide variety of independent import, use, discard and/or subsequent transport events.     

Documenting Differences between Early Stone Age Flake Production Systems: An Experimental Model and Archaeological Verification

This study investigates morphological differences between flakes produced via “core and flake” technologies and those resulting from bifacial shaping strategies. We investigate systematic variation between two technological groups of flakes using experimentally produced assemblages, and then apply the experimental model to the Cutting 10 Mid -Pleistocene archaeological collection from Elandsfontein, South Africa. We argue that a specific set of independent variables—and their interactions—including external platform angle, platform depth, measures of thickness variance and flake curvature should distinguish between these two technological groups. The role of these variables in technological group separation was further investigated using the Generalized Linear Model as well as Linear Discriminant Analysis. The Discriminant model was used to classify archaeological flakes from the Cutting 10 locality in terms of their probability of association, within either experimentally developed technological group. The results indicate that the selected independent variables play a central role in separating core and flake from bifacial technologies. Thickness evenness and curvature had the greatest effect sizes in both the Generalized Linear and Discriminant models. Interestingly the interaction between thickness evenness and platform depth was significant and played an important role in influencing technological group membership. The identified interaction emphasizes the complexity in attempting to distinguish flake production strategies based on flake morphological attributes. The results of the discriminant function analysis demonstrate that the majority of flakes at the Cutting 10 locality were not associated with the production of the numerous Large Cutting Tools found at the site, which corresponds with previous suggestions regarding technological behaviors reflected in this assemblage.     

Testing the suitability of a morphological monitoring approach for identifying temporal variability in a temperate sponge assemblage

Sponges are a dominant component of benthic assemblages in hard substratum environments across the world, but despite the importance of sponges, they are generally poorly represented in most marine monitoring programmes. There is considerable need to develop effective monitoring tools to monitor changes in sponge assemblages. Morphological monitoring has been proposed as a suitable method to monitor sponges and while morphological monitoring has already taken place at Skomer Marine Reserve (MNR), Wales, here we investigate whether species level and morphological level data sets are correlated with respect to temporal variation. Furthermore, we examine the environmental factors that correlate with the patterns of temporal variability. Both species and morphological data sets revealed significant seasonal changes and spatial variation in sponge assemblages; these data sets were highly correlated and explained by a number of environmental factors. We conclude that morphological monitoring of sponge assemblages may represents a cost-effective method for assessing temporal and spatial variation in sponges, where full species level monitoring is not possible, as patterns identified from morphological data were a suitable surrogate of species-level data.     

Between simplicity and complexity: The knapping flexibility on cobbles at the early Neolithic site of Zengpiyan Cave (12–7 ka), Guangxi Zhuang Autonomous Region,southern China

The cobble industry in southern Chinese provinces was for a long time called large and simple “chopper-chopping tool tradition,” which persisted from the Early Pleistocene to the Middle Holocene. This recognition is 50% true and 50% false because it does not reflect the entire archaeological reality. On the one hand, the so-called “chopper-chopping tool” category often characterizes or dominates the lithic assemblages; on the other hand, a “chopper” from site A could be different from another “chopper” in site B. This distinction is important since the typological names lost their validity when conducting comparative studies inter- and intra-sites. The cobble tool industry of the Zengpiyan cave site (Guilin City, Guangxi Zhuang Autonomous Region, China) is an excellent example of this terminological dilemma, which presents some extent of both simplicity and complexity, underestimated in its production strategies. We could also find substantial flexibility between the two ends (simplicity and complexity) during the selection and exploitation of the cobble/pebble blanks. Although chopper-chopping tools are predominant at the site, the variability of their morpho-structures and techno-functional organization indicate that they are not just simple chopper-chopping tools sensu stricto but have their specific characteristics which could not be found in their typological names. In this context, we need to re-evaluate the previously so-called “choppers” in south China with a techno-functional approach before comparing different lithic assemblages. Compared with other contemporary sites, the Zengpiyan technical system is not far from those found in South China and mainland Southeast Asia. For example, it has similarities with the Sonvian phenomenon in northern Vietnam and some other sites in southern China. While very different from the Hoabinhian technocomplex, the Zengpiyan assemblage raises questions about the kinship with other Late Pleistocene lithic facies. Further technological studies on a larger number of sites in these regions are still needed to understand better the prehistoric techno-cultural relationships between the industrial assemblages of southern China and those bordering the Far East.     

Baume Flandin et Abri du Maras : deux exemples de débitage laminaire du début du Pléistocène supérieur dans la Vallée du Rhône (sud-est, France)

Assemblages with blades are rare in the south of Europe and most of them are dated from the isotopic stages 4 and 3. The blades are, otherwise, often produced on Levallois cores with unipolar and bipolar methods. According to the sites, the blades are more or less retouched as different kinds of tools. However, there are no specific links between this blank and the tools, and never between the artefacts and the fauna remains (activities). The hypothesis of traditions or needs among time and space are asked without answer, while the blades are present in the north of Europe and the Near East in far older periods. Several sites can be considered to describe the variability of the technical behaviour and the archaeological context of the blades in south-east France, extending to north Italy. The two sites, Abri du Maras and Baume Flandin, located in the middle Rhône valley, yield evidence of a laminar debitage linked to a Levallois method and a “direct” method. The technical choices used in the two sites are accorded to the specificity of the laminar assemblages in the southern Europe.     

Chaînes opératoires and resource-exploitation strategies in chimpanzee (Pan troglodytes) nut cracking

We apply archaeological methods to extend our knowledge of chimpanzee material culture. The cha?ne opératoire conceptual framework, as introduced by ethnography, established technology as a phased process. Prehistoric archaeology adopted this concept to elucidate technological variability in tool-making procedures, based on knowledge of tool functions or subsistence patterns. We focused on the detection of operational sequences by wild chimpanzees (Pan troglodytes verus) when nut cracking with lithic implements at the sites of Bossou and Diecké, Guinea, West Africa. Thus, while it has recently been claimed that chimpanzees leave behind recognizable assemblages of stone hammers that can be morphologically distinguished from Oldowan hammers, this is the first study to focus specifically on the existence of operational sequences during the utilization of stone tools by wild chimpanzees. By combining primatological and archaeological methods and examining ecological areas inhabited by different chimpanzee groups, we sought technological variability and identified variables influencing regional diversity in tool typology and technology. We compared three case studies: (1) Bossou-direct recording of experimental nut-cracking sessions; (2) Bossou- direct and indirect monitoring of nut-cracking sites in the wild; (3) Diecké-indirect monitoring of nut-cracking sites in the wild. Results suggest that chimpanzees perform sequences of repeated tool transport and nut cracking. Data show discrimination of tool functions based on tool features. We identified the most technologically complex tool for nut cracking, which was composed of four stones. We found regional diversity in chimpanzee stone assemblages. Raw-material type and tool mobility constrain technological development in human and nonhuman primates. Spatial analysis of tool distribution indicates a pattern of resource-exploitation strategy, revealing affinities with Oldowan.     

Mechanistic Approach to the Structure of Animal Communities: Anolis Lizards and Birds

The role of competition in structuring animal communities iscurrently hotly debated. One of the crucial assumptions underlyingthe concept of resource partitioning as a function of competitionis the jack-of-all-trades, master of none principle: differencesin morphology and behavior results in the different abilitiesto use different resources and hence results in the potentialto partition those resources. Examination of the manner by whichhabitat structure constrains resource use offers a mechanisticapproach to study community structure. Anolis lizards providea useful example to illustrate the approach since in the WestIndies they show consistent assemblages of ecomorphs from islandto island. The bases for a strong influence of habitat structureon the resource use pattern of anoles are reviewed as a preambleto a new analysis of the 2-species Anolis assemblages in theLesser Antilles. The two species on the island of Grenada showa strong morphological segregation associated with differentuse patterns of similar microhabitats. The two species on St.Kitts in a richer habitat show strong microhabitat segregationbut little morphological difference apart from size. A relatedpair of species on Antigua in a habitat similar to that on Grenadaappear to be constrained in the degree of habitat segregationand, perhaps as a consequence, show morphological segregationsimilar to that documented for the species on Grenada. Thesecomparisons provide further evidence of the strong influenceof habitat structure on the morphology and habitat partitioningpatterns of coexisting species. Field and experimental studiesof coexisting bird assemblages suggest that details of habitatstructure also have a strong influence on their morphology andresource use patterns. Studies focusing on the manner by whichanimals use different microhabitats provide a useful, mechanisticapproach to study the influence of competition on communitystructure     

Age and Date for Early Arrival of the Acheulian in Europe (Barranc de la Boella,la Canonja,Spain)

The first arrivals of hominin populations into Eurasia during the Early Pleistocene are currently considered to have occurred as short and poorly dated biological dispersions. Questions as to the tempo and mode of these early prehistoric settlements have given rise to debates concerning the taxonomic significance of the lithic assemblages, as trace fossils, and the geographical distribution of the technological traditions found in the Lower Palaeolithic record. Here, we report on the Barranc de la Boella site which has yielded a lithic assemblage dating to ∼1 million years ago that includes large cutting tools (LCT). We argue that distinct technological traditions coexisted in the Iberian archaeological repertoires of the late Early Pleistocene age in a similar way to the earliest sub-Saharan African artefact assemblages. These differences between stone tool assemblages may be attributed to the different chronologies of hominin dispersal events. The archaeological record of Barranc de la Boella completes the geographical distribution of LCT assemblages across southern Eurasia during the EMPT (Early-Middle Pleistocene Transition, circa 942 to 641 kyr). Up to now, chronology of the earliest European LCT assemblages is based on the abundant Palaeolithic record found in terrace river sequences which have been dated to the end of the EMPT and later. However, the findings at Barranc de la Boella suggest that early LCT lithic assemblages appeared in the SW of Europe during earlier hominin dispersal episodes before the definitive colonization of temperate Eurasia took place.     

Tracing microfossil residues of cereal processing in the archaeobotanical record: an experimental approach

Interdisciplinary methodological approaches are fundamental for studying tool use and crop processing patterns in the archaeological record. Many archaeological studies of plant microfossil evidence, primarily those of phytoliths, starch grains and pollen, are concerned with processing methods which can be replicated through experimentally produced plant residues. However, most of these studies rely on crop identification through the presence or absence of such microfossils while giving little or hardly any weight to taphonomy and formation processes, which are critical for interpreting archaeological contexts. An investigation of experimentally produced phytolith and pollen assemblages provides the opportunity to evaluate the impact of cereal processing on both microfossils. Controlled experiments were conducted at the Museum of Menorca, Balearic Islands, Spain, for assessing microfossil taphonomy using Iron Age Talayotic tools and Hordeum vulgare (hulled barley) grown nowadays on the island. For dehusking, a sandstone mortar and a wooden pestle were used outdoors, whereas grinding took place indoors using a limestone quern and handstone. The results indicate that the size of multicellular or anatomically connected phytoliths decreases as a result of mechanical degradation suffered through processing activities, whereas the proportion of cereal pollen grains increases through these processes. Additionally, experimental samples from dehusking and sieving provided abundant evidence of floral bracts, and also of other plant parts and even different plant species, such as phytoliths from leaves and stems and non cereal pollen taxa, which were also to be found on the surfaces of the ground stone tools. These findings highlight the importance of integrating different lines of microfossil evidence and taking into account formation and taphonomic aspects, as well as the value of experimentally produced data for a better understanding of tool use and crop processing.     

An experimental approach for tracing olive processing residues in the archaeobotanical record, with preliminary examples from Tell Tweini, Syria

To help the interpretation of possible olive processing residues at archaeological sites, this study examines the changes which occur in pre- and post-charring breakage surfaces of olive stones at tissue level. Fresh olives were experimentally broken before and after charring and heated to different temperatures (230, 330, 430°C) in oxidising and non-oxidising conditions. The structures obtained by the experiment were studied with scanning electron microscopy (SEM) and the characteristics for the different temperatures, oxidation regimes and pre- or post-charring fractures were recorded. Furthermore, the experimental specimens were compared with recent and possibly old fractures of several archaeological olive stones from Tell Tweini, Syria. Criteria to infer their formation are discussed. These criteria could be developed further through more experimental replication and additional charring variables, and verified extensively on archaeobotanical assemblages in future studies. The described structures and alterations in the olive stones can be observed using reflected light microscopy, which would allow the proposed approach to be rather easy to apply in practice.     

Turning a rock shelter into a home. Neanderthal use of space in Abric Romaní levels M and O

Abstract

Bone refits can be a useful tool to understand the archaeological assemblages. From a taphonomic point of view, they have been used to reinforce some aspects related to the formation processes of the sites. However, they can be used to check some palaeoeconomic and social interpretations of the assemblages as well, providing accurate temporal relationships between the objects. For all these reasons, bone refits are relevant to study the settlement dynamics and the intra-site behaviour of past human groups. The main aim of this work is to present the faunal refits data obtained from two different archaeological assemblages (M and O) of the Abric Romaní site (Capellades, Spain), dated around 55 ka. The study also includes taxonomic, anatomical, taphonomic and spatial analyses. The spatio-temporal proxy allows us to interpret the activities carried out by the Neanderthal groups in different areas during different moments. From here, it is possible to interpret site structure, using ethnoarchaeological data as a referent. The comparison between these two archaeological layers, and between them and other assemblages, both from Abric Romaní and from other sites, can contribute significantly to the knowledge of the intra-site behaviour and settlement patterns at the end of the Middle Palaeolithic.     

Coastal sponge communities of the West Indian Ocean: morphological richness and diversity

Sponges are a reasonably ubiquitous, abundant and highly morphologically plastic taxon. They are very unusual in showing considerable morphological plasticity, not only within higher taxa but within species and at macro and micro scales. In this study we determined the prevalence of sponge morphologies at four coastal study regions in the west Indian Ocean. We show that tropical and subtropical assemblages (in the present study) can be separated on the basis of morphological composition alone (by > 4% arborescent forms or presence of palmate forms in the former). Inter‐tidal sponge assemblages can also be separated from those in the subtidal to a high degree of certainty, also on prevalence of morphologies (absence of tubular or branching forms in the former). The species diversity of many sponge assemblages has been quantified in various environments but, typically, only by specialists. Rarely have equivalent measures been made of morphological diversity. Values of morphological diversity (Shannon H′) were similar, though more variable, to those measured in temperate waters. Substratum nature had a major influence on morphological diversity and evenness, in contrast to geography and bathymetry. As coral reefs were the most diverse and caves and boulders the least, we suggest that substratum heterogeneity and water‐flow complexity are probably the chief determinant of sponge assemblage morphological diversity.     

A comparative measure of biodiversity based on species composition

In conservation planning, species richness and species endemism are the most often used metrics for describing the biodiversity importance of areas. However, when it comes to prioritizing regions for conservation actions these measures alone are insufficient because they do not reveal how similar or different the actual composition of species may be from one area to another. For comparative analysis an additional useful metric would be one that indicates the degree to which the species assemblage in one area is also represented in—or is distinct from—species assemblages of other areas. Here we describe a method for quantifying the compositional representativeness of species assemblages among geographic regions. The method generates asymmetric pairwise similarity coefficients that are then used to calculate separate measures for the representativeness and the distinctiveness of species assemblages in the regions being compared. We demonstrate the method by comparing fish communities among freshwater ecoregions of the Mississippi Basin, and then among smaller hydrological units within two individual freshwater ecoregions. At both scales of analysis, our measures of representativeness and distinctiveness reveal patterns of fish species composition that differ from patterns of species richness. This information can enhance conservation planning processes by ensuring that priority-setting explicitly consider the most representative and distinctive species assemblages.     

垩质年轮法在旧石器时代动物考古学中的应用

在旧石器时代考古遗址研究中,动物的死亡年龄往往能够反映古人类的生存能力及猎物选择趋向;死亡季节则是研究古人类人口数量变化、聚居模式等生存适应问题的重要途径。牙齿垩质切片研究表明,垩质的沉积与树木年轮的生长相似,具有较强的季节规律性和终生生长的特点,可被用于同时推断动物的死亡年龄与季节。垩质年轮法在国外已经有不少较为成功的应用范例,但由于其自身局限及我国旧石器时代动物考古学的发展状况,这一方法至今并未得到较好的应用。本文主要介绍垩质年轮法原理及其在旧石器时代考古学研究中的应用实例,并对该方法在国内遗址相关研究中的应用前景做出展望。