Revision of the family Idelinellidae,with a review of the Permian Eoblattida (Insecta)

Permian Eoblattida, which include the families Tillyardembiidae, Protembiidae (=Sylvardembiidae), Soyanopteridae, and Idelinellidae, are reviewed. The latter family is revised; it includes Idelinella macroptera Storozhenko, 1997 from the Middle Permian (Lower Kazanian Substage) of northern European Russia, Permostriga augustalis Novokshonov, 1999, Sylvastriga miranda Aristov, 2004, Strigulla cuculiophora (Aristov, 2002) comb. nov., Cucullistriga cucullata Aristov et Rasnitsyn, gen. et sp. nov., Scutistriga scutata Aristov et Rasnitsyn, gen. et sp. nov. from the Lower Permian (Kungurian Stage) of the Urals, Permeoblatta borealis Rasnitsyn et Aristov, 2010 from the Upper Permian (Upper Severodvinian Substage) of northern European Russia, and probably also Rasstriga americana Aristov et Rasnitsyn, gen. et sp. nov. from the Upper Carboniferous (Desmoinesian Stage) of Illinois.     

New taxa of Mesorthopteridae (Insecta: Grylloblattida) from the Permian of Russia

New taxa of grylloblattids of the family Mesorthopteridae are described from the Permian of Russia: Parastenaropodites aquilonius sp. nov. from the Soyana locality (Middle Permian); P. circumhumatus sp. nov. from the Kul’chumovo locality; and Tshermyaninus biforis gen. et sp. nov., Permorthopteron foliaceus gen. et sp. nov., Parastenaropodites pannea sp. nov., P. exossis sp. nov., Mesoidelia procera sp. nov., and M. riphaea sp. nov. from the Isady locality (Upper Permian). Two species of the genus Khosaridelia Storozhenko, 1992 (family Permotermopsidae) and one species of the genus Megakhosarina Storozhenko, 1993 (family Megakhosaridae) are transferred to the genus Parastenaropodites Storozhenko, 1996 (family Mesorthopteridae); one species of the genus Parastenaropodites is transferred to the genus Austroidelia Riek, 1954; as a result, four new combinations are established: Khosaridelia rigida Aristov, 2005 =Parastenaropodites rigidus (Aristov, 2005), comb. nov. (Vorkuta locality, Lower Permian of Russia); Khosaridelia vyatica Aristov, 2009 =Parastenaropodites vyaticus (Aristov, 2009), comb. nov. (Karaungir locality, Middle Permian of Kazakhstan); Megakhosarina intricata Aristov, 2008 =Parastenaropodites intricatus (Aristov, 2008), comb. nov. (Soyana locality, Middle Permian of Russia); and Parastenaropodites nervosa Storozhenko, 1996 =Austroidelia nervosa (Storozhenko, 1996), comb. nov. (Madygen locality, Middle Triassic of Kyrgyzstan). An identification key to species of the genus Parastenaropodites is given.     

New and little-known Eoblattida (Insecta) from the Paleozoic of Russia

New taxa of the insect order Eoblattida from the Upper Carboniferous of Russia are described, including Narkemina kata sp. nov., Narkeminopsis inversa sp. nov., Carbonokata storozhenkoi gen. et sp. nov., Tshunoptera ampla gen. et sp. nov., and Evenkiophlebia collucata gen. et sp. nov. from the Chunya locality (Krasnoyarsk Region), Narkemulla sibirica gen. et sp. nov. from the Chunya and Izykhskie Kopi localities, Khakassia (all Cnemidolestidae), and Izykhia tridentis gen. et sp. nov. from Izykhskie Kopi (?Spanioderidae). Narkeminuta permiana gen. et sp. nov. (Cnemidolestidae) is described from the Kedrovka locality (Kemerovo Region; Lower Permian), and Issadische maximum gen. et sp. nov. (Eoblattida incertae familiae) is descriobed from the Isady locality (Vologda Region; Upper Permian). Permeoblatta borealis Rasnitsyn et Aristov, 2010 (Idelinellidae) from Isady is redescribed.     

New insects (Insecta: Eoblattida,Embiida) from the Permian of Russia and the Triassic of Kyrgyzstan,with observations on the origin of webspinners

Locustoblattina segmentata gen. et sp. nov. and L. marginata gen. et sp. nov., new members of the family Mesorthopteridae (Eoblattida) are described from the Middle Triassic of the Madygen locality (Ladinian Stage of the Kyrgyzstan). New members of the Alexarasniidae (Embiida), Alexarasnia limbata Aristov, sp. nov. and Nestorembia shcherbakovi sp. nov., are described from the Upper Permian Isady locality in the Vologda Region and the Madygen locality, respectively. Parastenaropodites fluxa Storozhenko, 1996 (Mesorthopteridae) is redescribed from Madygen. The origin of webspinners from within the Permian eoblattids and the composition of the superorder Blattidea are discussed.     

Revision of the family Epideigmatidae (Insecta: Grylloblattida)

The family Epideigmatidae is revised. New members of this family from the Lower Permian of Russia: Vilvaptera permyakovae, gen. et sp. nov. (Vilva locality, Artinskian Stage, Perm Region) and Tshekardeigma rasnitsyni, gen. et sp. nov. (Chekarda locality, Kungurian Stage, Perm Region) are described. Paraphenopterum unicolor Storozhenko, 1992 (Soyana locality, Kazanian Stage, Arkhangelsk Region) is redescribed. The family Stenoneuritidae is regarded as a synonym of Epideigmatidae; the genera Fayoliella Meunier, 1908 (Upper Carboniferous of France), Fabreciella Carpenter, 1934 (Upper Carboniferous of the USA), and Turbopterum Kukalová, 1964 (Lower Permian of the Czech Republic) are transferred to Epideigmatidae. Fabreciella allegheniensis Carpenter, 1934 is synonimized under F. pennsylvanica Carpenter, 1934.     

New insects (Insecta: Mecoptera,Grylloblattida) from the Middle Permian Chepanikha locality,Udmurtia

New scorpionflies, Asiachorista europaea sp. nov. and Petromantis udmurtica sp. nov. (Mecoptera: Permochoristidae), and new grylloblattids, Tshepanichoptera lacera gen. et sp. nov. (Grylloblattida: Aliculidae) and Miralioma urzhumica sp. nov. (Liomopteridae), are described from the Urzhumian of Udmurtia (Chepanikha locality). Liomopterites novissimus Aristov, 2004 (Liomopteridae) is redescribed.     

Review of the stratigraphic distribution of Permian grylloblattida (Insecta), with descriptions of New Taxa

New taxa of grylloblattid insects (Grylloblattida) are described: Tshekardomina mongolica, sp. nov. (Tshekardominidae) from Bor-Tologoi locality (Severodvinian Stage of Mongolia), Permoshurabia mesenensis, gen. et sp. nov. (Geinitziidae) from Soyana fossil site (Kazanian Stage of Arkhangelsk oblast), Khosaridelia vyatica, sp. nov. (Permotermopsidae) from Karaungir II locality (Vyatkian Stage of Kazakhstan), Euremisca kazanica, sp. nov. (Euremiscidae) from Soyana fossil site (Kazanian Stage of Arkhangelsk oblast), Megakhosarina vyaznikensis, sp. nov. (Megakhosaridae) from Balymotikha locality (Vyatkian Stage of Vladimir oblast), and Parascalicia prokopensis, gen. et sp. nov. (Skaliciidae) from Prokop’evsk fossil site (Kazanian Stage of Kemerovo oblast). The genera Lodevopterum Béthoux, Nel, Lapeyrie et Gand, 2005 (Lodève fossil site, France), Elmopterum Béthoux et Beckemeyer, 2007, and Sigmophlebia Béthoux et Beckemeyer, 2007 (Elmo and Midco fossil sites, United States) are transferred from Grylloblattida incertae sedis to families Euremiscidae, Aliculidae, and Tshekardominidae, respectively. The stratigraphic distribution of Permian grylloblattid families is reviewed.     

New Grylloblattida (Insecta) from the Upper Permian and Lower Triassic of European Russia and Kazakhstan

New grylloblattid insects are described from the Upper Permian and Lower Triassic of European Russia and Kazakhstan: Kuplya minutissima gen. et sp. nov. (Tshekardominidae) from the Severodvinian locality Novo-Aleksandrovka (Orenburg Region); Parachauliodites orthopteroides gen. et sp. nov. (Chaulioditidae), Permofossilis commasticatus gen. et sp. nov. (Permotermopsidae), Megakhosarodes tensilis sp. nov. (Megakhosaridae), and Baharellinus dilaceratus sp. nov. (Blattogryllidae) from the Severodvinian locality Isady (Vologda Region); Dvinopedes salariovensis gen. et sp. nov. (Chaulioditidae) from the Vyatkian locality Aristovo (Vologda Region); Klyazmia karasevi gen. et sp. nov. (Chaulioditidae) from the Vyatkian locality Sokovka (Vladimir Region); Megakhosarodes borealis sp. nov. (Megakhosaridae) from the Vyatkian locality Balymotikha (Vladimir Region); Sigmophlebia rugulosa sp. nov. (Tshekardominidae) from the Upper Permian locality Karaungir (East Kazakhstan Province); and Chauliodites kitshmengensis sp. nov. and C. nedubrovensis sp. nov. (Chaulioditidae) from the Induan locality Nedubrovo (Vologda Region).     

The latest record of mayflies of the family protereismatidae sellards (Ephemerida = Ephemeroptera) and a new species of the family misthodotidae in the Upper Permian of Europe

A new mayfly genus and species, Alexandrinia gigantea gen. et sp. nov., is described in the family Protereismatidae from the Upper Permian locality of Isady (Severodvinian Stage, Poldarsa Formation). Protereisma directum Carpenter, 1979, known from the Early Permian of Oklahoma, United States, is transferred to this new genus. Another new species, Misthodotes tshernovae sp. nov., is described in the family Misthodotidae.     

New and little known blattogryllidae (Insecta: Grylloblattida) from the Permian and Triassic of Russia,Kazakhstan, and Kyrgyzstan

New members of the family Blattogryllidae are described, including Permoblattogryllus praecox gen. et sp. nov. from the Middle Permian Soyana locality in Russia, Protoblattogryllus nedubrovensis from the Lower Triassic Nedubrovo locality in Russia, Madygenocephalus micropteron gen. et sp. nov. (the second known case of brachyptery in grylloblattids), and Costatoviblatta similis sp. nov. from the Middle Triassic Madygen locality in Kyrgyzstan. Protoblattogryllus zajsanicus Storozhenko, 1990 from the Upper Permian Karaungir II locality in Kazakhstan is redescribed. The genus Microblattogryllus Storozhenko, 1990 from Madygen is considered to be a synonym of Protoblattogryllus Storozhenko, 1990. Protoblattogryllus abruptus Storozhenko, 1990 from the Madygen locality is transferred to the genus Mesoblattogryllus Storozhenko, 1990.     

New insects (Insecta: Trichoptera,Eoblattida) from the Lower Permian of Russia

The insect fauna of the Lower Permian Tyulkino locality in the Perm Region of Russia is considered. New members of the orders Trichoptera and Eoblattida are described. Members of the primitive family Microptysmatidae (suborder Protomeropina) are typical of the Lower and Middle Permian deposits of Eurasia and are represented by the two genera: Microptysmella Kukalová-Peck et Willmann, 1990 and Kamopanorpa Martynov, 1928. Kamopanorpa tyulkiana sp. nov. and K. tamarae sp. nov. (Trichoptera, Microptysmatidae) and Parakhosara kamica sp. nov. (Eoblattida, Megakhosaridae) are described.     

New beetles (Coleoptera) from the terminal Middle Permian of China

New beetles are described from the Middle Permian (Upper Capitanian) locality Yinping (Anhui Province, China): Linicupes yinpinensis gen. et sp. nov. (Permocupedidae), Taldycupes pingi sp. nov. (Taldycupedidae), Archosyne permiana gen. et sp. nov., Chaocoleus limnebius gen. et sp. nov. (Ademosynidae) and Asiocoleopsis hongi sp. nov. (Asiocoleidae); the type material of Dikerocoleus divisus Lin, 1982 from the same type locality is redescribed. Comparison with beetles from the Permian of European Russia shows that Yinping beetles are similar to Late Severodvinian beetles rather than to Early Severodvinian and Urzhumian ones.     

New Tatarian representatives of the subfamily Chaulioditinae (Insecta: Grylloblattida: Chaulioditidae) from European Russia

New Late Permian grylloblattids of the subfamily Chaulioditinae (Chaulioditidae) are described: Chauliodites afonini sp. nov. (Sokovka locality, Vladimir Region; Vyatkian Stage), Ch. gomankovi sp. nov. (Novo-Aleksandrovka locality, Orenburg Region; Severodvinian Stage), and Ch. ponomarenkoi sp. nov. (Isady locality, Vologda Region, upper Severodvinian Stage). Stratigraphic distribution of the subfamily is discussed.     

New Grylloblattids of the family Megakhosaridae (Insecta: Grylloblattida) from the Permian of Russia

New Megakhosaridae (Insecta: Grylloblattida) are described from the Permian of Russia: Ivakhosara pectinimedia gen. et sp. nov., Megakhosarina intricata sp. nov. and M. minuscula sp. nov. from the Lower Kazanian of Soyana (Arkhangelsk Region), M. magna sp. nov. from the Urzhumian of Chepanikha (Udmurtia), Kargalokhosara terraefossa gen. et sp. nov. from the Urzhumian of Kargala (Orenburg Region), and Alekhosara reticulata gen. et sp. nov. from the Severodvinian of Novo-Aleksandrovka (Orenburg Region).     

长足虻亚科系统发育研究及三新属记述(双翅目,长足虻科)

采用支序分析的方法首次对古北和东洋区长足虻亚科的24属3亚属之间的系统发育关系进行了分析.结果表明,长足虻亚科为一严格的单系群,其中Ahercostomus、Allohercostomus、Tachytrechus和Aphalacrosoma为一单系群,Tachytrechus和Aphalacrosoma为姐妹群.原为寡长足虻属Hercostomus亚属之一的Gymnopternus与新属Setihercostomus的亲缘关系较近,为有效属.粗柄长足虻属Ludovicius与Sybistroma为一严格单系群,建议合并为一属.弓脉长足虻属Paraclius Coquillet应为Pelastoneurus的异名.建立3新属,即准长毛长足虻属Aphypophyllus gen nov,模式种Ahy-pophyllus sinensis(Yang,1996);准白长足虻属Aphalacrosoma gen nov,模式种Aphalacrosoma postiseta(Yang et Saigusa,2001);毛颜寡长足虻属Setihercostomus gen nov.,模式种Setihercostomus zonalis(Yang,Yang et Li,1998).原为寡长足虻属的亚属Ahercostomus提升为属,模式种Hercostomus(Ahercostomus)jiangchenganus(Yang et Saigusa,2001).建立的新组合为:Ahypophytlus sinensis(Yang,1996)comb. nov. , Aphalacrosoma hubeiense (Yang, 1998) comb. nov., A. postiseta (Yang et Saigusa, 2001) comb. nov.,A. sichuanense (Yang et Saigusa, 1999) comb. nov., Seti hercostomus setifacies (Stackelberg, 1934) comb. nov., S. zonalis (Yang, Yang et Li, 1998)comb. nov., S. wuyangensis (Wei, 1997) comb. nov., S. huangi (Zhang, Yang et Masunaga, 2004) and Ahercostomus jiangchenganus (Yanget Saigusa, 2001) comb. nov. .     

Review of grylloblattid insects (Insecta: Grylloblattida) from the Solikamsk deposits of the Perm Region

The Solikamsk faunas of the Vorkuta series of the Pechora Basin and the Solikamsk Formation of the Perm Region of Russia are reviewed. New taxa are described from Tyul’kino locality (Solikamsk District): Artinska ufimica sp. nov., Sylvaprisca tyulkinensis sp. nov., Ufimoprisca gen. nov. (the type species Sylvaprisca alba Aristov, 2005), U. mica sp. nov., U. kamensis sp. nov. (family Lemmatophoridae), Sylvaella ovalis sp. nov. (family Liomopteridae), Ivakhosara prima sp. nov., and Megakhosarella prisca sp. nov. (family Megakhosaridae). Additionally, Uralioma sp. is described from Shchugor locality (Cherdyn’ District). Two species, Paraprisca solikamskensis Aristov, 2009 (family Lemmatophoridae) from Tyul’kino and Kolvidelia curta G. Zalessky, 1950 (family Ideliidae) from Pokcha locality (Cherdyn’ District)), are redescribed.     

Revision of the Nearctic species of the genus Stiropius Cameron (=Bucculatriplex Auct.) with the description of a new related genus (Hymenoptera: Braconidae)

Abstract. The New World rogadine genus Stiropius Cameron is revised for the Nearctic region and a new related genus from the Americas, Viridipyge , is decribed. Bucculatriplex Viereck is found to be a junior subjective synonym of Stiropius Cameron. Three Nearctic species are recognized: S.bucculatricis (Ashmead) comb.n., S.californicus sp.n. and S.wagneri sp.n. A single Nearctic specis of Viridipyge is described as new: V.prunicola , and a Neotropical species, V.letifer (Mann) comb.n., is found to be congeneric. The biology and systematic relationships of both genera are discussed; the two genera appear to represent a relatively basal branch of the Rogadini lineage.     

New and Little-known Nymphs of Gryllones (Insecta) from the Lower Permian of Russia

New nymphs of the gryllones insects (Insecta; Gryllones) Iblatta attrepida gen. et sp. nov. (Eoblattida incertae sedis) and Tshekardushka artenatis gen. et sp. nov. (Reculida incertae sedis) from the Lower Permian Chekarda locality (Kungurian Stage of Russia) are described. The nymph of Czekardia blattoides Martynov, 1940 (Eoblattida incertae sedis) from Chekarda is redescribed.     

The earliest Mesopsychidae and revision of the family Mesopanorpodidae (Mecoptera)

The family Mesopanorpodidae is revised. Most taxa referred to this family are not related to the type genus Mesopanorpodes Tillyard, 1918. The type species of the latter, Mesopanorpodes wianamattensis,is closely related to Mesopsyche Tillyard, 1917. Therefore Mesopanorpodes is transferred to Mesopsychidae Tillyard, 1917 (= Mesopanorpodidae Tillyard, 1918, syn. n.).The earliest Mesopsychidae are described from the Upper Permian of European Russia (Severodvinian; Isady locality, Vologda Province): Permopsyche issadensisgen. etsp. n. (type species) and Permopsyche rasnitsynisp. n. Two species described under Mesopanorpodes from the Upper Permian of Australia are also included into Permopsyche: Permopsyche belmontensis (Riek, 1953), comb. n., Permopsyche robustus (Riek, 1953) comb. n. The first pre-Triassic Mesopsyche, Mesopsyche incompletasp. n. is described from the uppermost Permian (the town of Vyazniki, Vladimir Province). Bittacopanorpa javorskii Zalessky, 1935 from the uppermost Permian or basal Triassic of Kuznetsk Basin is identified as a hindwing of Mesopsyche: Mesopsyche javorskii (Zalessky, 1935) comb. n. The origin, evolutionary history, and stratigraphic occurrence of Mesopsychidae are discussed.     

New and little known grylloblattida (Insecta) from intertrappean deposits of the Tunguska Basin of Sibiria

New grylloblattids from the Triassic of the Tunguska Basin (Evenkia, Russia) are described, including Chauliodites eskovi sp. nov. (Chaulioditidae) from the Nirungdakan-1 locality, Shurabia lukashevichae sp. nov. and Sh. bashkuevi sp. nov. (Geinitziidae) from the Lower Lyulyuikta-1 locality, Megakhosarodes shcherbakovi sp. nov. (Megakhosaridae) and Protoblattogryllus vittatus sp. nov. (Blattogryllidae) from the Anakit-3 locality. Chauliodites durus (Aristov, 2003) (Chaulioditidae) from the Anakit-1 locality is redescribed.