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1.
    
Studies on elevation gradients in Panama and Costa Rica have shown that leaf‐litter ants exhibit a mid‐elevation peak in diversity. This diversity pattern has been observed in other groups and regions, but uncertainty remains as to just how pervasive it is and what might explain it. Here we examine the robustness of the mid‐elevation peak in ant diversity across the entire Middle American corridor, from Veracruz, Mexico, to Costa Rica. We sampled 56 sites distributed throughout Middle America. All were in closed‐canopy evergreen wet forest, spanning 11° latitude, from near sea level to 2600 m elevation. Ants were extracted from 100 litter samples from each site and identified to genus or species. Model selection was performed on richness and diversity variables to test if ant diversity best fits a linear model or one allowing for a mid‐elevation peak. Linear models were also used to examine the relationships among diversity measures and temperature, precipitation, and seasonality. Species richness measures and diversity indices that incorporate relative abundance show a similar relationship to elevation throughout the region: a truncated bell curve with a mode near 400 m. A cubic relationship is statistically favored over quadratic or linear. Temperature is a significant correlate with diversity, but does not predict a bell‐curve. Precipitation and precipitation seasonality fail to explain much of the variability, and no combination of environmental variables predicts a bell curve. Potential causes of the truncated bell curve include lowland biotic attrition, mid‐point attractors, and ecotonal transitions from lowland to montane communities. Analysis of 17 subclades within ants mostly showed the same truncated curve but six clades were anomalous. Distinctive behavioral or historical features potentially explain their patterns.  相似文献   

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Studies of biodiversity along environmental gradients provide information on how ecological communities change in response to biotic and abiotic factors. For instance, distance to water is associated with several factors that shape the structure and the functioning of ecosystems at a range of spatial scales. We investigated the influence of distance to a perennial water source on ant communities in a semi‐arid savanna in northern Botswana. Ant abundance, taxonomic richness, and both alpha and beta diversity were generally higher during the wet than the dry season. However, there were strong seasonal influences on the effects of distance to water, with more pronounced effects during the wet season. While both abundance and beta diversity declined with increasing distances to water during the wet season, there was a contrasting increase in alpha diversity. There was no major effect of distance to water on taxonomic richness during either season. Beta diversity was as high across as along gradients, and we found support for modular rather than nested community structures along gradients. Our study demonstrated that small‐scale gradients in distance to water can influence several aspects of ant communities in semi‐arid savannas. However, our results also point to strong effects of small‐scale environmental variation, for instance associated with vegetation characteristics, soil properties, and plant community structure that are not directly linked to water access.  相似文献   

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研究群落物种组成和多样性的时空动态对揭示生物多样性的分布规律以及预测全球变化情景下生物多样性的变化趋势具有重要意义。然而,在山地生态系统中,不同海拔梯度的森林群落物种多样性和系统发育多样性如何随着时间尺度的变化仍不清楚。该研究以高黎贡山南段东、西坡海拔梯度(960~2 878 m)森林群落固定监测样带的17个样方为研究对象,基于2004、2008和2013年乔木层(DBH≥5 cm)重调查数据,分析样方内物种组成、物种多样性和系统发育多样性的时空动态变化。结果表明:(1)沿着海拔梯度,物种多样性呈现单峰分布格局,系统发育多样性呈现上升的趋势,系统发育结构呈现聚集到离散或者随机的结构。(2)在时间尺度上,森林乔木层在物种多样性和系统发育多样性上并未发生显著性变化。然而,系统发育结构随着时间的推移呈现更加聚集的趋势。(3)在海拔梯度上,东坡低海拔区域(960~1 381 m)的森林群落样方呈现显著的物种丧失,其植被完全被耕地所替代。其中,诃子(Terminalia chebula)、麻栎(Quercus acutissima)、清香木(Pistacia weinmanniifolia)、枳椇(Hovenia acerba)和假香冬青(Ilex wattii)等为主要的丧失物种。相反,物种获得主要集中在西坡低海拔的样方,群落中丰富度显著增加的物种主要为曼青冈(Cyclobalanopsis oxyodon)、华山矾(Symplocos chinensis)和台湾杉(Taiwania cryptomerioides)等。据此,我们推测高黎贡山海拔梯度森林乔木层的群落结构和多样性的动态变化在中高海拔受群落演替和气候变化的制约,而在低海拔主要受人类活动的影响。该研究结果加深了对高黎贡山亚热带常绿阔叶林植物群落动态变化的认识,也有助于该地区精准保护策略的制定。  相似文献   

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The recently described trait‐based approach is becoming widely popular for a mechanistic understanding of species coexistence. However, the greatest challenge in functional analyses is decomposing the contributions of different ecological and evolutionary processes (e.g., niche‐based process, neutral process, and evolutionary process) in determining trait structure. Taking rodents (Rodentia) in the Hengduan Mountains as our study model, we aim to (1) quantify the vertical patterns of functional structure for head–body length (HL), tail/body ratio (TR), animal component in diet (ACD), and all traits; (2) disentangle the relative importance of different assembly processes (environment, space, and phylogeny) in structuring trait dispersion; and (3) assess the feasibility of Bergmann's rule and Allen's rule along elevational gradient. Our results have suggested that the vertical functional structure pattern varied across these three traits, indicating distinct functional roles in the community assembly process. These nonrandom vertical patterns of HL, TR, and terminal ACD have demonstrated these traits were dominated by different ecological process along environmental gradient. In variance partitioning, high proportion of the spatial variations in trait dispersion was explained by environmental and spatial models, which have provided supporting strong evidence for niche‐based and neutral processes in leading species coexistence. Although the three traits all exhibited apparent phylogenetic signals, phylogenetic relationship within community failed to predict the spatial variations of functional dispersion, confirming the enormous inference of phylogenetic signals in predicting trait structure. By assessing the vertical patterns of HL and TR at order and family levels, we argued that functional adaptation along an environmental gradient is a surrogate of series of complex processes (e.g., environmental filtering, interspecific interaction, and neutral dispersal) acting on multiple functional axes, which results in inconsistence with the empirical rules along elevational gradient.  相似文献   

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为了进一步了解丛枝菌根(AM)真菌群落对不同海拔环境的响应,基于孢子形态学鉴定,研究了西藏高原不同海拔区域主要草本植物AM真菌的群落特征、菌根侵染及其变化.结果表明: 藏东南低海拔区(2200~3400 m)、藏中中海拔区(3400~3900 m)和藏北高海拔区(4300~5300 m)的AM真菌分别为11属31种、11属20种和6属14种.随着海拔上升,孢子密度(r=0.978,P<0.01)、物种丰度(r=0.462,P>0.05)均趋提高,优势种、特有种比例大幅增加,Shannon指数(r=-0.945,P<0.01)极显著下降.不同海拔区之间AM真菌群落Sorensen相似性系数(0.526~0.592)较为接近,仅在总体上随海拔差异扩大略趋下降;藏东南低海拔区、藏北高海拔区菌根侵染率无显著差异,但均显著低于藏中中海拔区.各海拔区内,不同海拔梯度对AM真菌群落、根系侵染亦具显著影响,但影响程度、影响趋势因整体海拔环境不同而异.说明西藏高原AM真菌群落趋于生境选择,受控于海拔所主导的水热环境及土壤环境.  相似文献   

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为揭示森林群落系统发育结构在海拔梯度上的变化及其驱动因素, 本研究以云南哀牢山西坡的亚热带森林群落为研究对象, 以APG III系统为基础框架, 结合DNA条形码序列信息解决末端分类单元亲缘关系的方法, 构建了哀牢山森林群落系统发育进化树, 采用净亲缘指数(net relatedness index, NRI)和最近亲缘指数(nearest taxon index, NTI), 探讨了不同植被类型的森林群落系统发育结构和沿海拔梯度的变化规律。结果表明, 从整体的海拔变化趋势上来看, 哀牢山森林群落系统发育结构随海拔上升由系统发育聚集(phylogenetic clustering)走向发散(phylogenetic overdispersion)或聚集程度降低。在低海拔地区, 群落表现为系统发育聚集, 表明生态位理论中的生境过滤作用在群落构建和生物多样性的维持中起着主导作用; 在中海拔地区, 出现了聚集与发散两种群落系统发育结构并存的现象, 推测可能是生境过滤和竞争排斥两种生态过程共同作用的结果; 在高海拔地区, 群落的系统发育结构因选择的指数不同而出现相反的结果, NRI表现出系统发育聚集, 而NTI却表现为随机或发散, 考虑到高海拔地区的环境胁迫可能促使植物发生趋同进化, 推测其群落构建的生态学过程需要更为综合的研究。本研究揭示群落系统发育结构沿海拔梯度确实存在显著的变化, 证明在生态群落的构建过程中非随机过程起到促进乃至关键作用。  相似文献   

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Along elevational gradients, phylogenetic relatedness patterns constitute a considerable source of information and may shed light on ecological processes that structure communities. This study focuses on community phylogenetic structure of planthoppers, specifically the species-rich and abundant Fulgoromorpha families (Hemiptera, Auchenorrhyncha), Cixiidae and Derbidae + Achilidae, along an elevational gradient on Mount Wilhelm (Papua New Guinea). In order to assess the factors driving planthoppers community composition, we recorded abundance data for planthoppers species at each elevation and we generated a molecular phylogeny of the local species, using Bayesian inference. We analyzed 168 individuals representing 59 local morphospecies. Using a fully resolved and well-supported phylogeny, we then investigated the phylogenetic structure of the communities by performing a Spatial Analysis of Community Diversity. We show that Cixiidae are phylogenetically clustered along the elevational gradient, whereas Derbidae + Achilidae harbor a random structure, suggesting that local adaptation to elevation shapes community structure of Cixiidae, but not that of Derbidae + Achilidae. Our findings highlight the importance of phylogenies in the study of tropical elevational gradients.  相似文献   

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Background: Treeline ecotones represent environmental boundaries that fluctuate in space and time and thus induce changes in plant taxonomic and functional diversity.

Aims: To study changes through time in taxonomic and functional plant diversity patterns along the treeline ecotone.

Methods: In 2002, vegetation was sampled along a gradient from upper montane forest to the treeline–alpine transition in the South Ural Mountains, Russia. In 2014, vegetation was resampled and plant functional traits were collected. We studied spatial and temporal changes in plant species composition, functional composition and functional diversity.

Results: Species composition and diversity changed along the elevational gradient. The functional composition in height, leaf area, specific leaf area and leaf nitrogen content decreased with elevation, whereas functional composition of leaf carbon content increased. We found a temporal shift towards shorter plants with smaller leaves in treeline sites. Functional richness varied in several traits along the elevational gradient, while functional dispersion showed a trend towards increased functional dispersion in height, specific leaf area and leaf nitrogen in the treeline–tundra transition.

Conclusions: Tree encroachment across the treeline ecotone has resulted in a shift in plant species relative abundances and functional diversity, possibly affecting plant community assembly patterns.  相似文献   

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Models applying space-for-time substitution, including those projecting ecological responses to climate change, generally assume an elevational and latitudinal equivalence that is rarely tested. However, a mismatch may lead to different capacities for providing climatic refuge to dispersing species. We compiled community data on zooplankton, ectothermic animals that form the consumer basis of most aquatic food webs, from over 1200 mountain lakes and ponds across western North America to assess biodiversity along geographic temperature gradients spanning nearly 3750 m elevation and 30° latitude. Species richness, phylogenetic relationships, and functional diversity all showed contrasting responses across gradients, with richness metrics plateauing at low elevations but exhibiting intermediate latitudinal maxima. The nonmonotonic/hump-shaped diversity trends with latitude emerged from geographic interactions, including weaker latitudinal relationships at higher elevations (i.e. in alpine lakes) linked to different underlying drivers. Here, divergent patterns of phylogenetic and functional trait dispersion indicate shifting roles of environmental filters and limiting similarity in the assembly of communities with increasing elevation and latitude. We further tested whether gradients showed common responses to warmer temperatures and found that mean annual (but not seasonal) temperatures predicted elevational richness patterns but failed to capture consistent trends with latitude, meaning that predictions of how climate change will influence diversity also differ between gradients. Contrasting responses to elevation- and latitude-driven warming suggest different limits on climatic refugia and likely greater barriers to northward range expansion.  相似文献   

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Aim The factors affecting the distribution of dioecious species have received surprisingly little attention and, as a consequence, are poorly understood. For example, there is a well‐documented negative association between dioecy and latitude, for which we have no candidate mechanisms. Conversely, it has been suggested that the relative proportion of dioecious species should be positively correlated with changes in elevation. Location Costa Rica, Central America. Methods We made use of data on the distribution of 175 seed plant species from a series of plots along a transect in Costa Rica that ranged from 30 to 2600 m a.s.l. to test the prediction that dioecy is correlated with elevation. Specifically, we examined relationships between the proportion of dioecy, at the species and individual levels, and the elevation, species richness, number of individuals, and phylogenetic diversity (PD) of plots. For comparison, we repeated all analyses with monoecious species, which also have unisexual flowers but do not suffer from reduced mate assurance and the seed shadow effect that may be the outcomes of having spatially separated sexes. Results The relative proportions of dioecious species and individuals displayed a unimodal relationship with elevation, both peaking at 750 m a.s.l. In contrast, the relative proportions of monoecious species and individuals displayed negative associations with elevation. In addition, the proportion of dioecious species was significantly positively associated with species richness and with the PD of plots. The proportion of monoecious species was not associated with species richness and was weakly positively associated with the PD of plots. Main conclusions Our results suggest that the selection pressure of elevation on the pollinators and life‐history strategy of dioecious species is much less than expected, and is overshadowed by the as‐yet unexplained correlation of dioecy with species richness. Additional studies of the ecology of cosexual and unisexual (i.e. dioecious and monoecious) species along large environmental gradients are needed.  相似文献   

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兰科是单子叶植物最大的科和被子植物中的第二大科.兰科植物物种多样性的海拔梯度格局因地域或坡向不同而呈不同趋势.南岭国家级自然保护区地处南岭山脉中段南坡,植物资源丰富.作者对南岭国家级自然保护区兰科植物进行了为期4年的调查,记录了其种类、数量、生活型、海拔、生境,分析了其区系成分,并依据IUCN(2001)濒危物种红色名录及保护区的具体情况对其濒危等级进行了评估.从海拔230-1,902m,每200 m分段,共划分为9个海拔段分析其垂直分布格局.结果显示:(1)实地记录兰科植物38属70种,大多数种类数量较少,乳阳管理处的种类最为丰富;(2)随着海拔的上升,总体兰科植物以及不同生活型的兰科植物物种数量均呈现\"中间高度膨胀型\"而峰值偏向低海拔的特点;(3)濒危物种主要集中在600-1,000 m人为活动稀少、环境好的小生境中;(4)800-1,000 m的区系成分最丰富,热带性质的兰花多数分布在海拔1,600 m以下;(5)相邻海拔段的兰科植物种类的相似性系数较高;海拔段相隔越远,相似性系数越小.小生境和水分条件对于兰科植物的生长非常重要,中海拔具有良好的水热条件,因此物种多样性最高;而低海拔地区的人为破坏使得该海拔段兰科植物物种的多样性显著降低.  相似文献   

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1. Investigating how climatic niches change over evolutionary timescales is a necessary step to understanding the current distribution of lineages, yet few studies have addressed this issue using comprehensive datasets. In this study, the evolution of ant climatic niches is investigated at a global scale based on bioclimatic data associated with 163 481 ant occurrence records. The resulting dataset was subjected to principal component analysis, and the scores obtained were used to characterise the main axes of ant climatic niche evolution. 2. Principal component axis 1 (PC1) reflected variation in average temperature and seasonality – consistent with typical tropical/temperate gradients – whereas PC2 was associated with varying levels of aridity. Evolution along these two niche axes was markedly different: differences in the amount of explained variance between PC1 (65%) and PC2 (19%) suggest that climatic niche evolution was nearly three times more pronounced along a tropical–temperate climate axis. 3. There was statistically significant phylogenetic signal on PC1, with genera occupying more tropical conditions diversifying at a faster rate, yet neither of these results is significant on PC2. In addition, most of the ancient ant lineages are associated with conditions of low seasonality and high temperatures. 4. These results provide partial support for the tropical conservatism hypothesis as an explanation for geographical patterns of ant species richness.  相似文献   

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As one of the most important hypotheses on biogeographical distribution, Rapoport's rule has attracted attention around the world. However, it is unclear whether the applicability of the elevational Rapoport's Rule differs between organisms from different biogeographical regions. We used Stevens’ method, which uses species diversity and the averaged range sizes of all species within each (100 m) elevational band to explore diversity‐elevation, range‐elevation, and diversity‐range relationships. We compared support for the elevational Rapoport's rule between tropical and temperate species of seed plants in Nepal. Neither tropical nor temperate species supported the predictions of the elevational Rapoport's rule along the elevation gradient of 100–6,000 m a.s.l. for any of the studied relationships. However, along the smaller 1,000–5,000 m a.s.l. gradient (4,300 m a.s.l. for range‐elevation relationships) which is thought to be less influenced by boundary effects, we observed consistent support for the rule by tropical species, although temperate species did not show consistent support. The degree of support for the elevational Rapoport's rule may not only be influenced by hard boundary effects, but also by the biogeographical affinities of the focal taxa. With ongoing global warming and increasing variability of temperature in high‐elevation regions, tropical taxa may shift upward into higher elevations and expand their elevational ranges, causing the loss of temperate taxa diversity. Relevant studies on the elevational Rapoport's rule with regard to biogeographical affinities may be a promising avenue to further our understanding of this rule.  相似文献   

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While patterns in species diversity have been well studied across large‐scale environmental gradients, little is known about how species’ interaction networks change in response to abiotic and biotic factors across such gradients. Here we studied seed‐dispersal networks on 50 study plots distributed over ten different habitat types on the southern slopes of Mt Kilimanjaro, Tanzania, to disentangle the effects of climate, habitat structure, fruit diversity and fruit availability on different measures of interaction diversity. We used direct observations to record the interactions of frugivorous birds and mammals with fleshy‐fruited plants and recorded climatic conditions, habitat structure, fruit diversity and availability. We found that Shannon interaction diversity (H) increased with fruit diversity and availability, whereas interaction evenness (EH) and network specialization (H2) responded differently to changes in fruit availability depending on habitat structure. The direction of the effects of fruit availability on EH and H2 differed between open habitats at the mountain base and structurally complex habitats in the forest belt. Our findings illustrate that interaction networks react differently to changes in environmental conditions in different ecosystems. Hence, our findings demonstrate that future projections of network structure and associated ecosystem functions need to account for habitat differences among ecosystems.  相似文献   

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青藏高原草地群落组成和结构的海拔梯度格局青藏高原高寒草地是维持区域生态安全的天然屏障,也在一定程度上造就了该区域较高的生物多样性。然而,我们对青藏高原高寒草地植物群落组成和结构的海拔分布格局及其自身维持机制仍知之甚少。本研究在青藏高原东北部沿公路形成的海拔梯度设置了39个实验样地(海拔跨度为2800–5100m),每个样地设5个调查样方进行群落调查,包括物种组成、高度、盖度,评估青藏高原高寒草地植物群落的α和β多样性的海拔梯度格局及其影响因素。研究结果发现草地群落高度随着海拔的增加而显著降低,而群落盖度变化却不显著。随着海拔的增加,植物物种丰富度(α多样性)显著增加,而群落变异性(β多样性)显著降低。约束聚类分析表明,随海拔增加草地群落结构逐渐发生变化,基于此,在这种变化过程中,我们监测到3个渐变的海拔间断点,分别在海拔3640、4252和4333 m处。结构方程模型(SEM)表明,降水增加和温度降低对α多样性有显著的正向作用,但植物群落α多样性的变化显著改变群落变异性。以上结果表明,青藏高原的群落组成和结构沿海拔梯度发生了从量变到质变的过程。  相似文献   

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裸肉足虫作为联结微生物和大中型土壤动物的重要环节, 在土壤生态系统物质循环和能量流动过程中起着重要作用。为探明裸肉足虫群落沿海拔梯度的分布特征及其主要驱动因子, 作者在长白山北坡选择不同海拔梯度(700 m、1,000 m、1,300 m、1,600 m、1,900 m和2,200 m), 采用最大可能数法对裸肉足虫进行了培养计数, 并采用平板培养、标记、分离再培养的方法进行了分类鉴定, 分析比较了不同海拔梯度裸肉足虫的群落组成和结构特征。结果表明: 长白山北坡裸肉足虫物种丰富, 不同海拔梯度裸肉足虫丰富度指数存在显著差异, 且与土壤酸碱度呈极显著正相关关系。其中林分较为单一的岳桦(Betula ermanii)林带(1,900 m)裸肉足虫丰富度最低, 位于植被交错带的针阔混交林带(1,000 m)裸肉足虫丰富度最大, Shannon-Wiener多样性指数和Pielou均匀度指数在不同海拔梯度间不存在显著性差异, 但变化趋势与丰富度一致。聚类分析结果显示, 1,300 m、1,600 m和1,900 m海拔带以及700 m和2,200 m海拔带裸肉足虫群落组成较为相似。典范对应分析(canonical correspondence analysis, CCA)显示, 裸肉足虫群落组成和结构主要受土壤酸碱度、铵态氮以及碳氮比的影响, 而海拔和土壤含水量对其没有显著影响。综上, 裸肉足虫群落多样性随海拔梯度的增加并未呈现递减或单峰的变化趋势, 土壤的基本理化性质是驱动裸肉足虫群落分布的主要因素; 此外, 地上植被也可能通过凋落物和根系分泌物间接影响裸肉足虫的群落组成和多样性。  相似文献   

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