Brittle stars from the upper Cenomanian of the Preafrican platform: First ophiuroid remains for the Cretaceous of Algeria

While Late Cretaceous ophiuroids are relatively well known in Europe, these faunas have been much less studied in North Africa. With the exception of some Tunisian assemblages preliminary described at the turn of the 21st century, nothing is known about the Cretaceous brittle stars of the southwestern Tethyan margin. The present paper seeks to bring the first data about hitherto unknown ophiuroids recently found in the early upper Cenomanian succession of the eastern side of the Preafrican trough (Menaguir section, Algeria). This “community” of brittle stars comprises at least eight species. Most of them are probably new, but have not been formally named here. These are representatives of the families Hemieuryalidae, Amphiuridae, Ophiodermatidae, Ophiacanthidae, Ophiopezidae and probably also Ophiomyxidae and Ophiobyrsidae. Almost all vertebrae are zygospondylous; no streptospondylous vertebrae indicate the absence of the order Euryalida here. Most of the ophiuroids belong to the orders Amphilepidida and Ophiacanthida. Ophiotitanos serrata, Ophiomyxa? aff. jekerica, Ophiojagtus? sp. and some other taxa resembling ophiuroid assemblages from the Late Cretaceous of central, western and northern Europe. With respect to the late Cenomanian age, the depth of the sea and the taxonomic composition, there are some similarities with ophiuroids of the Bohemian Cretaceous Basin. The mid-ramp subtidal facies suggests that brittle stars lived here in a warm, euphotic and probably shallow sea.     

The evolution of the Brachycytheride Ostracoda in the context of the breakup of Pangea

The ostracode Family Brachycytheridae is one of the most common taxa of Late Cretaceous shallow marine microfossils, and occurs in the marginal basins of Africa, Madagascar, the Middle East, South and North America and the Caribbean region. Their migration pathways were controlled largely by the plate tectonic events associated with the breakup of Pangea and the formation of the North and South Atlantic Oceans, the Caribbean Sea and the Gulf of Mexico. This study is an attempt to combine specific information on the plate tectonic events as evidenced by the timings of continental-scale structural movements, sea level dynamics (largely the product of mantle processes), the creation and destruction of migration pathways, and taxonomy of the group, and thus relate deep-seated Earth processes, surficial expression of these movements, and biological evolution. Although the Late Cretaceous Brachycytheridae are very widely distributed, their use for correlation has been hampered due to their conservative external morphology, with all of them having a posteriorly-sloped dorsal margin and a swollen ventrolateral carapace, and all of them were long assigned to a single genus, Brachycythere. In 2002, Puckett defined a new genus, Acuminobrachycythere, based on a distinctive clade of North American brachycytherids. It was also recognized at that time that the members of the family from Gondwana (mainly Africa and South America) were distinct in having a single (unsplit) second adductor muscle scar, whereas all of those from North America had a split second adductor. This tiny difference holds significant paleogeographic—and presumably evolutionary—information. Two new genera are therefore described herein, Sapucariella, which is restricted to Gondwana, and Tricostabrachycythere, which is an early, short-lived genus that occurred around the margins of Gondwana. Three new species are described and one is re-described. In all, there are six genera in the Family Brachycytheridae, which include Acuminobrachycythere, Brachycythere, Kaesleria (which is restricted to the Middle East), Opimocythere (which evolved in the Paleogene of North America and is the last of the Brachycytheridae), Sapucariella and Tricostabrachycythere. Taxonomic information is presented for 85 species, including new images of many of the type specimens. A summary of the breakup of Pangea is also presented, with precise paleogeographic reconstructions based on global magnetic anomalies.     

Ammoniten des Cenomans vom Südrand der westfälischen Kreide zwischen Unna und Möhnesee

Systematically collected ammonites from the different levels of the transgressive Cenomanian of the Haarstrang Mountains between Unna and Möhnesee (southern border of the westfalian Cretaceous, NW Germany) have been investigated. Twenty-one taxa (genera and species), partly previously unknown from this region, have been found and described. The fauna from the immediate transgression level is preserved as phosphoritic internal moulds (steinkerns). Fragmentary preservation and fossil condensation levels indicate reworking processes in these basal Cretaceous deposits. The age of the Cretaceous transgression in the area investigated can be precisely established as Lower Cenomanian (=carcitanensis subzone). The Cenomanian sequence, which is lithofacially very variable, can be, on the basis of the ammonite fauna, subdivided according to the modern Cretaceous stratigraphy for the boreal realm.     

粤西怀集盆地白垩系马屋组的介形类化石

依据闸岗剖面地表及钻井样品的分析鉴定,粤西怀集盆地白垩纪闸岗群马屋组产介形类化石10属17种,本文描述其中9属14种,包括1新种。介形类组合以Eucypris最繁盛,次为Ziziphocypris和Darwinula,称Eucypris hubeiensis-Ziziphocypris simakovi-Darwinula leguminella组合。马屋组与湖北贾店组的介形类组合面貌非常相似,可以进行对比,层位大体相当,其时代应属早白垩世最晚期(Albian)-晚白垩世早期(Cenomanian)。     

Ontogeny and phylogeny of Upper Cenomanian rotaliporids (Foraminifera)

The fossil record of planktonic foraminifera is a key source of data on the evolution of marine plankton. One of the most distinctive groups of Cretaceous foraminifera, the rotaliporids, widely used as a stratigraphic index, has always been considered to be a monophyletic clade. New data on the coiling direction and persistent morphological features of the late rotaliporids from the Upper Cenomanian of the Western Interior Seaway, USA, and the Vocontian Basin of southeast France is used as a phylogenetic proxy. Dealing with key morphological features, the coiling pattern of these keeled morphotypes proves that the rotaliporids group is polyphyletic and composed of Thalmanninella, that displays a dextral-coiling preference, and Rotalipora s.s., that have a proportionate-coiling mode. The stratigraphically youngest rotaliporids with keels co-occur with globular forms; and all morphologies transitional between these morphotypes are observed. The ontogenetic relationships between them are investigated, indicating that loss of the keel was a selective advantage that enabled those rotaliporids to remain in the surface water, thereby avoiding the expansion of the oxygen minimum zone. Two species are observed: Thalmanninella multiloculata and Rotalipora planoconvexa. These species are interpreted as having arisen by neoteny from Thalmanninella greenhornensis and Rotalipora cushmani respectively.     

Découverte d’une nouvelle faune d’âge cénomanien dans la région de Forthassa (Atlas saharien occidental,Algérie) : Implications biostratigraphiques et paléoenvironnementales

The upper Cretaceous of the eastern part of the Ksour Mountains (Western Saharan Atlas, Algeria) is classically subdivided into three fossiliferous lithostratigraphic formations: El Rhelida (lower Cenomanian), Mdaouer (lower-middle Cenomanian) and Rhoundjaïa (upper Cenomanian-lower Turonian). The western part of the Ksour Mountains, which is less documented and poorly known, is here the subject of a detailed sedimentological and palaeontological study. Near Hassi Djeifa locality (Forthassa) the upper Cretaceous is represented by the upper part of Mdaouer Formation and by the lowermost part of Rhoundjaïa Formation. The deposits attributed to the Mdaouer Formation are composed of reddish or green marlstone intercalated by biolaminated limestone, bioclastic limestone with gastropods and bivalves, and sandstone. The selachians (Mafdetia tibniensis, Baharipristis bastetiae), pycnodontiform fishes, and ostracod fauna (Damonella spp., Ilyocypris? sp., Cypridea sp., Paracypria? sp.) coming from marly beds are studied in detail. The two selachian species (Order of Rajiformes) were so far restricted to the Early Cenomanian of Egypt. Our sedimentological and palaeontological data support a probable Early Cenomanian age of the studied deposits and indicate a coastal plain palaeoenvironment continually influenced by the action of tide.     

New hybodont shark assemblage from the Early Cretaceous of the Basque-Cantabrian Basin

Hybodont remains from the Early Cretaceous of the Basque-Cantabrian Basin (Cantabric Range) in the North of Spain are described for the first time. This assemblage represents a highly diversified shark fauna adapted to different feeding habits. Isolated remains of six hybodont genera, namely Hybodus, Egertonodus, Planohybodus, Lonchidion, Parvodus, and Lissodus, have been recovered and described from the freshwater Vega de Pas Formation in the locality of Vega de Pas (Cantabria, Northern Spain). Most of these taxa are known from the English Wealden, making the shark fauna of the Vega de Pas 1 site the most similar assemblage to English Wealden shark fauna among any other site known in Europe.     

Cenomanian-Turonian rudists from Western Sinai, Egypt: Systematic paleontology and paleoecology

The Cenomanian-Turonian sequence is well exposed in western central Sinai and contains a considerable number of rudist species. The identified rudists belong to Radiolitidae, Hippuritidae and Requieniidae. Fifteen species are described, belonging to ten genera: Requienia, Toucasia, Apricardia, Radiolites, Eoradiolites, Praeradiolites, Sphaerulites, Sauvagesia, Hippurites, and Vaccinites. Eleven species are reported in the Cenomanian and four species in the Turonian. Within the Cenomanian, two new species are described: Requienia tortuosi and Eoradiolites lenisexternus. The genus Requienia and the species Apricardia carentonenis d’Orbigny, Eoradiolites syriacus (Conrad), Sphaerulites agariciformis Delamétherie, Sphaerulites depressus Blanckenhorn and Vaccinites cf. grossouvrei (Douvillé) are reported for the first time from Egypt. The Cenomanian and Turonian rudists in the western Sinai show either elevator or clinger morphotypes, with the predominance of the former type; recumbent mode of life is rarely represented by some Requieniidae. The elevators are of isolated and clustered occurrences and more represented in the Cenomanian sequence. Rudists of the study area are mainly of parautochthonous fabrics with low to dense packing. Autochthonous fabrics are also achieved by some species, mainly in the Turonian. The disappearance of rudists from the middle part of the sequence and above the Cenomanian/Turonian boundary is due to a deeper setting that resulted from sea-level rise or seafloor subsidence (shelf drowning). The occurrence of rudists with oysters and other benthic fossils in the siliciclastic and carbonate sediments of the Cenomanian and Turonian sedimentary rocks in the Sinai indicate that the sequence was deposited on a broad, shallow shelf. Although the first marine transgression invaded the central Sinai in the late Cenomanian, transgressive deepening conditions continued until the Turonian. The rudists of central Sinai are of Tethyan affinity with significant relation with North Africa, the Middle East, and Southern Europe.     

Dinosaurs of Portugal

A synthesis on the state of art on dinosaur knowledge in Portugal is presented. The following genera have been recognized: Ceratosaurus, Torvosaurus, Lourinhanosaurus, Allosaurus, cf. Compsognathus, Stokesosaurus, cf. Richardoestesia, cf. Archaeopteryx, Euronychodon, cf. Paronychodon, Dinheirosaurus, Lourinhasaurus, Lusotitan, cf. Pleurocoelus, Lusitanosaurus, Dacentrurus, Dracopelta, Phyllodon, Hypsilophodon, Alocodon, Trimucrodon, Draconyx, Iguanodon, and Taveirosaurus. Most are from Late Jurassic localities at the Lourinhã area and Guimarota. A new genus, Lusotitan, is here raised to include the Late Jurassic ‘Brachiosaurus’ atalaiensis. Lower Cretaceous until Cenomanian material is scarce, except for dinosaur footprints. An interesting Late-Cretaceous, mostly small dinosaur association has been collected between Aveiro and Taveiro. To cite this article: M.T. Antunes, O. Mateus, C. R. Palevol 2 (2003) 77–95.     

Mise en evidence de Campanien continental (facies provencal) a Salavas (Ardeche,S-E de la France): Description,sedimentologie, paleogeographie

The continental sediments in the Northern Alès Basin are dated as Campanian age based on gastropod Melania praelonga MATHERON. The lithology is made up by red shales, conglomerates and pedologic limestones (crusts and nodules) with microcodiums. The conglomerates are heterometric and made up by limestone (Urgonian facies) and standstone boulders of Cenomanian and Turonian age. Also sandy limestones of coniacian age are found, all belonging to the local substratum. This discovery helps to understand the evolution of the Upper Cretaceous paleogeography.     

New Ichneumonidae (Hymenoptera) from the Upper Cretaceous ambers of the Taimyr Peninsula

Two new genera and four new species of Ichneumonidae are described from the Upper Cretaceous ambers of the Taimyr Peninsula: Agapia sukatchevae gen. et sp. nov., Agapteron popovi gen. et sp. nov., Eubaeus abdominalis sp. nov., and Urotryphon baikurensis sp. nov. New detailed diagnoses are provided for the genera Urotryphon and Eubaeus. The genera Catachora, Urotryphon, and Eubaeus, previously placed in the subfamily Tryphoninae, are transferred to the subfamily Labenopimplinae, as well as the new genera Agapia and Agapteron. Possible causes of the miniaturization in ichneumonid wasps in the Cretaceous are discussed.     

New Remains of the Multituberculate Mammal <Emphasis Type="Italic">Barbatodon</Emphasis> from the Upper Cretaceous of the Hațeg Basin (Romania)

The so-called “Ha?eg Island” (Transylvania, Romania) is well known for its rich and peculiar Late Cretaceous vertebrate assemblages, including dwarf dinosaurs, first reported at the end of the 19th century. Besides dinosaurs, other important members of the “Ha?eg Island” terrestrial ecosystems were the multituberculate mammals. Late Cretaceous (Maastrichtian) multituberculates are extremely rare in Europe, being known exclusively from the “Ha?eg Island.” Two genera are reported from this area: Kogaionon, with the single species K. ungureanui, and Barbatodon, with two species: B. transylvanicus and B. oardaensis. Both genera belong to the European endemic family Kogaionidae. We report herein new remains of B. transylvanicus from the Maastrichtian locality of Pui (Hunedoara District) in the Ha?eg Basin, including the most complete dentaries found to date with complete cheek teeth rows. Intraspecific variability of this species in dentary shape, tooth morphology, and size difference is highlighted.     

Palaeobiogeography of late cretaceous belemnites of Europe

A North Temperate Realm, characterized by Belemnitellidae Pavlov, and a South Temperate Realm, characterized by Dimitobelidae WHITEHOUSE, existed throughout the Late Cretaceous, while Tethyan belemnites belonging to Belemnopseidae Naef existed only in the Cenomanian and disappeared afterwards. The North Temperate Realm may be subdivided into North European and North American Provinces. The latter province includes Greenland, Canada, the Western Interior Region of North America, and the Atlantic and Gulf coasts of North America. The belemnites from the North American Province, consisting of populations of the generaActinocamax Miller andBelemnitella d’Orbigny, are closely related to the belemnites of the North European Province and appear to have migrated from this province to North America via Greenland and Arctic Canada. The North European Province extends from Ireland to the Ural Mountains. Belemnites from this province belong to the following genera:Neohibolites Stolley,Parahibolites Stolley,Belemnocamax Crick,Actinocamax Miller,Belemnellocamax Naidin,Gonioteuthis Bayle,Belemnitella d’Orbigny,Belemnella Nowak, andFusiteuthis Kongiel. Two subprovinces within the North European Province have been recognized: the Central European and Central Russian Subprovinces. These subprovinces are well-defined in the late Coniacian-Early Campanian and are characterized by theGonioteuthis stock andBelemnitella stock, respectively. The two subprovinces are less distinct in other periods of the Late Cretaceous and may disappear completely.     

Sur l'origine du genre Microceratina Swanson, 1980 (Ostracoda, Eucytherurinae) et sa présence téthysienne dès le Jurassique supérieur

The discovery in the uppermost Jurassic of Lebanon of a species of the ostracod genus Microceratina Swanson, 1980 (Eucytherurinae), of which the oldest known species was until now of Maastrichtian age, confirms the Tethyan origin of this genus. Two new species are created: Microceratina bhannesensis nov. sp. from the Upper Jurassic (Kimmeridgian) of Lebanon, and Microceratina azazoulensis nov. sp. from the Cenomanian of Morocco. The genera Chapmanicytherura Weaver, 1982 and Erratacytheridea Herrig et al., 1997, are considered as probable junior synonyms of Microceratina. An exhaustive bibliographical analysis allowed to identifying several other Cretaceous species susceptible to be assigned to the genus Microceratina.     

Discovery of charophyte flora across the Cretaceous–Paleocene transition in the Jiaolai Basin

Charophyte assemblages from the Cretaceous/Paleogene (K/Pg) boundary transition in the JZK-2 borehole of the Jiaolai Basin (eastern China) were studied. The charophyte flora includes Tolypella grambastii, Peckichara praecursoria, Microchara cristata, Microchara prolixa, Chara changzhouensis, Lamprothamnium ellipticum, Nodosochara (Turbochara) specialis, Lychnothamnus aff. vectensis, and Lychnothamnus lanpingensis n. comb. The discovery of the ostracod species Porpocypris sphaeroidalis Guan in the upper section of the JZK-2 borehole strongly indicates the arrival of the Paleocene. Three floras of the latest Cretaceous to the early Paleocene were compared among the Jiaolai, Pingyi, and Songliao basins. The Pingyi Basin is special because it is the only carbonate palaeolake that occurred throughout the K/Pg transition in China. It is characterized by the appearance of the Cretaceous brackish water element, Feistiella anluensis (Wang), and is dominated by the Paleocene species, Peckichara varians Grambast. The Songliao and Jiaolai floras grow in clastic lakes; the Jiaolai Basin is thought to correspond to an altitude lake, which was formed in the coastal mountains with high palaeoelevation (≥ 2.0 km), while the Songliao Basin is presumed to have been formed in a low elevation site according to previous studies. This difference in altitude explains why the Songliao Basin is more species-rich than the Jiaolai Basin.     

New xylological data and the biogeography of the Iberian Peninsula during the Early Cretaceous

The position of the Iberian Peninsula during the Early Cretaceous, sandwiched between Laurasia and Gondwana, makes it an important area for the biogeography study of terrestrial ecosystems. Relevant data are, however, scarce. The discovery of silicified wood in the west of the Cameros Basin, in the village Hacinas (Spain), is the first record of the genus Protopodocarpoxylon in Spain and the seventh of Agathoxylon. A new species, Protopodocarpoxylon haciniensis sp. nov., is described. The anatomy of the samples studied shows that paleoclimatic conditions during the Early Cretaceous in the western part of the Cameros Basin were favorable to tree growth, with good water supply during the growth season. Although much impoverished in comparison with Western Europe, the Iberian Early Cretaceous wood floras are clearly Laurasian in affinity.     

ARIADNAESPORITES AND GLOMERISPORITES IN THE LATE CRETACEOUS: ANCESTRAL SALVINIACEAE

Glomerisporites and Ariadnaesporites are two genera based on megaspores and microspores from the late Cretaceous. Each has megaspores with characteristics shared with other late Cretaceous Salviniaceae, including numerous floats, an acrolamella, and a complex perispore. Microspores are borne singly, however, not in characteristic salviniaceous massulae. Microspores of Glomerisporites have a pseudovacuolate basal region and a single spore in an apical neck. Microspores of Ariadnaesporites are structurally like the megaspores but of smaller size. Ariadnaesporites varius occurs in the Cenomanian Stage (early late Cretaceous) and is the oldest presumed member of the Salviniaceae.     

Early evolution in the Pinaceae

Nineteen species of structurally preserved ovulate cones of the Pinaceae are known from the Cretaceous. One of these belongs to Pinus, while the others contain anatomical features necessitating their classification in the organ genera Pityostrobus and Pseudoaraucaria. The six species of the latter group are very similar to one another and probably represent a natural, thought extinct, genus. By contrast, there is so much variety in the twelve Cretaceous species of Pityostrobus that when they are considered with respect to the uniformity of cone structure within each of the modern genera, each of the species of this organ genus may well reperesent an natural genus by itself. All expect one of these fossil forms contain features that are today characteristic of Pinus. This, combined with the Early Cretaceous occurrence of a structurally preserved Pinus cone, suggests that Pinus or something very close to it represents the phylogenetic centrum of the ancestral complex. Lack of cones showing distinct affinity with other modern genera supports this idea and further suggests that, while Pinus was in existence in the Early Cretaceous, other Recent genera of the Pinaceae may not have diverged from the complex until the Late Cretaceous or Early Tertiary.     

Entobia exogyrarum (Frič, 1883) from the Upper Cretaceous of the Bohemian Cretaceous Basin

The validity of the little-known ichnospecies Entobia exogyrarum (Fri?) is verified. Shells of an oyster, Rhynchostreon suborbiculatum (Lam.), represent the substrate for the sponge borings. The borings occur in many Upper Cenomanian to Middle Turonian localities of the Bohemian Cretaceous Basin. This study revealed that the sponges attacked shells of living oysters. Entobia exogyrarum (Fri?) also represents one of the shallowest occurrences of Entobia borings in the Upper Cretaceous rocks.