New enantiornithine bird (Aves, Ornithothoraces) from the Late Cretaceous of NW Argentina

A new enantiornithine bird, Intiornis inexpectatus gen. et sp. nov, is described here. It is based on a partial hind limb found in beds of the Upper Cretaceous Las Curtiembres Formation (Campanian), North-West Argentina. The new taxon is referred to the family Avisauridae on the basis of its cranially convex third metatarsal. Several features suggest close relationships between Intiornis and the avisaurid Soroavisaurus, from the Lecho Formation (Maastrichtian; North-West Argentina). Intiornis was the size of a sparrow, thus representing the smallest Enantiornithes known from South America. The new species shows adaptations for a perching mode of life. Moreover, the hypothesis suggesting that the flying pterosaur reptiles decrease in taxonomic diversity due to competitive interaction with birds is discussed. The new species shows adaptations for a perching mode of life. Moreover, the hypothesis suggesting that the flying pterosaur reptiles decrease in taxonomic diversity due to competitive interaction with birds is discussed.     

Late Turonian (Cretaceous) climate cooling in Europe: faunal response and possible causes

Isochronous variations of δ¹⁸O curves within several European basins indicate a period of Late Turonian climate cooling, which is characterized by two distinct cooling phases, separated by a period of climate stability. Literature data for macrofauna (ammonites, echinoids, and belemnites) indicate that the cooling phases are associated with a southward shift of taxa. Concomitant Late Turonian events (volcanism and relative sea-level changes) suggest the migration to be triggered mainly by relative sea-level falls. The inferred cooling phases are seen in context with a general cooling trend due to the decrease in Mid-Cretaceous volcanogenic CO₂ emission. Short-term stagnation of cooling in the Late Turonian has been probably triggered by renewed volcanism. Due to the general high temperatures during Mid-Cretaceous times, a glacio-eustatic explanation for the coincidence of cooling and sea-level fall is considered unlikely.     

An Early Late Cretaceous Dinosaur Tracksite in Central Yukon Territory,Canada

Dinosaur tracks were first reported from the coal-bearing clastic sequences of the Ross River Block in 1999 by members of a University of Alaska Museum field party, and track sites were confirmed by a joint Alaska-Yukon team in 2000. This fault-bounded sedimentary block is 3 kilometers west of Ross River, in the Yukon Territory. The discovery was followed by two years of field mapping and collection. This research has resulted in the documentation of 251 individual tracks at two separate but stratigraphically related sites, as well as a short (four-footprint) trackway at one of the sites. Six ichnogenera were identified. Ornithomimipus, Amblydactylus, and Gypsichnites were recognized at one site. At a stratigraphically higher site, four ichnogenera were documented including Tetrapodosaurus, Irenesauripus, Amblydactylus, and Columbosauripus. This ichno-assemblage is compared with those of Aptian to Cenomanian age from Alberta, British Columbia, and Alaska.

The discovery of unequivocal dinosaur evidence in a small structural inlier in the Tintina Trench that was previously assumed to be Eocene in age resulted in a restudy of the palynology and biostratigraphy of this coal-bearing sequence and the recent assignment of a middle Albian to early Cenomanian age to the upper part of the dinosaur-bearing interval.     

On Late Jurassic and Early Cretaceous oysters (Bivalvia,Ostreidae) from northern Siberia

The morphology and taxonomic value of morphological features of Late Jurassic and Early Cretaceous oysters from northern Siberia are considered. A new species, Deltoideum exoticum Kosenko, sp. nov. is described.     

Development of calcrete and clinoforms during emergence and flooding of the Late Cenomanian carbonate platform, Jordan

The Late Cenomanian Hummar Formation was studied in three sections in north and central Jordan, at Aameriyya, northeast of Na’ur and the Wadi Haur areas. The base in the Aameriyya area is marked by a subaerial unconformity overlain by a calcrete and a paleokarstic horizon, separating the underlying Fuheis Formation marl from the overlying Hummar Formation limestone. The emergent Aameriyya area is interpreted to have been a paleohigh, as a response to tectonism, and a basin and swell topography is invoked for the Late Cenomanian carbonate platform in this region. The Hummar Formation is believed to form one complete depositional sequence; the calcrete-karst represents a lowstand systems tract, the overlying 2-m massive rudstone/floatstone represents the transgressive systems tracts (TST), and the cortoid grainstone/packstone with clinoforms the highstand systems tracts. The topmost miliolid limestone is probably the late highstand topset of the sequence, followed upwards by the TST of the Shueib Formation marl of the next sequence. The sequence boundary at the upper contact of the Hummar Formation can be correlated regionally whereas the sequence boundary at its base with subaerial exposure has not been reported elsewhere in Jordan, the Negev, or Sinai.     

Late Cretaceous (Cenomanian to Campanian) paleoenvironmental history of the Eastern Canadian margin of the Western Interior Seaway: bonebeds and anoxic events

Upper Cretaceous strata in the Pasquia Hills of the northern Manitoba Escarpment, eastern Saskatchewan, Canada provide a detailed paleoenvironmental and sea-level record of the eastern margin of the Western Interior Seaway. Sediments deposited during the Cenomanian/Turonian Greenhorn marine cycle are dominantly black mudstones deposited in a stratified water column, with bottom-water anoxia recurrently reaching into the photic zone. A middle Cenomanian sea-level lowstand event followed by transgression left a series of bonebeds within the Belle Fourche Member of the Ashville Formation, indicating a sedimentary environment starved of coarse siliciclastics. Maximum sea level resulted in the formation of limestone beds within the Favel Formation, further favoured by reduced terrigenous sediment input compared to the western margin. Limestone sedimentation was followed by a phase of increased freshwater input under lower sea level conditions, and reducing zoo- and phytoplankton diversities. During final Greenhorn regression, eastern Saskatchewan probably turned into a restricted basin severely limiting marine circulation. Poor or absent benthic foraminiferal assemblages and biomarker analysis suggest prevailing watermass stratification throughout the Cenomanian/Turonian transgressive/regressive cycle. This was caused either by a freshwater lid, stratification of Boreal and Tethyan-derived watermasses, or both, to various intensities affected by changing sea level. Basin oxygenation during Niobrara time varies between localities along the eastern margin as documented by presence/absence of benthic and planktic foraminifera.     

Late Jurassic sea-level fluctuations in NW Switzerland (Late Oxfordian to Late Kimmeridgian): closing the gap between the Boreal and Tethyan realm in Western Europe

In Late Jurassic times, the Swiss Jura carbonate platform occupied the transition between the Paris Basin and the Tethys and thus connects the Boreal and Tethyan realm. Up to now, the lack of index fossils in the Reuchenette Formation prevented a reliable correlation between both areas (its sediments are characterised by a prominent sparseness of index fossils). Now, seven recently in situ collected species of ammonites helped to establish a new sequence-stratigraphical frame for the platform sediments of the Reuchenette Formation in NW Switzerland. Based on biostratigraphical data, five third-order sedimentary sequences were assigned to the Late Oxfordian to Late Kimmeridgian. The upper three third-order sequences correspond to the Boreal sequences Kim3–5 of Hardenbol et al. (1998). The deduced large-scale sea-level fluctuations match those from other European regions (Spain, Russia). This biostratigraphically based sequence-stratigraphical frame is a prerequisite to refine correlations within a wider area covering the Swiss Jura and parts of adjacent France and Germany. Electronic Supplementary Material Supplementary material is available for this article at     

Aptychi microstructure in Late Cretaceous Ancyloceratina (Ammonoidea)

The microstructure of aptychi (bivalved calcareous coverings on lower jaws) of three genera of Late Cretaceous Ancyloceratina, Baculites, Polyptychoceras and Jeletzkytes is described for the first time on the basis of well-preserved and in situ material from the Western Interior of the USA and Hokkaido, Japan. Optical and scanning electron microscope observations of aptychi on polished median and cross-sections reveal some variation in their relative size, shape and microstructure among the three genera. The aptychus of Baculites is composed of two calcitic layers: one with tilted lamellae and the other one with horizontal lamellae, whereas those of Polyptychoceras and Jeletzkytes consist of a thin layer with horizontal lamellae. Comparison with aptychi (e.g. Laevaptychus) of Jurassic Ammonitina shows that the aptychi of Ancyloceratina differ from those of Jurassic Ammonitina in the smaller number of layers and the absence of a sponge-like structure. We propose for the first time growth models for a sponge-like aptychus of Jurassic Ammonitina and the lamellar aptychus of Cretaceous Ancyloceratina. The remarkable microstructural variation of aptychi observed in Mesozoic Ammonoidea is probably related to the diversity of their modes of feeding and the secondary function of the lower jaws as opercula.     

Sedimentary and biofacies records in calciturbidites at the Devonian-Carboniferous boundary in Moravia (Moravian-Silesian Zone, Middle Europe)

Summary At the Devonian/Carboniferous boundary, major climatic and oceanographic changes influenced sedimentation on carbonate platforms and in peri-platfrom asreas. Three deep-water carbonate successions in Moravia, which were selected to represent different paleotectonic settings, have been studied with the aim of testing the influence of eustatic, climatic and tectonic controls on sedimentation and conodont paleoecology and taphonomy. On the slopes of the wide carbonate platforms of the Moravian Karst Development (Lesní lom and Grygov sections), an exemplary highstand shedding systems developed in the upper Famennian (expansa Zone), marked by a pronounced thickness of their respective calciturbidite successions and an abundance of shallow-water skeletal grains.Palamatolepis— andBispathodus-dominated conodont assemblages contain an admixture ofPolygnathus representing a transported, near-shore component. The eustatic sea-level fall in the praesulcata Zone and the lowstand conditions at the D/C boundary resulted in a decline of carbonate platform production and condensed deposition or nondeposition. In the Lesní lom section, a condensed sequence of turrbiditic calcarenites and shales (Middle praesulcata—lowermost sulcata Zone) was followed by lime mud calciturbidites (sulcata and duplicata Zones). In the conodont assemblages, the first event in the Lower praesulcata Zone was associated with the reduction of ‘mesopelagic’Palmatopic and a bloom of epipelagicPolygnathus communis. The second event in the Middle praesulcata Zone corresponds to the onset of polygnathidprotogranthodid biofacies, indicating a carbonate slope environment. In the Grygov section, a pronounced thickening and upward-coarsening succession of tubiditic calcilutites through calcarenites and intraclast breccias, with poor palmatolepid-bispathodid connodont assemblages (expansa Zone), indicates a progradation of the calciturbidite system associated with sea-level highstand. After a break in sedimentation, covering the interval from the Lower praseulcata to the base of Lower crenulata Zone, thick-bedded, fine-grained calciturbidites were deposited in the Lower crenulata Zone, and are associated with poor, mixed assemblages where siphonodellids and polygnathids predominate. At the isosticha-Upper crenulata/Lower typicus boundary, coasre grained, turbiditic calcarenites and breccias rich in clastic quartz grains and mixed conodont assemblages with reworked Frasnian and Famennian conodonts indicate a deep erosion of the source area, presumably due totectonic uplift (relative lowstand). In the Jesenec section, on the flanks of the volcanic seamount (the Drahany Development), a deep-water Upper Famennian condensed succession of calciturbidites and presumably winnowed pelagic limestones is marked by conodont assemblages of palmatolepid-bispathodid biofacies. More proximal calciturbidites with mixed deep-water and shallowwater conodonts prograde at the top of the Upper Famennian succession (Middle to Upper expansa Zone). A striking hiatus, covering the interval from the Early preaesulcata to the base of Lower crenulata Zone, resulted from extreme condensation and submarine bottom current erosion due to sea-level lowstand in the late Famennian and early Tournaisian. The renewed middle Tournaisian calciturbidite sedimentation with strong evidence of erosion at the source area indicates global eustatic rise and tectonic uplift of the Drahany Development seamounts (relative lowstand). The earlier occurrence of the uplift in the Jesenec area, relative to the Grygov section, shows the advance of tectonic processes over time in the Moravian-Silesian basin (orogenic polarity) as a consequence of Variscan orogenic movements.     

Skeletal anatomy of the Late Cretaceous shark,Squalicorax (Neoselachii: Anacoracidae)

The paleobiology of the Cretaceous neoselachian shark,Squalicorax, has largely been based on isolated teeth. We examined partial and nearly complete skeletons of three species ofSqualicorax, S. falcatus (Aoassiz),S. kaupi (Agassiz), andS. pristodontus (Agassiz), that were collected from the U.S.A. These specimens suggest that the total body length (TL) ofS. falcatus typically measured 1.8–2.0 m, and probably did not exceed 3 m. Moderatesized individuals ofS. kaupi andS. pristodontus perhaps measured about 3 m TL. AlthoughS. pristodontus was the largest form among the three species examined, this taxon possessed a set of large jaws (with large but fewer teeth) relative to its body size compared toS. falcatus orS. kaupi. This suggests that tooth size is not an accurate indicator of the TL if one compares oneSqualicorax species to another. Neurocranial features suggest that the vision ofSqualicorax was not as acute as that of a contemporaneous macrophagous lamniform shark,Cretoxyrhina mantelli (Agassiz) , but olfaction ofSqualicorax may have been better thanC. mantelli. The morphology of placoid scales suggests thatSqualicorax was capable of fast swimming. New skeletal data support the view that the feeding dynamics ofSqualicorax was similar to the modern tiger shark (Galeocerdo Müller & Henle). The present data do not allow for exact ordinal placement, but, contrary to some previous interpretations,Squalicorax can be excluded from the Hexanchiformes and Orectolobiformes. The taxon should more appropriately be placed within the Lamniformes or Carcharhiniformes.      

Sauropod diversity in the Late Cretaceous of southwestern Europe: The lessons of odontology

Recent discoveries in southern France and northern Spain suggest that the morphology of titanosaurian teeth shows much greater variations that previously thought. It is suggested that the different morphotypes are informative at specific or generic level and that titanosaurian genera may indeed be recognized by their isolated teeth. It is also confirmed that juvenile titanosaurian teeth have a rather uniform, cylindrical morphology. Four different morphotypes are described for the Ibero-Armorican Island in the Late Cretaceous.     

Reconstruction of Late Cretaceous coastal paleotemperature from halite deposits of the Late Cretaceous Nongbok Formation (Khorat Plateau,Laos)

Cretaceous evaporites of the Maha Sarakhan Formation in Thailand (e.g., the Nongbok Formation, Laos) have been studied for almost a century as the huge potash deposits in the world. The consistently high local paleotemperatures should lead to huge salt deposits during the evaporation process. Primary fluid inclusions in halite can provide surface brine water temperatures directly and quantitatively. Until now, there have been no data published from paleotemperature of primary fluid inclusions of Cretaceous halite. The non-marine halite from the Cretaceous Nongbok Formation (Laos) precipitated from shallow brine waters with temperatures of 17.7–42.3 °C.     

The first record of Late Cretaceous blackflies (Diptera: Simuliidae) in Asia (Yantardakh,Taimyr)

The blackfly Simuliites yantardakh sp. nov. from the Taimyr amber is described. It is distinguished from the only known Late Cretaceous blackfly from the New Jersey amber Archicnephia ornithoraptor Currie et Grimaldi by the twice larger size, the short pubescent antenna, only slightly expanded antennomeres 1 and 2; the small palpomere 3, which is 1.5 times as wide as palpomere 4; and the shorter mouth appendages.     

Campanian-Maastricthian (Late Cretaceous) veneroids (Bivalvia: Heterodonta) from the Ariyalur Group,South India

Eleven veneroid (Bivalvia) species are systematically described from the Campanian-Maastrichtian (Late Cretaceous) sediments of Ariyalur sub-basin of South India. Among these, only Crassatella (Crassatella) macrodonta (Sowerby) and Arctica lacianata (Stoliczka) were earlier recorded by Stoliczka. Protocardia (Pachycardium) madagascariensis (Colignon), Protocardia (Pachycardium) pauli (Cocquand), Epicyprina angulata (Sowerby), Venilicardia truncata (Sowerby), and Calva (Egelicalva) buttensis (Anderson) were previously unknown from the Late Cretaceous horizons of the Indian sub-continent. The present record also includes a new species Palaeomoera stoliczkai n. sp. erected on the basis of its unique hinge characters and surface features. Lucina (Lucina) cf. fallax Forbes, Nicaniella (Nicaniella) aff. trigonoides (Stoliczka), and Corbicula? sp. indet., are tentatively identified because of their imperfect preservation. The present record may add useful information to the understanding of the Late Cretaceous palaeobiology of the region.     

Cenomanian (Upper Cretaceous) bivalves from the Hameimat Massifs,north of Tebessa,Algeria: Systematics,biostratigraphy, palaeoecological and taphonomical remarks

In the eastern Saharan Atlas, particularly in the northern area of Tebessa Province (NE Algeria), the widely outcropping Cenomanian strata display a highly diversified macrofauna, among which bivalves are prominently represented. Twenty-eight bivalve species are here reported for the first time from the Cenomanian of Hameimat Massifs. Based on the stratigraphic distribution of these bivalves, five bivalve zones were recognized, i.e., Costagyra olisiponensis - Gyrostrea delettrei, Rhynchostreon suborbiculatum - Exogyra conica, Ceratostreon flabellatum, Ilymatogyra africana, and Pycnodonte vesicularis vesiculosa - Rastellum carinatum zones. Correlation to the ammonite biozones of the same region as follows: the Costagyra olisiponensis - Gyrostrea delettrei and the Rhynchostreon suborbiculatum - Exogyra conica zones occur respectively in the Sharpeiceras schlueteri and Mantelliceras saxbii subzones of the lower Cenomanian Mantelliceras mantelli Zone. The Ceratostreon flabellatum Zone is correlated with the middle Cenomanian Acanthoceras rhotomagense Zone. The Ilymatogyra africana Zone is correlated with the upper Cenomanian Calycoceras naviculare and the Metoicoceras geslinianum zones. Finally, the Pycnodonte vesicularis vesiculosa - Rastellum carinatum Zone represents the uppermost Cenomanian. Detailed analysis of biometrical and morphological features of these bivalve specimens provides the most reliable tool within the scope of palaeo-environmental reconstitution and the many palaeo-ecological variables that had driven the development and distribution of these macro-invertebrates. Comparison of these new data to those of adjacent south Tethyian areas supports the homogeneity of the Cenomanian bivalve faunas. Such an affinity underlines more vividly the favorable marine communications and currents driving the geographic dispersal of these bivalves during the Cenomanian.     

Late Messinian rodents from Verduno (Piedmont,NW Italy): Biochronological,paleoecological and paleobiogeographic implications

The stratigraphic and paleoenvironmental context of the Verduno fossil vertebrate locality is discussed herein based on its rodent record. The Verduno section crops out in the southern part of the Tertiary Piedmont Basin (TPB), and can be included in the Messinian post-evaporitic Cassano Spinola Fm., chronologically corresponding to the so-called Lago-Mare event. Rodents are represented by a relatively rich assemblage. Murids are by far the most diverse and abundant, with at least four taxa, including the common Centralomys benericettii and Paraethomys meini, and the rare Apodemus gudrunae and Occitanomys sp. Cricetids are represented by a single species, Apocricetus cf. A. barrierei. Muscardinus aff. M. vireti appears to be the only glirid present at Verduno. The Verduno rodent assemblage shares some taxa with other Messinian post-evaporitic localities from Italy bearing continental vertebrate remains, such as Brisighella (central Italy) and Moncucco Torinese (NW Italy) (e.g., C. benericettii, P. meini) and, possibly, with Ciabòt Cagna (NW Italy). However, the general structure of these four Messinian assemblages displays substantial differences, which may reflect different palaeoenvironmental conditions.     

Continental, brackish and marine carbonates from the Lower Cretaceous of Kolone–Barbariga (Istria, Croatia): stratigraphy, sedimentology and geochemistry

During the Early Cretaceous, wide areas of the Dinaric–Adriatic Carbonate Platform emerged for long periods. The Hauterivian–Barremian carbonates from Kolone–Barbariga show a few typical examples of lacustrine facies with dinosaur bones and brackish/palustrine facies. The sequence of the platform is made for the most part by subtidal and intertidal limestones. The bone levels are located in a large depression few meters deep in the uppermost Hauterivian marine limestones. The filling facies of this depression are made by oncolitic rudstones and algal boundstones, which represent marginal lacustrine facies, and by laminated limestones, thin stromatolitic levels and distal fringes of rudstones which represent relatively open lacustrine facies. The fossil content is characterized by rare charophyte stems, ostracods, gastropods and plant remains, while typical marine fauna is absent. At the Hauterivian–Barremian boundary a major emersion event has been observed, then a slow transgressive phase occurred. The transgressive facies are primarily made by mudstones with ostracods, charophytes and Spirillina (brackish and probably freshwater facies), wackestones with Ophtalmidiidae and rare dasyclad algae, storm layers with gastropods and miliolids and breccia-like dinoturbated beds. Wackstones, packstones and very rich in dasyclad grainstones outcrop at the top of the section, representing the maximum of the transgression. Trace elements content, carbon and oxygen stable isotope analyses have been performed to aid the palaeoenvironmental interpretation. In this geological setting, Barium seems to discriminate between brackish and freshwater facies. The isotopic values of the marine carbonates appear to depend on early diagenetic processes, meanwhile lacustrine facies seem to show a weak signal of the depositional environment.     

Nematode (Nematoda: Mermithidae) and hairworm (Nematomorpha: Chordodidae) parasites in Early Cretaceous amber

The present report describes a mermithid nematode (Nematoda: Mermithidae) and a gordiid hairworm (Nematomorpha: Chordodidae) from Early Cretaceous Burmese amber dated at 100-110 million years. The mermithid, Cretacimermis protus sp. n., is emerging from a biting midge (Diptera: Ceratopogonidae) while the hairworm, Cretachordodes burmitis, gen. n., sp. n. had already emerged from its host. These rare specimens represent the first fossil mermithid parasite of a ceratopogonid midge and second oldest described nematode and the earliest known and only Mesozoic fossil of the phylum Nematomorpha. A list of previously described fossil mermithids is included.     

A Late Cretaceous Chimaerid (Chondrichthyes, Holocephali) from Seymour Island, Antarctica

A chimaerid holocephalian, Chimaera zangerli sp. nov., is described from both palatine tooth plates, two tooth plate fragments, and part of the chondrocranium in a nodule from the Maastrichtian of Antarctica. Possibly the oldest known chimaerid, C. zangerli sp. nov. exhibits a tooth plate which is a morphological intermediate between that of the Jurassic Ganodus and the Oligocene C. rupeliensis . The presence of C. zangerli sp. nov. before the end of the Cretaceous is evidence of the early evolution of the Chimaeridae, considered the most derived of the chimaeroid fishes.