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1.
SUMMARY 1. A new method is described for sampling the drift in large rivers continuously and automatically, using equipment based on the marine, continuous plankton sampler of Hardy (1936).
2. Between April and November 1989, about 790 h were sampled continuously, equivalent to a water volume of 2937m3 and the total catch was 527 drifting macroinvertebrates, in forty-nine taxa. Twenty-five of these taxa are recorded in the drift for the first time and twenty-eight of the drifting taxa were present in benthos samples from the Danube.
3. Total drift density varied between 12 and 31 animals per 100m3 of water sampled throughout the year, with a maximum of 31 animals in May, The composition of major faunal groups showed a significant seasonal pattern, with Oligochaeta and Diptera predominant in spring, Crustacea and Insecta in summer and only Crustacea in autumn. The overall density of the macrozoobenthos from October 1986 to December 1987 was about 19360 animals m−2 and the proportion of total benthos animals, drifting at any instant in time, ranged from 0.0026 to 0.0064%.
4. The relationship between drift density day−1 and mean daily discharge was described by a power-function. Total mean drift rate of macroinvertebrates in the Danube was estimated to be 13 600 000 animals per 24 h and the mean drift distance was estimated to vary between about 4 and 31 m, dependent on the animal group and the water velocity.
5. No obvious consistent diel pattern could be established from the continuous samples, and no marked diel rhythm could be detected for Oligochaeta, Diptera larvae and Crustacea.  相似文献   

2.
Water abstraction modifies the environmental conditions of stream ecosystems, which can affect invertebrate assemblages by altering drift. We examined this issue with a before–after–control–impact design experiment in which we diverted 90% of the natural flow from a headwater stream. We measured flow-reduction effects on drift densities (animals/m3), total drift rates (animals leaving the reach per hour) and net balance of invertebrates entering or leaving the Impact reach. We also identified the specific taxa and traits that drove these responses. The sudden decrease in flow promoted a 12-fold increase in overall drift density at the Impact reach, which was primarily driven by filterers, shredders and taxa associated with fast velocities, such as simulids. By contrast, drift densities of other abundant taxa, such as Baetis, Esolus and chironomids, increased less than what could be expected from the magnitude of flow reduction. While drift rates remained unchanged after water abstraction, the Impact reach became a net invertebrate exporter indicating that many taxa drifted actively as a response to stressful conditions rather than passively, which would be reduced by water abstraction. Therefore, our results suggest that water abstraction influences drift, with potentially important consequences for the invertebrate assemblages and ecosystem processes further downstream.  相似文献   

3.
Artificial substrates were used to monitor the invertebrate communities below a power peaking impoundment and a flow re-regulating impoundment on the Missouri River on northeastern Nebraska, USA. Invertebrate communities on shallow samplers subjected to exposure from diel fluctuations in flow averaged 3 taxa per sampler and 91 organisms per square meter. In the absence of diel fluctuations, number of taxa per sampler increased to 12 and mean densities increased to 743 per square meter. Fluctuating discharges had no significant effect on numbers of taxa or densities on continually submerged artificial substrates. However, greater numbers of taxa and total densities were observed on deep (107 cm) versus shallow (30 cm) samplers below both impoundments.Polycentropodidae (Trichoptera), Chironomidae (Diptera) and Oligochaeta were observed to tolerate diel fluctuations and exposure below Fort Randall Dam while Hydropsychidae (Trichoptera) and Heptageniidae (Ephemeroptera) were extremely intolerant.Collector-gatherers and predator-engulfers were the predominant functional groups colonizing samplers below both impoundments. Higher numbers of collector-gatherers, collector-filterers and scrapers were observed from samplers in the absence of diel flow fluctuations.This study corroborates results from a number of other investigations, separated in space and time. Examined collectively, these studies provide the framework for understanding power peaking as a stress to benthic stream communities.  相似文献   

4.
The composition and abundance of the meiofauna and macrofauna were studied in a survey carried out within 6 locations in a mangrove at the Island of Santa Catarina, South Brazil. Nine meiofaunal taxa were registered with densities ranging between 77 and 1589 inds.10 cm?2. The nematodes, composed by 94 putative species (86 genera), largely dominated the meiofauna. The most abundant genera were Haliplectus (Haliplectidae), Anoplostoma (Anoplostomatidae) and Terschillingia (Linhomoidae). Contrary to the meiofauna, the macrofauna showed a low number of taxa (only 17 recorded) and abundance (up 7250 inds.m?2). The macrofauna was mainly composed by deposit feeders, and numerically dominated by oligochaetes and capitellid polychaetes. For both components, differences in the composition and abundance along the sampling sites were significant but not primarily related to the typical variations along estuaries, such as salinity. The results of the stepwise multiple regression analyses showed that the detritus biomass (ash-free dry weight) was the most important predictor of faunal densities and diversity. The clear relationship between detritus and fauna, together with the contrasting community structure of the two component of the benthos suggest that the meiofauna showed a high efficiency in exploiting the micro-habitat created by the presence of the detritus. Yet the macrofauna, potentially the main consumer of the debris, is negatively affected by their low palatability and poor nutritive value.  相似文献   

5.
The effects of pulsed acidification on invertebrate densities and drift, and water chemistry, in a high altitude Sierra Nevada stream were measured using artificial stream channels. Water was diverted from the Marble Fork of the Kaweah River, California, U.S.A., through twelve replicate channels; however, low flow in the summer of 1985 eliminated all but four of these channels. Channels were stocked with natural substrates and organisms from the Marble Fork of the Kaweah River. After a three week acclimation period, we simulated a low pH rain event by adding acid (H2SO4 and HNO3) to two of the channels, reducing pH to 5.0 for 6 hours. The other two channels acted as controls (pH 6.4). During acid additions, Baetis spp. drift in acidified channels was ca. 7 times higher than in control channels (F = 39.02, p < 0.025; data fourth root transformed, ANOVA), and the percentage of drifting baetids that was dead was significantly higher in acidified than control channels (46% vs. 0%, F = 29.86, p < 0.05; arcsine square root transformed data, ANOVA). Other taxa showed no significant drift responses, and benthic densities of all taxa showed no effects two days after acidification, probably owing to rapid recolonization by invertebrate drift in influent waters. Stream chemistry data are presented; heavy metal concentrations did not significantly increase in the 2 m stream channels.  相似文献   

6.
The Yukon Delta, a low alluvial tundra in western Alaska, has more than 105 thaw-basin ponds within its 70000 km2 area. In 1984 and 1985, 68 ponds in three interior areas of the Delta were surveyed to determine limnological features, macroinvertebrate fauna, and trophic character. Ponds ranged up to 90 ha in area, 2 m in depth, and 17 m in elevation, and occurred in various temporal stages of growth and senescence. Among the 18 major invertebrate taxa collected, in order of decreasing frequency of occurrence, Trichoptera, Hemiptera, Diptera, Pelecypoda, Isopoda, Coleoptera, Gastropoda, and Oligochaeta were found in over 50% of the ponds. Trichoptera, the only taxon occurring in all ponds, was represented by 22 species of 6 families. The average Delta pond had 6.6 of the nine more common taxa. This measure of faunal richness was similar among study areas but was higher in low-tundra (sea level) ponds and in older ponds on raised tundra. In comparison, lentic invertebrate communities in five other areas of Alaskan and Canadian tundra had fewer taxa and also lower average richness based on occurrence of the same nine taxa.  相似文献   

7.
The headwaters of karst rivers experience considerable hydrological variability, including spates and streambed drying. Extreme summer flooding on the River Lathkill (Derbyshire, UK) provided the opportunity to examine the invertebrate community response to unseasonal spate flows, flow recession and, at temporary sites, streambed drying. Invertebrates were sampled at sites with differing flow permanence regimes during and after the spates. Following streambed drying at temporary sites, dewatered surface sediments were investigated as a refugium for aquatic invertebrates. Experimental rehydration of these dewatered sediments was conducted to promote development of desiccation-tolerant life stages. At perennial sites, spate flows reduced invertebrate abundance and diversity, whilst at temporary sites, flow reactivation facilitated rapid colonisation of the surface channel by a limited number of invertebrate taxa. Following streambed drying, 38 taxa were recorded from the dewatered and rehydrated sediments, with Oligochaeta being the most abundant taxon and Chironomidae (Diptera) the most diverse. Experimental rehydration of dewatered sediments revealed the presence of additional taxa, including Stenophylax sp. (Trichoptera: Limnephilidae) and Nemoura sp. (Plecoptera: Nemouridae). The influence of flow permanence on invertebrate community composition was apparent despite the aseasonal high-magnitude flood events. Flow permanence was also critical in determining the community response to the spate flows. Following streambed drying at temporary sites, the surficial sediments overlying the karstic bedrock functioned as an effective refugium for several taxa. The development of aquatic insects following experimental rehydration indicated that these taxa survived in dewatered sediments as desiccation-resistant eggs. Electronic supplementary material  The online version of this article (doi:) contains supplementary material, which is available to authorized users. Handling editor: Sonja Stendera  相似文献   

8.
Despite frequent disturbances from flow, stream meiofauna form diverse and abundant assemblages suggesting that they are resistant and/or resilient to flow disturbances. Stream flow profoundly influences benthic invertebrate communities but these effects remain poorly understood. We examined the influence of flow on meiofauna colonization at small spatial scales (2–3 m) using artificial streams in conjunction with similar sites (flow, depth, substrates) in the reference stream (Illinois River, Arkansas). Colonization of meiofauna was found to be rapid and generally increased with flow rates examined (1–2, 6–7, and 11–12 cm s−1). Six of the 10 most abundant taxa successfully completed colonization in artificial channels (equaled or exceeded reference benthic densities) within 5 days. Benthic meiofauna were more abundant in fast flows in artificial channels and in fast and slow flows in reference stream sites. A diverse assemblage of meiofauna was collected from the plankton which was dominated by rotifers, copepods (mostly nauplii), dipterans, and cladocerans. Densities of drifting meiofauna (potential colonists of the benthos) were low (5 no. l−1) and similar among artificial channels and reference sites regardless of flow rates (F 1,18 = 2.19, p = 0.1407). Although densities were low, the numbers of drifting meiofauna were more than sufficient to colonize the benthos. Less than 0.65% of the drifting meiofauna were needed to colonize the substrates of artificial streams. The benthic assemblage paralleled that of the plankton, consisting mainly of rotifers, copepods (mainly nauplii), and dipterans. Evidence for active control over dispersal was observed as meiofauna densities varied between the plankton and benthos over the diel cycle (F 1,18 = 6.02, p = 0.0001 and F 1,18 = 9.88, p = 0.006, respectively). Rotifers, copepods, and nematodes were more abundant in the plankton during the day and in the substrates at night. These results suggest that meiofauna assemblages can change rapidly in response to alterations of habitat patches by disturbance.  相似文献   

9.
Permanent meiofauna taxa and portions of the population of other invertebrates that are temporarily in the meiofauna size class are often precluded from stream studies and assessments. This study was designed to determine the identity, density, and distribution of major meiofauna taxa relative to substrate size in a set of similar headwater streams. Using a coring technique, meiofauna (80 μm–1 mm) and substrate samples were collected from 11 Ozark headwater streams in the Boston Mountain ecoregion of Arkansas, USA. Mean meiofauna density among streams was 1739 ± 436 organisms per l. Permanent meiofauna taxa (Copepoda, Cladocera, Ostracoda, Rotifera, Nematoda, Hydrachnida, and Tardigrada) comprised 22.5% of the organisms collected with a mean density of 394 ± 233 organisms per l; temporary meiofauna taxa (Oligochaeta, Turbellaria, Hydroidea, Chironomidae, Ephemeroptera, and other insects) comprised the remainder with a density of 1346 ± 308 organisms per l. Chironomidae was the most numerous temporary meiofauna taxon, and Hydrachnida was the most numerous permanent taxon. Streams were found to differ significantly in substrate composition and densities of major taxonomic categories. Substrate size was found to predict densities for most of these taxonomic categories. Meiofauna patchiness was reflected in high variability within streams. Canonical correspondence analysis revealed positive associations between Copepoda and Nematoda and silt, and between Copepoda, Nematoda, and Rotifera and fine sand. Hydrachnida and Rotifera were negatively associated with silt and coarse sand, respectively. The potential value of inclusion of meiofauna in stream environmental assessments is discussed.  相似文献   

10.
1. Small permanent streams are coming under increasing pressure for water abstraction. Although these abstractions might only be required on a short‐term basis (e.g. summer time irrigation), the highest demand for water often coincides with seasonal low flows. 2. We constructed weirs and diversions that reduced discharge in three small streams (<4 m width) to test the hypotheses that short‐term water abstractions would decrease habitat availability and suitability for invertebrates, resulting in increased invertebrate drift, reduced taxonomic richness and decreased benthic invertebrate densities. 3. We sampled benthic invertebrates, invertebrate drift and periphyton at control (upstream) and impact (downstream) sites on each stream before and during 1 month of discharge reduction. 4. Discharge decreased by an average of 89–98% at impact sites and wetted width decreased by 24–30%. Water depth decreased by 28–64% while velocity decreased by 50–62%. Water conductivity, temperature and dissolved oxygen showed varying responses to flow reduction among the three streams, whereas algal biomass and pH were unaffected in all streams. 5. The densities of invertebrate taxa tended to increase in the impact reaches of these streams, even though invertebrate drift increased at impact sites in the first few days following discharge reduction. There were a higher proportion of mayflies, stoneflies and caddisflies at the impact site on one stream after flow reduction. There were no changes to the number of taxa or species evenness at impact sites. 6. Our results suggest that for these small streams, the response of invertebrates to short‐term discharge reduction was to accumulate in the decreased available area, increasing local invertebrate density.  相似文献   

11.
The data on seasonal and interannual changes in the taxonomic, structural, and quantitative characteristics of macrozoobenthos communities in rivers with a high salinity gradient are given. A total of 91 benthic invertebrate taxa have been revealed, which were dominated by Cricotopus salinophilus, Chironomus salinarius, C. aprilinus, Tanytarsus kharaensis, Microchironomus deribae, Glyptotendipes salinus (Diptera: Chironomidae), Culicoides (M.) riethi, Palpomyia schmidti (Diptera: Ceratopogonidae), Paranais simplex (Oligochaeta), and Ephydra sp. (Ephydridae) in different years. The fauna of benthic communities is mainly represented by eurybiontic halotolerant species with different ranges of resistance to salinity. The taxonomic composition and diversity of macrozoobenthos communities are closely correlated with water salinity in the range from 4 to 41 g/L; the complex of hydrological and hydrophysical factors (depth, overgrowing, water temperature, pH, etc.) control the distribution and abundance of species.  相似文献   

12.
Drift as a low-energy cost means of migration may enable stream invertebrates to leave risky habitats or to escape after encountering a predator. While the control of the diurnal patterns of invertebrate drift activity by fish predators has received considerable interest, it remains unclear whether benthivorous fish reduce or increase drift activity. We performed a large-scale field experiment in a second-order stream to test if invertebrate drift was controlled by two benthivorous fish species (gudgeon Gobio gobio and stone loach Barbatula barbatula). An almost fishless reference reach was compared with a reach stocked with gudgeon and loach, and density and structure of the invertebrate communities in the benthos and in the drift were quantified in both reaches. The presence of gudgeon and stone loach reduced the nocturnal drift of larvae of the mayfly Baetis rhodani significantly, in contrast to the findings of most previous studies that fish predators induced higher night-time drift. Both drift density and relative drift activity of B. rhodani were lower at the fish reach during the study period that spanned 3 years. Total invertebrate drift was not reduced, by contrast, possibly due to differences in vulnerability to predation or mobility between the common invertebrate taxa. For instance, Chironomidae only showed a slight reduction in drift activity at the fish reach, and Oligochaeta showed no reduction at all. Although benthic community composition was similar at both reaches, drift composition differed significantly between reaches, implying that these differences were caused by behavioural changes of the invertebrates rather than by preferential fish consumption. The direction and intensity of changes in the drift activity of stream invertebrates in response to the presence of benthivorous fish may depend on the extent to which invertebrate taxa can control their drifting behaviour (i.e. active versus passive drift). We conclude that invertebrate drift is not always a mechanism of active escape from fish predators in natural streams, especially when benthos-feeding fish are present.  相似文献   

13.
Quantitative samples of benthic invertebrates were collected from a sandy riverbed of a mountainous stream (Kozu site of Takami-gawa stream, Nara Prefecture), central Japan by core samplers in five sampling occasions through the years 2008–2009. A total of 120 taxa were identified, representing 55 families and 97 genera. Insects formed about 92% of the total recorded taxa and 88% of individuals’ abundance. A total of 111 taxa of aquatic insects, belonging to 49 families and 92 genera, were identified and represented by ten orders. Oligochaeta and Acari were dominant non-insect invertebrates. Diptera was the most diverse insect group, followed by Trichoptera and Ephemeroptera. Dominant taxa were mesoinvertebrates, younger stages of macroinvertebrates, both of which predominantly inhabit the interstitial zone of a sandy riverbed. Both taxon richness and invertebrate abundance were higher in February 2009 and lower in April and August 2008. A few major invertebrate taxa demonstrated distinct seasonal trends; i.e. Nymphomyia alba, Rheosmittia, and Corynoneura were abundant in February 2009. Newly hatched larvae of Larcasia akagiae were abundant in May 2008. This study also demonstrated the effectiveness of core samplers to collect small-sized benthic fauna that inhabit the interstitial or hyporheic zone of the sandy riverbed.  相似文献   

14.
In this study we quantified invertebrate drift and related it to the structure of the benthic community, over a 6–8 month period, in a 4th-order tropical stream in Costa Rica. Relative to reports from similar-sized temperate and tropical streams, drift densities were high (2-fold greater: mean 11.2 m−3; range 2.5–25 m−3), and benthic insect densities were relatively low (>3-fold lower: mean 890 m−2; range 228–1504 m−2). Drift was dominated by larval shrimps that represented more than 70% of total drift on any given date; the remaining 30% was composed of 54 insect taxa. Among insects, Simuliidae and Chironomidae (Diptera) and Baetidae, Leptohyphes and Tricorythodes (Ephemeroptera) comprised 24% of total drift. Drift periodicity was strongly nocturnal, with peaks at 18:00 h (sunset) and 03:00 h. Our results, and those of previous experiments in the study stream, suggest that nighttime drift is driven by the presence of predatory diurnal drift-feeding fishes and nocturnal adult shrimps. There were no clear seasonal patterns over both ‘dry’ and wet seasons, suggesting that benthic communities are subject to similar stresses throughout the year, and that populations grow and reproduce continuously. This revised version was published online in July 2006 with corrections to the Cover Date.  相似文献   

15.
16.
1. Two sampling techniques were used to characterize invertebrate communities in eight, low-order streams along an altitudinal gradient in Costa Rica that represents the last continuous tract of primary forest spanning such extremes in elevation (i.e. near sea level to 2900 m a.s.l.) along the Caribbean Slope of Central America. A standard Surber sampler was used to sample invertebrates on the stream bottom, and drift sampling nets were used to sample invertebrates drifting in the stream flow. 2. Sites were established at 30, 50, 700 1800 and 2700 m a.s.l. In one to two streams per site, six Surber samples were collected, and drift was sampled every 3 h over one 24-h period between April and August 1994. All sites were in primary forest, with the exception of the lowest elevation site (30 m) which was located in banana plantations. 3. Both sampling techniques indicated that Diptera (Chironomidae) and Ephemeroptera were the dominant insect groups at all sites. Disturbed streams draining banana plantations were dominated by Chironomidae and had lower taxon richness and diversity than other sites. 4. While data from benthic samples indicated that insects were the major faunal component (> 90%) at all sites, drift samples were dominated by larval shrimps (> 50%) at the 30 m and 50 m sites. 5. Drift periodicity of invertebrates was observed at those sites characterized by predaceous fishes: nocturnal drift densities were higher than diurnal densities at 30, 50 and 700 m a.s.l., however, no periodicity was observed at 1800 and 2700 m a.s.l. where fish were absent. 6. This study shows the importance of measuring invertebrate drift, in addition to directly sampling the benthos. Drift sampling provided data on a major community component (shrimps) of lowland tropical streams, that would have been overlooked using traditional benthic sampling techniques, and in some cases provided additional information on taxon richness. 7. Based on results of the present study, it is recommended that drift sampling be included as a standard complementary tool to benthic sampling in biological assessments (e.g. bioassessment protocols) of tropical streams, which are often characterized by migratory invertebrate species such as shrimps. Drift samples provide critical information on the presence or absence of shrimps and also on the timing and magnitude of their migration which is an important link between many tropical rivers and their estuaries.  相似文献   

17.
We conducted two experiments to assess drift and benthic invertebrate responses to stepwise and abrupt changes in non-scouring flow in gravel-bed experimental streams. Intuitively, a stepwise flow increase should allow aquatic invertebrates more time to seek refuges than would an abrupt increase. We hypothesized that abrupt flow increases would result in larger increases in taxon richness and in the number of invertebrates in the drift, and a larger decrease in benthic density than would stepwise flow increases. Two kinds of drift response to flow increases were observed in the stepwise experiment: (1) no response (e.g. Caenissp. [Ephemeroptera] and Sphaeriidae [Pelecypoda]); (2) threshold response of some aquatic (e.g. Crangonyx pseudogracilis[Amphipoda]) and semiaquatic (e.g. Ormosiasp. [Diptera]) taxa. Drift richness and drift density in both experiments declined after reaching a peak. The peak was reached almost immediately in the abrupt treatment and later in the stepwise treatment. Maximum richness of taxa and taxon composition in the drift were similar in both experiments. Despite significant increases in drift, stepwise and abrupt increases in flow did not have a significant effect on benthic density. However, relative to reference streams, the percentage of total benthic invertebrates in the drift increased 10× in the stepwise experiment and 33× in the abrupt experiment. These non-scouring increases in flow were non-trivial. Our results suggest that several high flow events of the same magnitude (i.e. 2.5–3.0 fold increases) may cause considerable losses of benthic populations to the drift. The rate of increases in flow appears to be important: abrupt increases in flow had a stronger effect on invertebrate drift than did stepwise increases.  相似文献   

18.
We investigated recolonization by insects of a small headwater stream in the southern Appalachian Mountains that was treated along its entire length with an insecticide (methoxychlor). Initial treatment (December 1985) resulted in massive insect drift. Applications continued seasonally for three years, and drift was measured during each treatment. Taxonomic composition of the drift indicated several responses: (1) Some taxa were eliminated. (2) A number of taxa occurred only sporadically following initial treatment. (3) Early instars for some taxa showed seasonal occurrences which closely paralleled known life cycles and flight periods of adults. Groups which provided strong evidence for aerial recolonization included several Ephemeroptera, Plecoptera (Peltoperlidae and Isoperla spp.), and Trichoptera (Parapsyche cardis, Diplectrona modesta, Pycnopsyche spp., and Lepidostoma spp.). (4) Some long-lived taxa survived and exhibited distinct growth through several treatment periods. These include Odonata (Lanthus and Cordulegaster), some Ephemeroptera, and some Diptera (Tipulidae, Ceratopogonidae, and Tanypodinae). (5) Some taxa which were not present at the time of initial treatment appeared during the experiment. Chironomidae dominated the drift in all samples, and the number of genera did not decrease during the three-year treatment period. Of the 27 chironomid genera identified, only Micropsectra decreased in proportional abundance during treatment. In contrast, several genera (Corynoneura, Meropelopia, Parametriocnemus, and Tvetnia) showed little change in relative abundance. Larsia (Tanypodinae) increased in proportional abundance during the treatment period.  相似文献   

19.
The invertebrate drift leaving the lower end of a small coastal stream on the east coast of Vancouver Island was recorded during low flow conditions. 20,156 animals, weighing 0.96 g (dry weight), were collected in drift nets over a 5 day period in early summer. High and low drifting taxa are listed. Considerable daily variation in total captures occurred and was attributed to a few dominant taxa (primarily harpacticoid copepods, mites and chironomid larvae) exhibiting atypical drift patterns. 0.004% of the stream's invertebrate standing crop was estimated to be in the water column at any instant in time. The possible use of the outgoing animals as food for juvenile salmon in the estuary is discussed.  相似文献   

20.
Highly specific environmental factors such as the presence of strongly mineralized water, oxygenation of the water down to 10 m, a low trophic status of the water and low organic matter content in the bottom sediments, were found in the studied post-exploitation Lake Piaseczno. Of the 42 macroinvertebrate taxa found, Oligochaeta (Tubificidae) and Chironomidae predominated. Almost exclusively ubiquitous species were found, except for some chironomid taxa recorded for the first time in Poland. The most diversified faunal communities occurred at 1 m water depth whereas the highest densities were found at 5 m. At the deepest points studied only oligochaetes and Chaoborus flavicans were found. There was no correlation between organic matter content and macrofauna densities. Differences in inhabitation at the studied transects could result from various environmental factors such as the degree of macrophyte cover and protection from the wind. Undisturbed transects overgrown by macrophytes had a similar community composition and higher faunal densities than those which were less protected.  相似文献   

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