Multi-scale analysis on wintering habitat selection of Reeves's Pheasant (Syrmaticus reevesii) in Dongzhai National Nature Reserve, Henan Province, China

Reeves's Pheasant (Syrmaticus reevesii), endemic to China, is an endangered species of pheasants. The wintering habitat selection by the species was investigated at three scales (10, 115 m and 250 m) in Dongzhai National Nature Reserve from 2000 to 2002. At each scale, a range of habitat variables were compared between the used and the control sites. At the smallest scale (10 m), the variables influencing wintering habitat selection were slope, tree cover, and the interaction between the cosine of slope aspect and the shrub height. At the mid-scale (115 m), the area of shrub, the area of broad-leaved forest, and the area of conifer forest were the key factors. At the largest scale (250 m), broad-leaved and conifer forest coverages and their interaction were the key factors. According to the lowest AIC and AIC_c values at the mid-scale, the characteristics at this scale were stated as the ultimate factors influencing the habitat selection of the bird. When a range of habitat variables at all scales within a multivariate regression were considerred, the most important variables were conifer forest coverage at the mid-scale, broad-leaved forest coverage, and the interaction between the conifer forest and shrub coverages at the large-scale, and the distance to beach and farmland. These results highlight the importance of multiscale analyses when habitat selection by pheasants are considerred.     

江西官山自然保护区四种雉类的生境选择差异

2009年10月—2010年11月在江西官山国家级自然保护区对白颈长尾雉、白鹇、勺鸡和灰胸竹鸡4种雉类的生境选择进行了研究:共调查了6条样线上的388个样方,选取了与这四种雉类栖息地相关的17个生态因子进行观测和测量。结果显示,4种雉类均偏好阔叶林和针阔混交林、阳坡或半阴半阳坡的生境。Kruskal-Wallistest检验表明,4种雉类在海拔、坡度、乔木盖度、灌木盖度、灌木数量、灌木高度、草本盖度、草本种类、草本数量、落叶层盖度和水源距离11个生态因子上存在极显著差异(P<0.01),乔木种类存在显著差异(P<0.05),其余生态因子无显著差异(P>0.05)。典则判别函数图显示,4种雉类在生境选择上存在一定程度的重叠,又有比较明显的差异。逐步判别表明在区分4种雉类生境选择差异方面有一系列的生态因子发挥作用,依照贡献值的大小依次为海拔、草本数量、灌木高度、水源距离、灌木盖度、乔木盖度、坡度、落叶层盖度、灌木数量、草本盖度。由这10个变量构成的方程对4种雉类生境选择差异的正确区分率为74.7%。     

山西芦芽山褐马鸡越冬栖息地选择的多尺度研究

1998～2000年在山西芦芽山自然保护区对褐马鸡的越冬栖息地选择进行了研究。采用4种空间尺度(10m、100m、300m和距离尺度),对影响褐马鸡越冬栖息地选择的主要因子进行了深入分析,并建立了褐马鸡越冬栖息地选择的逻辑斯谛回归模型。在300m尺度上．活动点和非活动点的生境类型有针叶林、针阔混交林、灌木林和草丛等。活动点周围针叶林面积显著高于非活动点(F=-3．116,P=0．002),虽然针阔混交林在两者中的面积比例都较小,但活动点周围针阔混交林的面积明显地低于非活动点(F=-2．255,P=0．024)．在灌木林和草丛的面积上两者无显著差异。这表明褐马鸡在300m尺度上喜欢活动于针叶林较多的地域,由于冬季针阔混交林不如针叶林能提供很好的隐蔽条件,褐马鸡避免选择针阔混交林;在100m尺度上,活动点和非活动点的生境类型有针叶林、针阔混交林和草丛,无灌木林生境,活动点的针叶林面积明显地高于非活动点(F=-2．931,P=0．003)。这表明褐马鸡在100m尺度上虽然倾向于选择针叶林,但对其它类型的生境如针阔混交林和草丛是可以利用的,这可能与其广泛取食活动有关。褐马鸡大尺度上的隐蔽条件满足以后,在小尺度上主要是为了获取更为丰富的食物。在距离尺度上活动点距居民点的距离、距道路的距离显著大于非活动点(F=15．621;6．048,P=0．000;0．018)。通过逐步逻辑斯谛回归分析,发现距灌草丛的距离、距居民点的距离、100m范围内针叶林的面积、树高以及食物的丰盛度是冬季褐马鸡栖息地选择的重要因子。以另外一个研究地收集的数据对所建立的栖息地选择模型的可靠程度进行了检验,结果表明该模型能有效地对褐马鸡的越冬栖息地进行预测。     

大熊猫的生境选择特征

基于王朗国家级自然保护区1997—2009年的连续监测数据,利用分布频率法和Bailey法,从地形因子、森林群落结构和主食竹3个方面研究了大熊猫的生境选择特征.结果表明：王朗国家级自然保护区的大熊猫对生境具有明显的选择性.在地形上,多选择海拔在2500～3000 m的山体脊部、上部和中部的均匀坡和凸坡,坡向西南,坡度在6°～30 °,与水源距离＞300 m的环境;森林群落结构上,多选择起源为次生林、针阔混交林,微生境为竹林的生境,乔木平均高度在20～29 m,灌木盖度在0～24%;主食竹多选择平均高度在2～5 m,竹丛盖度＞50%,混生,生长状况良好的缺苞箭竹.     

大熊猫的生境选择特征

基于王朗国家级自然保护区1997—2009年的连续监测数据,利用分布频率法和Bai-ley法,从地形因子、森林群落结构和主食竹3个方面研究了大熊猫的生境选择特征.结果表明:王朗国家级自然保护区的大熊猫对生境具有明显的选择性.在地形上,多选择海拔在2500～3000 m的山体脊部、上部和中部的均匀坡和凸坡,坡向西南,坡度在6°～30°,与水源距离>300 m的环境;森林群落结构上,多选择起源为次生林、针阔混交林,微生境为竹林的生境,乔木平均高度在20～29 m,灌木盖度在0～24%;主食竹多选择平均高度在2～5 m,竹丛盖度>50%,混生,生长状况良好的缺苞箭竹.     

浙江天童地区常绿阔叶林退化对土壤养分库和碳库的影响

为了解常绿阔叶林退化对土壤碳库和养分库的影响,采用空间代替时间的研究方法,以常绿阔叶林顶级群落为参照,选择了次生常绿阔叶幼年林、次生针阔混交林、次生针叶林、灌丛和灌草丛代表不同的退化类型,分别对其土壤氮磷养分库、碳库进行了调查和分析。结果表明：土壤氮库贮量从大到小依次为,成熟常绿阔叶林、次生常绿阔叶幼年林、灌丛、次生针叶林、灌草丛和次生针阔混交林;土壤总磷含量也是在成熟林最高,次生针阔混交林和次生针叶林的总磷含量显著高于次生常绿阔叶幼年林和灌丛;土壤有机碳含量从高到低依次为：成熟常绿阔叶林,次生针叶林、次生常绿阔叶幼年林、灌丛、灌草丛和次生针阔混交林;土壤铵态氮在成熟林、灌丛和灌草丛的库容量最大,其次分别为次生幼年常绿阔叶林、次生针阔混交林,最小的为次生针叶林;硝态氮则在灌草丛的库容量最大,其次分别为次生针叶林、次生针阔混交林和成熟林针叶林,最小的为次生常绿阔叶幼年林和灌丛。统计显示,常绿阔叶林退化不仅导致土壤有机碳库含量的显著下降,也使得土壤氮磷养分库含量显著下降。可以认为,砍伐导致的大量生物量输出和森林管理措施的影响,植物种类组成的改变,土壤物理性质的改变以及养分和有机碳的主要生物化学转化环节发生改变是导致此类变化的主要因素,常绿阔叶林顶极群落土壤是该地区土壤的最大养分库和碳库。     

河南董寨白冠长尾雉繁殖期栖息地选择

2001年至2003年春季,采用样线调查和媒鸟招引,在河南董寨国家级自然保护区对我国特有珍稀雉类白冠长尾雉（Syrmaticus reevesii）的栖息地选择进行了调查,结合RS和GIS分析了在景观水平上对栖息地的选择性,并借助逐步逻辑斯谛回归分析了影响繁殖期白冠长尾雉栖息地的关键尺度和主要因素。结果表明,在白云保护站,占区雄性白冠长尾雉在不同栖息地类型中的出现频率不同,出现最多的是混交林,其次是杉木林,随后是松林、灌丛、阔叶林;在董寨自然保护区内,在115 m尺度和250 m尺度上,针叶林的面积比例均是影响其栖息地选择的关键因子,而到农田的距离是距离因素中最重要的因素。根据回归分析和AIC_C及ΔAIC_C值,115 m尺度上栖息地变量对白冠长尾雉繁殖期的栖息地选择影响最大。综合分析表明,影响白冠长尾雉繁殖期栖息地选择的主要因子为115 m尺度上针叶林的面积比例和到农田的距离。建议在制定白冠长尾雉栖息地保护策略时,应加强现有适宜栖息地的管理,改善栖息地布局,并从景观尺度上开展针叶林对白冠长尾雉种群影响方面的研究工作。     

河南董寨国家级自然保护区发冠卷尾的巢址选择

2004年6~9月于河南董寨国家级自然保护区对发冠卷尾(Dicrurus hottentottus)的巢址选择进行了研究。野外调查中共发现发冠卷尾的巢47个,分布于针叶林、阔叶林和针阔混交林等3种植被类型中。我们在巢址样方(n=47)和对照样方(n=47)中对地形、距离与植被等因素(共15个变量)进行了调查,最终通过逐步判别分析表明,距山脊距离、距水源距离、乔木均高和灌木密度是影响其巢址选择的主要因子。     

广西黑颈长尾雉对夜宿地的选择

2003年12月、2004年5月和12月、2005年4月,通过野外调查及设置样方的方法对广西金钟山鸟类保护区黑颈长尾雉的夜宿地选择进行了研究。共记录到12个夜宿地,以夜宿树为中心做一个10m×10m和5个1m×1m的样方,记录夜宿树特征(种类、胸径、栖枝高度、栖枝直径、栖枝上方盖度),测量每个样方的11个生境因子(海拔、林型、坡向、坡度、乔木层盖度、灌木层盖度、草本层盖度、距水源距离、距林缘距离、距道路距离和乔木密度);并随机设10个对照点,测量相同的生境因子。结果表明,黑颈长尾雉夜宿地主要位于阔叶林中,一般,夜间在乔木上休息,一树栖息一只,也有2—3只同宿一树。黑颈长尾雉选择有一定坡度、林下灌木和草本较少以及在森林内部远离林缘并且乔木密度较大的地方作为夜宿地,其夜栖树主要为生境内的中等大小的阔叶树,栖枝直径约为2·5—4cm,栖处高度为2—5m,黑颈长尾雉夜宿地的选择与距水源距离的远近无关。影响黑颈长尾雉夜宿地选择的因素依次为:位置及林下植被、栖枝、天气。影响黑颈长尾雉夜宿地选择的最终原因可能是安全、舒适和栖息地转换的方便程度。     

黄泥河自然保护区狍冬季栖息地选择

2009年12月—2010年1月,在黄泥河自然保护区采用样线法对狍冬季栖息地选择进行了研究。共设置了47条样线,调查了109个狍利用样方和109个对照样方,测定了海拔、坡度、坡向、坡位、隐蔽级、郁闭度、食物丰富度、灌丛盖度、树胸径、人为干扰距离、林型、裸岩、雪深、倒木和针叶树15类栖息地因子。Baileys法和Mann-WhiteyU检验表明:在黄泥河自然保护区冬季,狍喜欢在坡度平缓、平均海拔602m,位于阳坡或半阴半阳坡的中坡位上雪被较浅,食物丰富度、灌丛盖度和隐蔽级都较高的针阔混交林生境中活动,对倒木因子表现出明显的选择性,避开选择阴坡、裸岩和针叶林生境。在主成分分析中,前5个主成分中的累积贡献率达到88.6%,第一主成分贡献率为30.8%,主要反映利用样方的林型、海拔和隐蔽级的影响,第二主成分贡献率为26.5%,主要反映利用样方的食物丰富度和灌丛盖度影响。     

Habitat preferences of medium/large mammals in human disturbed forests in Central Japan

It is a great concern whether human disturbed forest, i.e., secondary forest or monoculture forest, can be a habitat for the wildlife. We examined habitat preference of medium/large terrestrial mammal species in a human-disturbed mountainous settlement area of Akiyama Region, Central Japan, by using camera-traps, which were placed at conifer plantation forest, high/low-disturbed deciduous broad-leaved forest, and broad-leaved/conifer mixed forest in 2008 and 2009. Camera-traps were operational for 4568.6 trap-nights producing 740 photo-captures of 13 medium/large terrestrial mammal species. Japanese serow and Japanese hare dominated 54 % of all photo-captures. Low-disturbed deciduous broad-leaved forest showed the richest mammal fauna (12 spp.). We suggested that fruit trees and understory vegetation provided suitable habitat for frugivorous as well as herbivorous mammals. The mixed forest showed the lowest mammal fauna (6 spp.), which located higher elevation. High-disturbed deciduous broad-leaved forest (9 spp.) was also supposed to be a better habitat for some frugivorous mammals and herbivorous mammals, but tended to be avoided by Japanese marten and Japanese macaque. Conifer plantation forest (7 spp.) with understory vegetation was supposed to be a better habitat for Japanese serow and Japanese hare, but not for the other species. Without fruit trees, conifer plantation forest was supposed to be non-attractive habitat for frugivorous mammals. We demonstrated that, if a forest had been disturbed by human, the forest can be a habitat for terrestrial herbivores, but this was not always true for frugivores.     

Habitat selection and movement ecology of eastern Hermann’s tortoises in a rural Romanian landscape

Understanding wildlife movements and habitat selection are critical to drafting conservation and management plans. We studied a population of eastern Hermann’s tortoise (Testudo hermanni boettgeri) in a traditionally managed rural landscape in Romania, near the northern edge of the species geographic distribution. We used telemetry to radio-track 24 individuals between 2005 and 2008 and performed a Euclidian distance-based habitat selection analysis to investigate habitats preferred by tortoises at both landscapes (second-order order selection) and individual (third-order selection) home range scales. The home range size for tortoises in our study area was 3.79?±?0.62 ha and did not differ by gender or season (pre- and post-nesting seasons). Their movement ecology was characterized by short-distance movements (daily mean?=?31.18?±?1.59 m), apparently unaffected by habitat type. In contrast to other studies, movements of males and females were of similar magnitude. At the landscape (population home range) scale, grasslands and shrubs were preferred, but tortoises also showed affinity to forest edges. At the individual home range scale, tortoises selected grassland and shrub habitats, avoided forests, and used forest edges randomly. Creeks were avoided at both spatial scales. Our results suggest that tortoise home ranges contain well-defined associations of habitats despite a higher selection for grasslands. As such, avoiding land conversion to other uses and maintaining habitat heterogeneity through traditional practices (e.g., manual mowing of grasslands, livestock grazing) are critical for the persistence of tortoise populations.     

Environmental variation and moose Alces alces density as determinants of spatio-temporal heterogeneity in browsing

Understanding temporal variation in habitat selection and browsing intensity by large herbivores is fundamental because of their large impact on the ecosystems. I studied the annual variation in winter browsing pressure on young trees and habitat selection by moose Alces alces over a ten year period. Specifically, the relationships between browsing pressure on Scots pine Pinus sylvestris and two birch species ( Betula ssp.) and three explanatory variables – 1) availability of forage, 2) moose density (estimated by pellet group counts) and 3) snow cover was studied. At a larger spatial scale (forest stand level) the relationship between moose habitat selection between three different habitat types (forest <30 yr, forest>30 yr and mire) and two explanatory variables, 1) snow condition and 2) moose density, were studied. Browsing pressure on Scots pine, the dominating food plant, was related to forage availability, moose density and snow condition. No significant relationships between any of the three explanatory variables and browsing pressure on the two birch species were found. Moose selection for certain habitats varied between years and was affected by number of days with >0.10 m of snow.
Habitat selection was not significantly related to moose density and the relationship between overall moose density and habitat specific moose densities was proportional within the studied density range. These findings have implications for understanding varying browsing patterns – and will affect both the ability to predict herbivores' effect on the forest ecosystem. A snow dependent browsing pattern also indicates that one can expect a long term decrease in browsing pressure on the tree and shrub layer as a consequence of the ongoing large-scale climate change.     

广西金钟山自然保护区主要植被类型的特征

金钟山自然保护区计有种子植物101科273属514种,落叶栎林分布面积最广。随海拔升高,植被依次呈现出4个分布带:沟谷落叶阔叶林、沟谷常绿阔叶林、常绿落叶阔叶混交林和落叶阔叶林、山地苔藓矮林和山地常绿阔叶林。该区主要有4个分布区类型:世界分布、热带分布、温带分布和中国特有分布,其中热带分布占总属数的75.21%,表明本保护区的植物分布具有热带性质。其天然植被类型可划分为5个植被型组,7个植被型含暖性针叶林、暖性落叶阔叶林、常绿落叶阔叶混交林、常绿阔叶林、竹林和草丛,2个植被亚型含南亚热带山地常绿阔叶林、山顶阔叶矮林,以及33个群系。     

Measures of giant panda habitat selection across multiple spatial scales for species conservation

Examining ecological processes across spatial scales is crucial as animals select and use resources at different scales. We carried out field surveys in September 2005, March–September 2006, and April 2007, and used ecological niche factor analysis to determine habitat preferences for the giant panda (Ailuropoda melanoleuca) across 4 spatial scales: daily movement, core range, home range, and seasonal elevational migration. We found that giant pandas prefer conifer forest and mixed forest at higher than average elevation (2,157 m) of study area in the 4 scale models. However, we also observed significant scale differences in habitat selection. The strength of habitat preference increased with scale for the 2 disturbed forests (sparse forest and fragmented forest), and decreased with scale for 0–30° gentle slope and south- and north-facing aspect. Furthermore, habitat suitability patterns were scale-dependent. These findings highlight the need to determine species–environment associations across multiple scales for habitat management and species conservation. © 2012 The Wildlife Society.     

凉水自然保护区松鼠和星鸦贮食生境选择差异

2003 年9 月30 日～2005 年10 月8 日,在黑龙江凉水国家级自然保护区,应用样方调查法, 采用Vanderploeg和Scavia 选择系数Wi和选择指数Ei作为衡量指标,对松鼠和星鸦贮食生境选择进行了研究。结果表明, 二者贮食生境选择优先顺序略有不同,松鼠偏爱的贮食生境依次是: 云杉林、原始红松林、人工红松林、针阔混交林、人工云杉林、白桦林、针叶混交林、人工落叶松林、阔叶混交林、冷杉林和其它。星鸦对贮食生境的偏爱程度依次为: 人工红松林、原始红松林、云杉林、人工云杉林、针阔混交林、阔叶混交林、白桦林、针叶混交林、人工落叶松林、冷杉林和其它。在对贮食微生境因子的选择利用上, 二者大致相同,只是在对优势灌丛的选择上略有差异, 松鼠优先选择在狗枣猕猴桃优势灌丛内贮藏红松种子, 而星鸦优先选择在刺五加优势灌丛内贮食。松鼠和星鸦贮食生境选择的差异将对随后的红松天然更新过程产生不同的影响。     

官山保护区白颈长尾雉栖息地适宜性评价

2009年4月-2009年5月和2009年12月-2010年1月,在江西官山国家级自然保护区对白颈长尾雉(Syrmaticus ellioti) 繁殖期和越冬期的栖息地进行调查。通过实地调查,结合已有研究结果,确定了官山自然保护区白颈长尾雉栖息地适宜性评价因子,即植被类型、海拔、坡度和坡向等4个栖息地因子。基于"被利用生境-可获得生境比较法",以对各评价因子不同梯度的资源选择指数为依据,确定了白颈长尾雉适宜性评价准则。应用ArcGIS 9.3空间分析功能和栖息地片断化指数,对官山保护区白颈长尾雉栖息地进行了适宜性评价。结果表明:在官山自然保护区适宜白颈长尾雉繁殖的栖息地斑块有502个,总面积为3307 hm²,占保护区总面积的28.6%,分离度指数和片断化程度指数分别为0.365和0.912;适宜白颈长尾雉越冬的栖息地斑块有514个,总面积2755 hm²,占保护区总面积的23.8%,分离度指数和片断化程度指数分别为0.443和0.939。官山保护区白颈长尾雉栖息地片断化较严重,片断化现象在白颈长尾雉越冬期加剧。因此,迫切需要采取措施白颈长尾雉现存栖息地进行保护,特别是要高度重视其越冬栖息地的保护。     

Modeling spring habitat requirements of the endangered brown eared pheasant Crossoptilon mantchuricum in the Huanglong Mountains, Shaanxi Province, China

A detailed understanding of the habitat needs of brown eared pheasants (Crossoptilon mantchuricum) is essential for conserving the species. We carried out field surveys in the Huanglong Mountains of Shaanxi Province, China, from March to June in 2007 and 2008. We arrayed a total of 206 grid plots (200 × 200 m) along transects in 2007 and 2008 and quantified a suite of environmental variables for each one. In the optimal logistic regression model, the most important variables for brown eared pheasants were slope degree, tree cover, distance to nearest water, cover and depth of fallen leaves. Hosmer and Leweshow goodness-of-fit tests explained that logistic models for the species were good fits. The model suggested that spring habitat selection of the brown eared pheasant was negatively related to distance to nearest water and slope degree, and positively to cover of trees and cover and depth of fallen leaves. In addition, the observed detected and undetected grids in 2007 did not show significant differences with predictions based on the model. These results showed that the model could well predict the habitat selection of brown eared pheasants. Based on these predictive models, we suggest that habitat management plans incorporating this new information can now focus more effectively on restrictions on the number of tourists entering the nature reserve, prohibition of firewood collection, livestock grazing, and medicinal plant harvesting by local residents in the core areas, protection of mixed forest and sources of the permanent water in the reserve, and use of alternatives to firewood.     

长白山不同演替阶段针阔混交林群落物种多度分布格局

为解释长白山温带森林群落构建和物种多度格局的形成过程, 该文以不同演替阶段的针阔混交林监测样地数据为基础, 采用中性理论模型、生物统计模型(对数正态分布模型)和生态位模型(Zifp模型、分割线段模型、生态位优先模型)拟合森林群落物种多度分布, 并用χ ²检验、Kolmogorov-Smirnov (K-S)检验和赤池信息准则(AIC)选择最佳拟合模型。结果显示: 中性模型能很好地预测长白山温带森林不同演替阶段植物群落的物种多度分布。在10 m × 10 m尺度上, 5种模型均可被χ ²检验和K-S检验接受, 但中性模型拟合效果不如对数正态分布模型、Zifp模型、分割线段模型和生态位优先模型, 表明小尺度上中性过程和生态位过程均能解释群落物种多度分布, 但生态位过程的解释能力相对较大。而在中大尺度上(30 m × 30 m、60 m × 60 m和90 m × 90 m), 中性模型为最优拟合模型, 并且随着研究尺度增加, 生态位模型和生物统计模型逐渐被χ ²检验拒绝, 表明中性过程在长白山针阔混交林群落物种多度分布格局形成中的作用随着研究尺度增加而逐渐增大。该文证实了中性过程在长白山温带针阔混交林群落结构形成中具有重要作用, 但未否认生态位机制在群落构建中的贡献。因此, 温带森林群落构建过程中中性理论和生态位理论并非相互矛盾, 而是相互融合的。在研究森林群落物种多度分布时, 应重视取样尺度和演替阶段的影响, 并采用多种模型进行拟合。     

青藏高原东缘高寒森林小型兽类地表洞穴选择与生境特征

高寒森林地表的小型兽类洞穴是生态系统重要的景观特征,对于认识小型兽类的分布格局及其生态功能具有重要意义.为此,以青藏高原东缘典型高寒森林生态系统小型兽类洞穴为对象,于2015年夏季(7—9月)研究了其地表分布与生境特征.结果表明: 小型兽类洞穴平均洞口密度为182.8个·hm^-2,不同森林植被类型显著影响洞穴密度,表现为针叶林(328.6个·hm^-2)>针阔混交林(160.0个·hm^-2)>灌木林(125.0个·hm^-2)>阔叶林(81.5个·hm^-2)>竹林(66.7个·hm^-2). Mann-Whitney U Test和主成分分析表明,小型兽类栖息样地的乔木郁闭度、乔木均高、乔木胸径、乔木密度、地表植被盖度、倒木密度、倒木覆盖度、灌木均高都显著大于非栖息样地;乔木因子是决定小型兽类生境选择的首要因子,贡献率最大(44.8%),其次是倒木因子(23.2%)和灌草丛因子(13.9%).说明高寒森林地表有明显的小型兽类洞穴分布,其分布格局主要受植被和倒木的影响.