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1.
  总被引:1,自引:0,他引:1  
We review the fossil history of seed plant genera that are now endemic to eastern Asia. Although the majority of eastern Asian endemic genera have no known fossil record at all, 54 genera, or about 9%, are reliably known from the fossil record. Most of these are woody (with two exceptions), and most are today either broadly East Asian, or more specifically confined to Sino-Japanese subcategory rather than being endemic to the Sino-Himalayan area. Of the "eastern Asian endemic" genera so far known from the fossil record, the majority formerly occurred in Europe and/or North America, indicating that eastern Asia served as a late Tertiary or Quaternary refugium for taxa. Hence, many of these genera may have originated in other parts of the Northern Hemisphere and expanded their ranges across continents and former sea barriers when tectonic and climatic conditions allowed, leading to their arrival in eastern Asia. Although clear evidence for paleoendemism is provided by the gymnosperms Amentotaxus, Cathaya, Cephalotaxus, Cunninghamia, Cryptomeria, Glyptostrobus, Ginkgo, Keteleeria, Metasequoia, Nothotsuga, Pseudolarix, Sciadopitys, and Taiwania, and the angiosperms Cercidiphyllum, Choerospondias, Corylopsis, Craigia, Cyclocarya, Davidia, Dipelta, Decaisnea, Diplopanax, Dipteronia, Emmenopterys, Eucommia, Euscaphis, Hemiptelea, Hovenia, Koelreuteria, Paulownia, Phellodendron, Platycarya, Pteroceltis, Rehderodendron, Sargentodoxa, Schizophragma, Sinomenium, Tapiscia, Tetracentron, Toricellia, Trapella, and Trochodendron, we cannot rule out the possibility that neoendemism plays an important role especially for herbaceous taxa in the present-day flora of Asia, particularly in the Sino-Himalayan region. In addition to reviewing paleobotanical occurrences from the literature, we document newly recognized fossil occurrences that expand the geographic and stratigraphic ranges previously known for Dipelta, Pteroceltis, and Toricellia.  相似文献   

2.
Fossil wood is subject to different taphonomic, sampling and recognition biases in the palaeobotanical record when compared with leaves and palynomorphs. Wood therefore provides a systematically independent source of information that can increase our knowledge of past biodiversity and environments. Increase in fossil wood records from Cretaceous and Tertiary sediments helps further the understanding of trends in anatomical specialization through geological time. These data can then be used to distinguish such specialization from anatomical response to environmental change. Two case studies, a Late Cretaceous early Tertian' wood flora from Antarctica and a lower Tertiary w ood flora from southern England, have been used to exemplify the importance of studying the fossil wood component of palaeofloras.  相似文献   

3.
    
Cedrelospermum Saporta is an extinct genus in the Ulmaceae with abundant fossil records in North America and Europe. However, so far, fossil records of this genus from Asia are sparse, which limits the interpretations of the morphological evolution and biogeographical history of the genus. Here we report well‐preserved fruits (Cedrelospermum tibeticum sp. nov.) and a leaf (Cedrelospermum sp.) of Cedrelospermum from the upper Oligocene Lunpola and Nyima basins in the Qinghai–Tibetan Plateau (QTP). This is the first fossil record of Cedrelospermum in the QTP, showing that this genus grew in this region during the late Oligocene. Cedrelospermum tibeticum fruits are double‐winged, morphologically similar to the Eocene and Oligocene double‐winged Cedrelospermum species from North America. This supports the hypothesis that Cedrelospermum migrated to Asia from North America by way of the Bering Land Bridge. Given that Cedrelospermum was a typical element of Northern Hemispheric flora in the Paleogene and Neogene, the presence of this genus indicates that the central region of the QTP was phytogeographically linked with other parts of the Northern Hemisphere during the late Oligocene. The morphological observations of C. tibeticum fruits and other double‐winged Cedrelospermum fruits suggest an evolutionary trend from obtuse to acute apex for the primary wing. Cedrelospermum tibeticum likely had warm and wet climatic requirements. This type of an environment possibly existed in the central QTP in the late Oligocene, thereby supporting the survival of C. tibeticum.  相似文献   

4.
Sauropod dinosaurs are poorly represented in the Lower Cretaceous of eastern Asia. Here, we describe a number of isolated sauropod teeth from the Kuwajima Formation (?Berriasian–?Hauterivian) of Shiramine, Japan. The mosaic of shared derived characters and symplesiomorphies displayed by the teeth indicate that they are referable to a basal member of the titanosauriform radiation. A taxonomic review of previously described sauropod specimens from eastern and south–eastern Asia reveals that a diversity of sauropods (including a titanosaurian, a basal titanosauriform and a ?euhelopodid, as well as several forms of indeterminate systematic position) was present in this region in the Early Cretaceous. This diversity conflicts with previous suggestions that eastern Asia was biogeographically isolated from the rest of Laurasia until the late Early Cretaceous and that the sauropod fauna was limited to the endemic East Asian clade Euhelopodidae. The presence of titanosauriform sauropods in the basal Cretaceous of Japan and Thailand indicate that the proposed faunal isolation of eastern Asia ended approximately 20 myr earlier than usually suggested.  相似文献   

5.
    
Fossil records of endemic plants play an important role in recognizing the floristic history of East Asia and thereby facilitate the conservation of plant diversity in the region. However, the fossil record of many extant East Asian endemic genera remains poorly documented thus far. Here, we report an infructescence fossil of an East Asian endemic genus, Sladenia (Sladeniaceae), from the early Miocene of southeastern Yunnan, China. The fossil is characterized by: (i) dichasial cymes; and (ii) flask‐shaped ovary with dense subparallel ribs on the surface extending from the base to the distal end of the united style. It represents the first fossil record of Sladenia in Asia, showing that the genus was established in the region at least by the early Miocene. Given that a much older fossil record of Sladeniaceae has been reported from Africa and the sister group of Sladenia is distributed only in Africa, Sladenia is not likely of East Asian origin. The present endemic status of Sladenia was possibly achieved by regional extirpation in Africa and taking refuge in East Asia. This case thus supports the “Museum” rather than “Cradle” hypothesis for the genesis of high plant species in the flora of East Asia. A comparison of the present fossil with extant Sladenia infructescence shows morphological stasis from the early Miocene to present. Such evolutionary tardiness might have resulted in the reduced fitness of the genus, which further caused its current endangered situation.  相似文献   

6.
  总被引:4,自引:0,他引:4  
Specimens of Metasequoia from the Late Cretaceous or Early Tertiary collected from the Wuyun Formation, Jiayin County, Heilongjiang Province, north-east China, include shoots, female and male cones. Vegetative and reproductive features as well as details of leaf cuticle are described. Comparative studies on fossil and living specimens of the genus suggest a new approach to the classification of species of Metasequoia. Our investigations indicate that extant M. glyptostroboides is distinct from all of the recognized fossil species. The specimens described here, in addition to those previously regarded as M. disticha, M. japonica, M. chinensis, M. cuneata, M. papillapollenites, M. miocenica, M. kimurae, M. onukii, M. europaea, M. asiatica and M. nathorstii , are now considered to belong to a single species, namely M. occidentalis. A further valid species is M. milleri , which can be distinguished from M. occidentalis by the morphology of the male cones and the vegetative organs.  相似文献   

7.
    
The new discovery of angiosperm remains in the Jehol Biota of northeastern China contributes to our understanding of the origin and early evolution of flowering plants. The earliest eudicot genus with reproductive organs, Leefructus, was recently documented from the Lower Cretaceous Yixian Formation at 125.8–123.0 Ma, and was reconsidered to be close to the extant family Ranunculaceae based on gross morphology. However, this hypothesis has not been tested using a cladistic approach. To determine the possible allies of Leefructus within extant eudicots, we constructed a 66 morphological data matrix. Molecular and morphological analyses of extant Ranunculales combined with the fossil suggest that it has an affinity with the Ranunculaceae. The earliest fossil record of the eudicots is 127–125 Ma based on tricolpate pollen grains. Thus, we suggest a hypothesis that the basal eudicots might have experienced an accelerated evolution and diversification during the latest Barremian and earliest Aptian, leading to the stem groups of at least six extant families or lineages, 10–15 Myr earlier than currently documented. Angiosperms have undergone multiple uneven pulses of radiation since their origin. Many key character innovations occurred in different stages that could have triggered those radiations in concert with various biotic and abiotic factors.  相似文献   

8.
    
Reproductive and vegetative remains of Eucommia from 25 localities in the United States, Canada, and Mexico document the wide distribution of this genus in North America during the Cenozoic. Autofluorescent elastic latex filaments bearing capitate termini are preserved in nearly all of the remains and provide conclusive evidence of their affinity to Eucommia. Four species of Eucommia are recognized on the basis of the characteristic samaras: E. eocenica from middle Eocene strata of the Mississippi Embayment in Missouri, Tennessee and Mississippi; E. montana from early Eocene to early Oligocene localities in British Columbia, Washington, Oregon, Utah, Colorado, and Montana; E. constans from Neogene rocks in central Mexico; and E. jeffersonensis n. sp. from the latest Eocene or earliest Oligocene John Day Formation of Oregon. Atypical specimens of E. montana and E. eocenica are the first records of two-seeded fruits for the genus. Eucommia leaves from Eocene localities in British Columbia and Mississippi are the first records of Eucommia foliage in North America whose identifications are confirmed by the presence of capitate latex strands. These leaves and a specimen from Oregon are referred to E. rolandii n. sp. Fruit evolution in Eucommia may have involved increases in samara size and symmetry, and reduction in seed number from two to one, perhaps as adaptations for wind dispersal. All fossil Eucommia samaras from North America are smaller and less symmetrical than those of the living species, E. ulmoides. Preliminary flight tests of E. ulmoides samaras and of models of the fossils suggest that E. ulmoides fruits are aerodynamically better suited for wind dispersal than the fossils.  相似文献   

9.
10.
A fossil flower with affinities to the modern families of the saxifragalean complex is described. Fossils were collected at Old Crossman Pit, Raritan Formation, New Jersey, USA. These sediments are dated on the basis of palynology as Turonian (Upper Cretaceous, ~90 million years before present). Fossils are charcoalified and preserved with exceptional three- dimensional detail. The characters observed in these flowers, when compared with those of extant flowers of several families of the saxifragalean complex, suggest a close relationship with extant members of the Saxifragaceae and Hydrangeaceae. Hypotheses on the origin of petals and staminodes and a possible mechanism of pollination are discussed. This new taxon provides additional characters in the floral morphology of the fossil saxifragoids and extends their geographical distribution in the Cretaceous to North America.  相似文献   

11.
12.
Phylogenetic relationships within the genus Pieris (Ericaceae) were investigated based on the rbcL and matK genes along with five spacer sequences of chloroplast DNA to address questions regarding the phylogeography of the genus in association with insular plants on the Ryukyu Islands. The most parsimonious trees indicated that P. floribunda from eastern North America is a sister taxon to the remaining taxa examined, and suggested that the East Asian taxa examined are monophyletic. A morphologically cohesive group, section Pieris, was revealed to be paraphyletic. Within the East Asian clade, insular endemics from the Ryukyu Islands, Taiwan, and mainland Japan formed a sister group to P. formosa from the Himalayas and southern China. Our data suggest that the insular endemics of the Ryukyu Islands and Taiwan arose via allopatric divergence as a result of a paleogeographical land configuration of a landbridge during the early–middle Pleistocene in the Quaternary Period.  相似文献   

13.
1 全球气候变化研究新动向自 2 0世纪 80年代以来 ,全球气候变化研究至今仍方兴未艾。全球气温波动、大气中温室气体丰度变化、气候格局改变、气候带迁移和降水量增减 ,都影响到社会经济的发展和人类生活质量的提高。各国政府对全球气候变化的研究 ,如气候演变的过程、短期和长期气候变化的预测和自然灾害的防治都给予极大关注和财力支持[1] 。目前全球气候变化的研究正向两个方向发展。1.1 加大全球气候变化研究时间尺度 ,从研究当今气候发展到研究地质历史时期气候变化全球气候变化 ,如大气中CO2 浓度的变化、温度的升降和降水量的变…  相似文献   

14.
Recent discoveries of fossil reproductive structures from deposits of the Raritan Formation in New Jersey (Turonian, Upper Cretaceous, ~90 million years BP) include a previously undescribed representative of the Order Capparales. The fossils are usually charcoalified with three-dimensional structure and excellent anatomical details. In the present contribution, we introduce a taxon represented by fossil flowers that have a combination of characters now found in the families of the Order Capparales sensu Cronquist. The fossil species is characterized by an unique suite of characters, such as the presence of a gynophore, arrangement of the sepals, unequal petal size, monothecal anthers, and a bicarpellate gynoecium, that are found in extant families of the Order Capparales. This new taxon constitutes an important addition to our understanding of Cretaceous angiosperm diversity and represents the oldest known fossil record for the Capparales. Heretofore, the oldest known capparalean was from the Late Tertiary sediments of North America.  相似文献   

15.
    
There are many extant endemic plants in China, which were widely distributed in the North Hemisphere during Tertiary. The global cooling during the Tertiary caused a series of narrow distribution regions of the plants. Quaternary glaciation invaded most regions of North America and Eurasia where severe destruction was imposed onto vegetation. However, such destruction was lessened in China largely because of specific topographic and geographical and obviously, a number of other conditions accounted for an unusual refugee camp for the relics of plants in China, among which lots of endemic taxa exist. Recently, Chinese endemic species, such as Metaseqouia, Eucommia , have been employed to conduct multi-disciplinary comprehensive studies so as to analyze Tertiary climate changes quantitatively. Meanwhile, a rigorous method, i.e. climate analysis of endemic species (CAES) has come to maturation. This method is characteristic of some generality because it is supposed to be applicable to the endemic species in other regions of the world. CAES is involved in the following aspects: 1. Conduct multidisciplinary studies on living and fossil species of endemic plants and trace their evolutionary courses. 2. Compare fossil species with living one and clarify which is the nearest living relative (NLR) to fossil counterpart. 3. Fossils and their living counterparts (NLR) are supposed to have similar ecological requirements to meet their life cycles. 4. Investigate the geographic distribution of living and fossil plants within the same taxa and ascertain the dynamic changes of their distributions in geological age. 5. Analyze climate factors in the distribution of specific endemic taxa and obtain the data of climatic characters which are suitable for reconstruction of paleoclimate where fossil counterparts lived. 6. Further study the physio-ecology of living species and determinate paleoclimate where fossil counterparts lived. 7. Integrate analysis of the data from steps 4, 5 and 6, and quantitatively reconstruct the climate where fossil and living plants survive.  相似文献   

16.
    
The Turonian flora from Sayreville New Jersey includes one of the world's most diverse assemblages of Cretaceous angiosperm flowers. This flora is made even more interesting by its association with a large insect fauna that is preserved by charcoalification as well as in amber. Floral diversity includes numerous representatives of Magnoliidae, Hamamelididae, Rosidae, Dilleniidae, and Asteridae (Ericales sensu lato). Included are hypogynous, five-merous flowers with uniseriate hairs on the pedicels and stamens in bundles most frequently borne opposite the petals. There is considerable variation in filament length, and some filaments are branched. On some anthers, strands of residue, suggesting the former presence of a liquid of unknown nature, partially occlude the apparent zone of dehiscence. In other cases, open anthers are fully occluded by an amorphous substance. pollen is rarely found associated with anthers, but is common on stigmatic surfaces. pollen is prolate and tricolporate with reticulate micromorphology. The superior syncarpous ovary is five-carpellate with axile/intruded parietal placentation and numerous anatropous ovules/carpel. Ovary partitions have closely spaced, parallel ascending channels (secretory canals?), and there are apparent secretory canals/cavities in receptacles, sepals, and petals. Individual stigmas are cuneiform with a central groove and eccentrically peltate. Styles are short and fused. In aggregate, the stigmas form a secondarily peltate stigma. Seeds have a reticulate sculpture pattern, a pronounced raphe, and funicular arils with sculpture similar to the seeds. phylogenetic analyses of several data matrices of extant taxa place this fossil in a monophyletic group with the modern genera Garcinia and Clusia within the Clusiaceae. As such, these fossils represent the earliest fossil evidence of the family Clusiaceae. Some modern Clusiaceae are notable, in particular, for their close relationship with meliponine and other highly derived bee pollinators; the fossil flowers share several characters that suggest a similar mode of pollination. This possibility is consistent with other floral and insect data from the same locality.  相似文献   

17.
A multidisciplinary investigation of the eight annual taxa of Bromus sect. Genea throughout their total ranges has brought a fresh insight into their evolutionary pathways. These species, mainly in southwest Asia and the Mediterranean, are still actively in the process of speciation and their study illuminates evolutionary processes in annual grasses as a whole. Following a discussion of Genea features, a list of ancestral and derived characters in the group is presented. The recent emergence of a multiplicity of niches is probably the reason for the rapid emergence of many new kinds of plants in sect. Genea.  相似文献   

18.
    
Abstract

This study was conducted to identify areas of endemism for Acantholimon species using parsimony analysis of endemicity (PAE) and to detect endemic species richness of the genus in the region. The results obtained from the two methods used in this study were used in determining the priorities for the conservation of Acantholimon species in Iran. The distribution database of 62 endemic species belonging to this genus was formed by 1250 georeferenced observations in Iran. The study area was divided into 1?×1? grids of operative geographical units (OGUs) and the species?×?area matrix including presence/absence data was created. The endemic species richness was calculated using circular neighborhood with a radius of 50?km in 10?×?10?km2 raster cells using DIVA-GIS software. The results of PAE analysis have shown four areas of endemism (AOEs) in Iran. AOE1: including Alborz and Zagros mountains, the mountains of central Iran. AOE2 and AOE3 are located in Khorassan subregion and AOE4 contains parts of western Iran. The map of endemic species richness indicated that the highest number of endemic species occurs in central Alborz region as well as Kerman, Chahar-Mahal and Bakhtiari, and Isfahan provinces.  相似文献   

19.
    
Over the past 25 years, discoveries of Early Cretaceous fossil flowers, often associated with pollen and sometimes with vegetative parts, have revolutionized our understanding of the morphology and diversity of early angiosperms. However, few of these fossils have been integrated into the increasingly robust phylogeny of living angiosperms based primarily on molecular data. To remedy this situation, we have used a morphological data set for living basal angiosperms (including basal eudicots and monocots) to assess the most parsimonious positions of early angiosperm fossils on cladograms of Recent plants, using constraint trees that represent the current range of hypotheses on higher-level relationships, and concentrating on Magnoliidae (the clade including Magnoliales, Laurales, Canellales, and Piperales) and eudicots. In magnoliids, our results confirm proposed relationships of Archaeanthus (latest Albian?) to Magnoliaceae, Endressinia (late Aptian) to Magnoliales (the clade comprising Degeneria, Galbulimima, Eupomatia, and Annonaceae), and Walkeripollis pollen tetrads (late Barremian?) to Winteraceae, but they indicate that Mauldinia (early Cenomanian) was sister to both Lauraceae and Hernandiaceae rather than to Lauraceae alone. Among middle Albian to early Cenomanian eudicots, we confirm relationships of Nelumbites to Nelumbo, platanoid inflorescences and Sapindopsis to Platanaceae, and Spanomera to Buxaceae. With the possible exception of Archaeanthus, these fossils are apparently not crown group members of living families but rather stem relatives of one or more families.  相似文献   

20.
Rhododendron seeds from the Palaeocene of southern England   总被引:2,自引:0,他引:2  
Seeds closely resembling those of some species of the extant genus Rhododendron L. have been recovered from the Reading Beds near Newbury, Berkshire. The fossils are assigned to Rhododendron newburyanum sp. noV. on the basis of detailed comparisons with seeds of extant Rhododendron species, other Ericaceae, and taxa with seeds of similar morphology.  相似文献   

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