记陕北子长晚三叠世一新的古鳕类

记述了陕北子长晚三叠世古鳕科（Ｐａｌａｅｏｎｉｓｃｉｄａｅ）一新属新种──子长瓦窑堡鳕（Ｗａｙａｏｂｕｌｅｐｉｓ ｚｉｃｈａｎｇｅｎｓｉｓ ｇｅｎ． ｅｔ ｓｐ． ｎｏｖ．）。其一般形态特征如身体长纺锤形、背鳍基长,后部对着腹鳍和臀鳍之间的空隙、鳍条粗壮、饰缘棘鳞细小、悬挂骨倾斜、鳃盖骨很高大、鳞片小及横列鳞数目多等,与澳大利亚中至晚三叠世的Ｍｙｒｉｏｌｏｐｉｓ很相似,但新属具有吻较尖、背鳍鳍条数目较少、鳞片具有众多的斜嵴、后缘成梳齿状以及具有背嵴鳞等显著特征区别于Ｍｙｒｉｏｌｏｐｉｓ。瓦窑堡鳕在体形、鳍的位置、鳃盖骨高大及鳞片的纹饰等方面也与欧洲中至晚三叠世的 Ｇｙｒｏｌｅｐｉｓ相似,但新属以背鳍基较长、臀鳍基较短、背峪鳞发达及鳍条粗壮等特征区别于Ｇｙｒｏｌｅｐｉｓ。根据瓦窑堡鳕的性质和化石层位的情况,将合鱼化石地层的时代定为晚三叠世。     

云南墨江晚三叠世Burmesia一新种

云南墨江孟弄晚三叠世路马组中发现的墨江缅甸蛤（Ｂｕｒｍｅｓｉａｍｏｊｉａｎｇｅｎｓｉｓｓｐ．ｎｏｖ．）,以其壳面装饰前部的反Ｎ字形斜褶脊,同已发表的缅甸蛤各种相区别,从而丰富了缅甸蛤的内容。     

云南罗平中三叠世翼鳕属一新种及早期辐鳍鱼类系统发育关系（英文）

辐鳍鱼亚纲是现存脊椎动物中最大的类群,包括腕鳍鱼次亚纲、辐鳍鱼次亚纲(包括软骨硬鳞类和新鳍鱼类)和亲缘关系密切的化石类群。已灭绝的翼鳕属(Pteronisculus)是隶属于辐鳍鱼亚纲的一个干群,包括产于马达加斯加、欧洲和北美下三叠统的11个种和中国中三叠统的一个种。根据滇东罗平中三叠世(安尼期)海相地层中发现的5块保存完好的化石,命名翼鳕属一新种,张氏翼鳕(Pteronisculus changae sp. nov.)。这是翼鳕属在中三叠世的第二个确切种,最大体长达295 mm,代表了罗平生物群中已知体型最大的辐鳍鱼亚纲干群物种。新种具有翼鳕属的独特衍征,泪骨具有牙齿,但它又有明显区别于本属其他种的自近裔特征,如间颞骨中部有一个内突起,21根上神经骨,83列侧线鳞。分支分析结果为早期辐鳍鱼类系统发育关系提供了新的见解,认为翼鳕属是Cyranorhis的姐妹群。根据体型和口缘牙齿等特征推测张氏翼鳕是一个快速游动的捕食者,以浮游无脊椎动物和体型较小的鱼类或鱼类幼体为食。作为翼鳕属最年轻的成员之一,张氏翼鳕的发现进一步表明翼鳕的多样性比我们过去认识的要高,古特提斯洋东缘可能是该属在中三叠世早期的避难所。     

西藏沙蜥属一新种—泽当沙蜥

根据形态特征,泽当居群与西藏沙蜥、红尾沙蜥有显著区别,为沙蜥属一新种：泽当沙蜥Ｐｈｒｙｎｏｃｅｐｈａｌｕｓ ｚｅｔａｎｇｅｎｓｉｓ ｓｐ．ｎｏｖ．Ｗａｎｇ,Ｚｅｎｇ ｅｔ Ｗｕ鉴别特征为鼻鳞１枚;鼻间鳞４－５枚;背鳞数少于腹鳞数;雄性尾末端黑色,雌性灰白色。     

广西金线Ba属一新种

描记采于广西凌云县泗城镇沙洞地下河中的金线Ｂａ属１新种,以采集地县名命名为凌云金线ＢａＳｉｎｏｃｙｃｌｏｃｈｅｉｌｕｓ ｌｉｎｇｙｕｎｅｎｓｉｓ ｓｐ．ｎｏｖ．。新种以侧线鳞７１－７８对４８－５０;背鳍最末不分枝鳍条下半部为硬刺,后缘有锯齿对纤细柔软,光滑无锯齿等特征区别于相近种季氏金线ＢａＳ．ｊｉｉ Ｚｈａｎｇ ｅｔＤａｉ。     

云南、贵州中三叠世一种大型的肋鳞鱼(辐鳍鱼亚纲:肋鳞鱼目)（英文）

肋鳞鱼目是新鳍鱼类的一个干群,包括肋鳞鱼科和胸鳍鱼科,生活于欧洲和华南中、晚三叠世海洋环境。其中肋鳞鱼科鱼类通常个体较小,标准体长很少超过55 mm。根据最近发现于云南富源和贵州兴义中三叠世拉丁期(~240 Ma)海相地层的3块保存良好的鱼化石,命名了肋鳞鱼属一新种,霸王肋鳞鱼(Peltopleurus tyrannos sp.nov.)。该发现代表了中三叠世个体最大的肋鳞鱼属物种(体长达72 mm),为了解肋鳞鱼科的骨骼形态和摄食分异提供了新的信息。霸王肋鳞鱼不仅上、下颌口缘具有颇为粗壮的牙齿,在口内的颚翼骨上还生长有许多不同大小的钝牙;推测这些钝牙可能用于压碎一些带壳的无脊椎动物(如甲壳动物)和被硬鳞的小鱼,这与此前在兴义生物群中发现的东方肋鳞鱼形成鲜明的对比:东方肋鳞鱼体型很小(标准体长为26 mm)、口缘无牙、个体数量丰富,可能代表了一种集群游动、捕食细小浮游无脊椎动物的肋鳞鱼物种。新的发现表明华南中三叠世兴义生物群中的肋鳞鱼属已进化出两种不同的生态类型。     

大山口低等四足类动物群中的两栖类──甘肃玉门晚二叠世脊椎动物群系列报道之五

首次记述了大山口低等四足类动物群中的两栖类。其中走廊泰齿螈（Ｉｎｇｅｎｔｉｄｅｎｓｃｏｒｒｉｄｏｒｉｃｕｓ ｇｅｎ．ｅｔ ｓｐ．ｎｏｖ．）和祁连兄弟迟滞螈（Ｐｈｒａｔｏｃｈｒｏｎｉｓ ｑｉｌｉａｎｅｎｓｉｓ ｇｅｎ．ｅｔ ｓｐ．ｎｏｖ）为石炭蜥目迟滞鳄科（Ａｎｔｈｒａｃｏｓａｕｒｉａ, Ｃｈｒｏｎｉｏｓｕｃｈｉｄａｅ）的新成员。前者个体较大,具为数众多的犬齿状齿和齿骨表面的齿骨沟,冠状骨无齿,下颌表面纹饰蜂窝型;后者个体较小,侧生型齿,上颌仅有３个粗大的犬齿状齿。石油似卡玛螈（Ａｎａｋｍａｃｏｐｓ ｐｅｔｒｏｌｉｃｕｓ　ｇｅｎ．ｅｔ ｓｐ．ｎｏｖ．）是中国二叠纪离片椎目（Ｔｅｍｎｏｓｐｏｎｄｙｌｉ）的唯一代表。头骨中等大小,吻部扁平,前颌孔被前颌骨和锄骨包围,内鼻孔长大,锄骨表面粗糙,内鼻孔内缘光滑。这３个新属种的发现丰富了大山口动物群的组成,进一步证实了它与俄罗斯晚二叠世脊椎动物群的密切关系。     

贵州兴义晚三叠世贵州龙层新发现的鱼类

描述贵州兴义地区贵州龙层中新发现的鱼类4新属、4新种和一件新材料,它们是小鳞贵州鳕(新属、新种)Guizhouniscus microlepidus Liu gen．et sp．nov．,小短体鱼(新属、新种)Brachysomus minor Liu gen．et sp．nov．,秀丽兴义鱼(新属、新种)Xingyia gracilis Liu gen．et sp．nov．,臀鳞贵州真颌鱼(新属、新种)Guizhouella analilepidn Liu gen．et sp.nov,和优美贵州弓鳍鱼Guizhouamia bellula Liu et a1．,2002的一件新材料。新的发现和研究表明这是一个颇具特色的晚三叠世早期鱼群。文章还简要讨论各新属新种的系统分类位置和该鱼群在整个鱼类进化中的地位,着重论述贵州弓鳍鱼Guizhouamia的进化意义及弓鳍鱼科的系统发育问题。根据鱼群的特征贵州龙层的地质时代为晚三叠世。     

论晚二叠世的“克氏蛤”

系统评述目前世界上出现的晚二叠世克氏蛤类标本,并描述采自贵州,云南藏东的有关化石１属３种。其中１新属１新种,右壳足丝凹口特征在克氏蛤类研究中具有重要意义。晚二叠世的Ｃｌａｒａｉｏｉｄｅｓ ｇｅｎ．ｎｏｖ．以足丝凹口内端明显向腹方或后腹方扩展膨大成足丝凹湾而不同于早三叠世的Ｃｌａｒａｉａ．后者似由前者演化而来,它们在二叠－三叠系界线附近发生替代,可能与古生代末的绝灭事件相关。     

新疆玛纳斯古近纪—新的鲤科鱼类

记述了新疆玛纳斯古近纪安集海河组发现的鲤科(Cyprinidae)一新属、种——刘氏天山鱼(Tianshanicus liui gen.et sp.nov.)。新属具有臀鳍分叉鳍条多于7根、体长形稍侧扁、口端位、下咽齿近锥形而齿尖前后略侧扁、背鳍和臀鳍均无硬棘、背鳍起点略在腹鳍起点之后以及尾鳍深分叉等特征,表明它应属于雅罗鱼亚科(Leuciscinae sensu Cavender & Coburn,1992)。然而,它又具有额骨前宽后窄、顶骨长方形、膜质蝶耳骨很大、齿骨冠状突较低、鳃盖骨近长方形等特征而有别于本亚科中其他属。值得注意的是,这些特征却见诸于现生的胭脂鱼类Myxocyprinus中,这将对研究雅罗鱼类的起源很有意义。根据鱼化石和哺乳化石的证据,认为含鱼层的时代为晚始新世。     

Lessons from the first dorsal fin in atheriniforms—A new mode of dorsal fin development and its phylogenetic implications

The median fins in extant actinopterygians are the product of millions of years of evolution. During this time, different developmental patterns for the dorsal and anal fins emerged leading to a high variation in median fin morphology and ontogeny. In this study, the development of anal and dorsal fins in atheriniforms is described and its consequences for the current phylogenetic hypothesis are discussed. Developmental series of five atheriniform species were investigated using clearing and staining as well as antibody staining. The skeletal elements of the second dorsal fin and the anal fin emerge in a bidirectional pattern. The first dorsal fin, however, arises separately in front of the second dorsal fin after this one is almost completely formed. The pterygiophores of the first dorsal fin, including the interdorsal pterygiophores, develop from caudal to rostral, but the fin‐spines of the first dorsal fin form in the opposite direction. This new mode of fin development has been found in all examined atheriniform species with two dorsal fins. Several morphological characters of atheriniforms, including interdorsal pterygiophores, are also found in one other taxon: the Mugiliformes. Thus, several dorsal fin characteristics may provide evidence for a closer relationship of these two taxa.     

The median‐fin skeleton of the Eastern Atlantic and Mediterranean clingfishes Lepadogaster lepadogaster (Bonnaterre) and Gouania wildenowi (Risso) (Teleostei: Gobiesocidae)

Previous research on the osteology of the Gobiesocidae focused mostly on the neurocranium and the thoracic sucking disc (formed by the paired‐fin girdles). Little attention has been paid to the skeleton of the median fins. The dorsal‐ and anal‐fin skeleton of Lepadogaster lepadogaster and other gobiesocids (excluding Alabes, which lacks these fins) are characterized by the absence of spines, branched fin‐rays, and middle radials. In gobiesocids, the distal radials never ossify and consist of elastic hyaline‐cell cartilage. Gouania wildenowi is unique among gobiesocids in having further reductions of the dorsal‐ and anal‐fin skeleton, including a notable decrease in the size of the proximal‐middle radials in an anterior–posterior direction. Unlike L. lepadogaster, which exhibits a one‐to‐one relationship between the dorsal‐ and anal‐fin rays and proximal‐middle radials, G. wildenowi has a higher number of proximal‐middle radials than distal radial cartilages and fin rays in the dorsal and anal fins. In G. wildenowi, the dorsal‐ and anal‐fin rays do not articulate with the distal tip of the proximal‐middle radials but are instead positioned between proximal‐middle radials, which is unusual for teleosts. Previously unrecognized dorsal and ventral pads of elastic hyaline‐cell cartilage are also present in the caudal skeleton of L. lepadogaster, G. wildenowi, and all other gobiesocids examined. J. Morphol., 2010. © 2009 Wiley‐Liss, Inc.     

Leis' conundrum: homology of the clavus of the ocean sunfishes. 2. Ontogeny of the median fins and axial skeleton of Ranzania laevis (Teleostei, Tetraodontiformes, Molidae)

One of the most conspicuous characters of the ocean sunfishes, family Molidae, is the punctuation of the body by a deep, abbreviated, caudal fin-like structure extending vertically between the posterior ends of the dorsal and anal fins, termed the clavus by Fraser Brunner. Homology of the clavus has been a matter of debate since the first studies on molid anatomy in the early 1800s. Two hypotheses have been proposed: 1) It is a highly modified caudal fin; 2) It is formed by highly modified elements of the dorsal and anal fins. To resolve this homology issue, we studied the ontogeny of the molid vertebral column and median fins and compared it to that of a less morphologically derived gymnodont (see Part 1 of this study), a member of the family Tetraodontidae. We show that in molids the chorda never flexes during development, that the claval rays form from the posterior ends of the dorsal and anal fins toward the middle, thus closing the gap inward, and that elements of the molid clavus have an identical development and composition as the proximal-middle and distal radials of the regular dorsal and anal fins. We thus conclude that the molid clavus is unequivocally formed by modified elements of the dorsal and anal fin and that the caudal fin is lost in molids.     

Pectoral fin and girdle development in the basal actinopterygians Polyodon spathula and Acipenser transmontanus

The pectoral fins of Acipenseriformes possess endoskeletons with elements homologous to both the fin radials of teleosts and the limb bones of tetrapods. Here we present a study of pectoral fin development in the North American paddlefish, Polyodon spathula, and the white sturgeon, Acipenser transmontanus, which reveals that aspects of both teleost and tetrapod endoskeletal patterning mechanisms are present in Acipenseriformes. Those elements considered homologous to teleost radials, the propterygium and the mesopterygial radials, form via subdivision of an initially chondrogenic plate of mesenchymal cells called the endoskeletal disc. In Acipenseriformes, elements homologous to the sarcopterygian metapterygium develop separately from the endoskeletal disc as an outgrowth of the endoskeletal shoulder girdle that extends into the posterior margin of the finbud. As in tetrapods, the elongating metapterygium and the metapterygial radials form in a proximal to distal order as discrete condensations from initially nonchondrogenic mesenchyme. Patterns of variation seen in the Acipenseriform fin also correlate with putative homology: all variants from the "normal" fin bauplan involved the metapterygium and the metapterygial radials alone. The primary factor distinguishing Polyodon and Acipenser fin development from each other is the composition of the endoskeletal extracellular matrix. Proteoglycans (visualized with Alcian Blue) and Type II collagen (visualized by immunohistochemistry) are secreted in different places within the mesenchymal anlage of the fin elements and girdle and at different developmental times. Acipenseriform pectoral fins differ from the fins of teleosts in the relative contribution of the endoskeleton and dermal rays. The fins of Polyodon and Acipenser possess elaborate endoskeletons overlapped along their distal margins by dermal lepidotrichia. In contrast, teleost fins generally possess relatively small endoskeletal radials that articulate with the dermal fin skeleton terminally, with little or no proximodistal overlap.     

Development of the pectoral, pelvic, dorsal and anal fins in cultured sea bream

The pectoral fin girdle was the first element of the fins to develop in Sparus aurata. By 3·1mm L _N (notochord length) the cleithrum was ossified and the cartilaginous caracoid-scapula was present. The fin was fully developed at 11·6 mm L _S (standard length) and by 16·0 mm L _S most elements of the fin were ossified. The pelvic fins were the last pair to develop and rudiments of these were first detected at 7·9 mm L _S. The pelvic fin and girdle were completely formed and ossified at 16·0 mm L _S. The development of dorsal and anal fins began at c. 6·5–7·0 mm L _S with the formation of 10 cartilaginous dorsal proximal radials and eight cartilaginous ventral proximal radials. The three cartilaginous predorsals (supraneurals) appeared at 7·7 mm L _S and the ossification of dorsal and anal proximal and distal radials began, respectively, at 10·5 mm L _S and 11·3 mm L _S. Ossified structures in the fins were also classified according to their origin, as being either dermal or endochondral. Finally the chronology of appearance of fin structures in S. aurata was compared with that reported for other Sparidae, Engraulidae and Haemulidae.     

Larval development of Hypsophrys nicaraguensis (Pisces: Cichlidae) under laboratory conditions

The cichlid Hypsophrys nicaraguensis is a popular fish known as butterfly, and despite its widespread use as pets, little is known about its reproductive biology. In order to contribute to this knowledge, the study describes the relevant larval development characteristics, from adult and larval cultures in captivity. Every 12h, samples of larvae were collected and observed under the microscope for larval stage development, and every 24h morphometric measurements were taken. Observations showed that at 120h, some larvae had swimming activity and the pectoral fins development was visible; at 144h, the dorsal fin appear and all larvae started food intake; at 168h, the formation of anal fins begins, small rudiments of pelvic fins emerge, the separation of caudal fin from anal and dorsal fins starts, and the yolk sac is reabsorbed almost completely; at 288h, the pelvic fins starts to form; at 432h, the rays and spines of dorsal and anal fins can be distinguished, both the anal and the dorsal fins have the same number of spines and rays as in adults. After 480h larvae have the first scales, ending the larval stages and starting the transformation to fingerlings. Larvae were successfully fed with commercial diet.     

Anatomy and muscle activity of the dorsal fins in bamboo sharks and spiny dogfish during turning maneuvers

Stability and procured instability characterize two opposing types of swimming, steady and maneuvering, respectively. Fins can be used to manipulate flow to adjust stability during swimming maneuvers either actively using muscle control or passively by structural control. The function of the dorsal fins during turning maneuvering in two shark species with different swimming modes is investigated here using musculoskeletal anatomy and muscle function. White‐spotted bamboo sharks are a benthic species that inhabits complex reef habitats and thus have high requirements for maneuverability. Spiny dogfish occupy a variety of coastal and continental shelf habitats and spend relatively more time cruising in open water. These species differ in dorsal fin morphology and fin position along the body. Bamboo sharks have a larger second dorsal fin area and proportionally more muscle insertion into both dorsal fins. The basal and radial pterygiophores are plate‐like structures in spiny dogfish and are nearly indistinguishable from one another. In contrast, bamboo sharks lack basal pterygiophores, while the radial pterygiophores form two rows of elongated rectangular elements that articulate with one another. The dorsal fin muscles are composed of a large muscle mass that extends over the ceratotrichia overlying the radials in spiny dogfish. However, in bamboo sharks, the muscle mass is divided into multiple distinct muscles that insert onto the ceratotrichia. During turning maneuvers, the dorsal fin muscles are active in both species with no differences in onset between fin sides. Spiny dogfish have longer burst durations on the outer fin side, which is consistent with opposing resistance to the medium. In bamboo sharks, bilateral activation of the dorsal in muscles could also be stiffening the fin throughout the turn. Thus, dogfish sharks passively stiffen the dorsal fin structurally and functionally, while bamboo sharks have more flexible dorsal fins, which result from a steady swimming trade off. J. Morphol. 274:1288–1298, 2013. © 2013 Wiley Periodicals, Inc.     

Neolumpenus unocellatus,a new genus and species of stichaeid fish from Japan

Neolumpenus unocellatus gen. et sp. nov., a stichaeid fish (subfamily Lumpeninae,sensu Makushok, 1958) is described on the basis of a single specimen found in the stomach of the Pacific cod,Gadus macrocephalus Tilesius, caught off Akkeshi, Hokkaido, Japan. The new genus and species is distinguished from all other lumpenines in having the following combination of characters: 1) 51 dorsal spines, 33 anal fin rays, 57 total vertebrae; 2) broad pelvic fin with deeply-branched soft rays; 3) lower rays of pectoral fin branched and not prolonged backward; 4) prevomerine and palatine teeth present; 5) pungent spines present in pelvic and anal fins; 6) upper lip fused to snout anteriorly; 7) gill openings not extending forward beyond a vertical through posterior margin of eye; 8) minimal (fifth) hypural present; 9) first interneural spine inserted between first and second neural spines; 10) extremely large cephalic sensory pores present; 11) high, steep snout; 12) ocellus on dorsal base of caudal fin.