Chronological changes in stone tool assemblages from Krapina (Croatia)

This study presents the results of the first recent analysis of stone tool assemblages from Krapina (Croatia). All assemblages are Pleistocene in age and many are associated with human remains, the Krapina Neandertals. The assemblages are described typologically and technologically, and subtle chronological changes in raw material selection and technology of tool blank production are observed. These changes involve increasingly sophisticated and selective use of lithic materials. Changing artefact assemblages are considered in light of variability in the hominids from Krapina, and are interpreted as reflecting behavioral change among Neandertals rather than between Neandertal and modern human populations.     

Resource selection of lithic raw materials in the Middle Palaeolithic in southern France

The work reported here uses several approaches to examine the costs and benefits associated with exploiting potential sources of lithic raw material in the Vaucluse, southern France, and then tests the results against the proportions of raw materials from various sources found in the lithic assemblage of a Middle Palaeolithic site, the Bau de l’Aubesier. A previously published equation designed to quantify the attractiveness of each source proves to be significantly correlated with source use, but the results show that it can be improved. We then individually test the components of the attractiveness equation (raw material quality, source extent, terrain difficulty, and the size and abundance of raw material pieces at the source) and additional variables (Calories expended to get from the source to the site using a straight-line route, Calories expended using a least-cost path, surface distance of the source from the site, and distance to the closest used source) using generalized linear models. Since very similar raw materials can be found at more than one source, we combine such similar sources into source areas, and test the area of the source area (AOSA) and the total area of sources within the source area (AOSISA), as two additional independent variables. The best model shows that raw material quality, source extent, abundance of large and very large rocks, and AOSISA, are positively correlated with use of sources, while terrain difficulty, abundance of small and medium rocks, Calories, and AOSA, are negatively correlated with source use. This shows that the hominins at the Bau de l’Aubesier optimized their raw material procurement to get the most good quality material that they could, while minimizing the time and energy spent getting and using it.     

The origins of stone tool technology in Africa: a historical perspective

The search for the earliest stone tools is a topic that has received much attention in studies on the archaeology of human origins. New evidence could position the oldest traces of stone tool-use before 3.39 Myr, substantially earlier than previously documented. Nonetheless, the first unmistakable evidence of tool-making dates to 2.6 Ma, the period in which Oldowan assemblages first appear in the East African record. However, this is not an unchangeable time boundary, and considerations about the tempo and modo of tool-making emergence have varied through time. This paper summarizes the history of research on the origins of stone knapping in Africa and places the current evidence in a historical perspective.     

The major stages in the evolution of artiodactyl communities from the Pliocene-early Middle Pleistocene of northern Eurasia: Part 1

Seven main stages in the evolution of Pliocene-Pleistocene artiodactyl communities from Russia, Ukraine, Moldova, Georgia, Kazakhstan, and Tajikistan within the interval from 5.3 through 0.4 Ma are characterized. The dominant groups are briefly revised. The major biotic events and the development of the main phyletic lineages are discussed.     

The Early-Middle Pleistocene environmental and climatic change and the human expansion in Western Europe: A case study with small vertebrates (Gran Dolina, Atapuerca, Spain)

The dispersal of hominins may have been favored by the opening of the landscape during the Early-Middle Pleistocene transition (EMP) in Western Europe. The structure of the small-vertebrate assemblages of the archaeo-paleontological karstic site of Gran Dolina in Atapuerca (Burgos, Spain) shows important environmental and climatic changes in the faunal succession, across the Matuyama-Brunhes boundary at 780 ka. These changes are interpreted to indicate impoverishment of the forests, along with an increase in dry meadows, and open lands in general that entailed a tendency towards the loss of diversity in small-vertebrate communities above the EMP. We evaluate variation in diversity of the faunal succession of Gran Dolina using Shannon’s Second Theorem as an index of ecosystem structure. The long cultural-stratigraphic sequence of Gran Dolina during the EMP is somewhat similar in its completeness and continuity to that in the locality of Gesher Benot Ya'aqov in the Upper Jordan Valley. We also evaluate related data including faunal and floral (pollen) succession. Both localities present cold, dry and humid, warm fluctuations at the transition between the Early and the Middle Pleistocene. Comparisons between these sites present opportunities to understand large-scale climatic changes.     

Main stages in evolution of Artiodactyla communities from the Pliocene-Early Middle Pleistocene of northern Eurasia: Part 2

Changes in diversity of Artiodactyla from the Pliocene to Middle Pleistocene of northern Eurasia are considered. The major turnovers of artiodactyl communities correlate with the major global cooling events at the boundaries of epochs and ages and were accompanied by migrations. A correlation of the main biotic events in northern Eurasia and adjacent areas is presented. Zoogeographical features of Artiodactyla evolution in northern Eurasia and the role of Artiodactyla in the establishment of different bioms are addressed.     

Regional behaviour among late Neanderthal groups in Western Europe: A comparative assessment of late Middle Palaeolithic bifacial tool variability

Population dynamics between and within Pleistocene groups are vital to understanding wider behavioural processes like social transmission and cultural variation. The late Middle Palaeolithic (MIS 5d-3, ca. 115,000–35,000 BP [years before present]) permits a novel, data-driven assessment of these concepts through a unique record: bifacial tools made by classic Neanderthals. Previously, studies of late Middle Palaeolithic bifacial tools were hampered by a convoluted plethora of competing terms, types and regional entities. This paper presents a large-scale intercomparison of this tool type, and bridges typo-technological and spatio-temporal data from across Western Europe (Britain, Belgium, the Netherlands, France and Germany).     

生物制品原辅料及其质量控制

生物制品原辅料的质量控制是保证产品质量的重要因素,就生物制品原辅料分类及质量控制进行了阐述和归纳,并就国家对生物制品原辅料的质量控制要求进行了探讨。     

The Devil's Trails: Middle Pleistocene Human Footprints Preserved in a Volcanoclastic Deposit of Southern Italy

Three fossilized trackways of human footprints were discovered in a pyroclastic flow dated to 385–325 ka, along the western margin of the Roccamonfina volcanic complex in Southern Italy. The pyroclastic flow is zeolite-rich and was probably emplaced at a relatively low temperature, forming a nonwelded pyroclastic deposit. The well-preserved footprints show the interesting strategies used by ancient humans in descending steep slopes. The footprints, short and very broad, are entaxonic, plantigrade, with an average length of 24cm and an average width of 10 cm. The three trackways are narrow with an oblique pace of about 60 cm and a stride of about 120 cm.

If dating constraints are correct, the Roccamonfina human fossil footprints are the oldest known in Europe and among the oldest in the world. Moreover, they are preserved on a steep slope lying in a subaerial setting and are associated with handprints. For their potential in preserving evidence of behavior, these tracks provide unique insight into the activities of some of the earliest known Europeans.     

Neandertal mobility and large-game hunting: The exploitation of reindeer during the Quina Mousterian at Chez-Pinaud Jonzac (Charente-Maritime, France)

Neandertals were effective hunters of large ungulates throughout their geographic and temporal ranges. Equipped with this knowledge, researchers in paleoanthropology continue to seek insight on the relationships between hunting and subsistence strategies with other components of the Neandertals’ niche, such as mobility, site use, and lithic technology. The Quina Mousterian deposits from the rockshelter site of Chez Pinaud Jonzac (Charente-Maritime, France; hereafter Jonzac) offer an excellent opportunity to pursue these issues. This paper focuses on the extensive and well-preserved skeletal remains of reindeer (Rangifer tarandus) recovered from recent excavations of the site, representing at least 18 individuals that were hunted by Neandertals during the fall through winter. Our zooarchaeological results indicate that all ages of reindeer were hunted but adult individuals predominate. No bias is evident in the comparable frequencies of males and females. These prey were butchered on-site, with abundant evidence of meat filleting and marrow exploitation. In the excavated sample, the absence of hearths and the almost complete lack of burned bones or stones suggest that Neandertals were not using fire to assist with processing the reindeer carcasses. The zooarchaeological results presented here indicate that reindeer were hunted during a restricted window of time when they were seasonally abundant in the local area near Jonzac. Taken together with the lithic industry based on bifacial elements, the evidence is consistent with a pattern of site use by highly mobile hunter-gatherers making frequent, short-term visits. Ongoing research at Jonzac and other Quina Mousterian localities will contribute to a better understanding of Neandertal behavior during cold climate phases.     

The Middle Stone Age of the northern Kenyan Rift: age and context of new archaeological sites from the Kapedo Tuffs

Rift Valley sites in southern Ethiopia and northern Kenya preserve the oldest fossil remains attributed to Homo sapiens and the earliest archaeological sites attributed to the Middle Stone Age (MSA). New localities from the Kapedo Tuffs augment the sparse sample of MSA sites from the northern Kenya Rift. Tephrostratigraphic correlation with dated pyroclastic deposits from the adjacent volcano Silali suggests an age range of 135-123 ka for archaeological sites of the Kapedo Tuffs. Comparisons of the Kapedo Tuffs archaeological assemblages with those from the adjacent Turkana and Baringo basins show broad lithic technological similarity but reveal that stone raw material availability is a key factor in explaining typologically defined archaeological variability within this region. Spatially and temporally resolved comparisons such as this provide the best means to link the biological and behavioral variation manifest in the record of early Homo sapiens.     

The relative importance of local, regional and historical factors determining the distribution of plants in fragmented riverine forests: an emergent group approach

Aim To investigate how local, regional and historical factors shape the herbaceous plant communities in fragmented riverine forests, and how the community composition and species richness of these fragments is related to the interplay between the environmental factors and specific plant life‐trait combinations. Location Riverine forest fragments in the Grand‐duché de Luxembourg. Methods Forest fragments were surveyed for their abundance in herbaceous plant species. All plant species where clustered into Emergent Groups (EG) by means of a formal classification based on 14 life‐history traits. Within each EG, the local, regional and historical factors were related to the community composition using partial Canonical Correspondence Analyses (pCCA) and to the species richness using Generalized Linear Models (GLMs). The EG colonization ability was characterized by means of logistic regressions. Results We defined and characterized seven EGs, among which three consisted of forest specialist species (barochorous perennials, short geophytes and zoochorous perennials), which exhibited specific life‐trait combinations: large and short‐lived seeds and/or vernal phenology. Differences in EG composition between forest fragments were mainly explained by local environmental factors such as soil productivity and pH. The richness of barochorous perennials and short geophytes was well predicted by the historical and regional factors. The colonization ability appeared very low for barochorous perennials and short geophytes. Main conclusions Local environmental conditions appear to drive the differentiation of the riverine forest plant communities owing to the specific habitat requirements of many forest species. Spatial and temporal forest discontinuities affect the richness of forest specialist species, due to dispersal and/or recruitment limitations. The emergent group approach enhances the understanding of the relative influence of local, regional and historical factors by distinguishing between forest specialists from generalists or ‘matrix’ species, which have a masking effect.     

Membrane vesicles from brown adipose tissue: A tool for the study of amino acid transport. The case of L-alanine

A density gradient method is used to isolate membrane vesicles from brown adipose tissue. These respond to changes in osmolarity and show the classical overshoot pattern when L-alanine uptake is assayed. Transport is shown to be effected by two components: a linear (K_d=0.498 min⁻¹) and Na⁺-dependent saturable component (K_m=2.3 mM) and a V_max=19.9 pmol/μg protein·min). This pattern is similar to that shown by cells isolated from brown adipose tissue.     

The geographical and chronological pattern of herpetofaunal Pleistocene extinctions on the Iberian Peninsula

Despite the postulation of the “herpetofaunal stability hypothesis”, which suggests that the herpetofauna underwent relatively few changes during the Pleistocene, the amphibian and squamate faunas of western Europe are known to have progressively diminished during the Pliocene and the beginning of the Pleistocene. Iberian Early Pleistocene sites continue to document the presence of “exotic” herpetofaunal elements that are supposed to have disappeared from the mainland, such as Oriental vipers, agamid lizards, the anguids Pseudopus and Dopasia, and a representative of the green toad group (Bufo viridis sensu lato), as well as possibly gekkonids, a scincid lizard (Lygosominae), the anguid Ragesaurus and blind snake (Scolecophidia). The geographical and temporal pattern of these progressive southward withdrawals and extirpations on the Iberian Peninsula shows that extirpation events occurred in northern Spain at roughly the Olduvai paleomagnetic event and in southern Spain just before the Jaramillo paleomagnetic reversal, thus permitting us to establish a framework that can be used to complement the biochronological zonation of the Iberian Early Pleistocene.     

The role of atmospheric dispersion models and ecosystem sensitivity in the determination of characterisation factors for acidifying and eutrophying emissions in LCIA

Background, aim and scope  The methodological choices and framework to assess environmental impacts in life cycle assessment are still under discussion. Despite intensive developments worldwide, few attempts have been made hitherto to systematically present the role of different factors of characterisation models in life cycle impact assessment (LCIA). The aim of this study is to show how European average and country-dependent characterisation factors for acidifying and eutrophying emissions differ when using (a) acidifying and eutrophying potentials alone, (b) depositions from an atmospheric dispersion model or (c) critical loads in conjunction with those depositions. Furthermore, in the latter case, the contributions of emissions, an atmospheric transport model and critical loads to changes in characterisation factors of NO₂ are studied. In addition, the new characterisation factors based on the accumulated exceedance (AE) method are presented using updated emissions, a new atmospheric transport model and the latest critical loads. Materials and methods  In this study, characterisation factors for acidifying and eutrophying emissions are calculated by three different methods. In the ‘no fate’ (NF) methods, acidifying and eutrophying potentials alone are considered as characterisation factors. In the ‘only above terrestrial environment’ (OT) approach, characterisation factors are based on the deposition of the acidifying or eutrophying substances to terrestrial land surfaces. The third method is the so-called AE method in which critical loads are used in conjunction with depositions. The results of the methods are compared both at the European and the country level using weighted mean, weighted standard deviation, minimum and maximum values. To illustrate the sensitivity of the AE method, changes in European emissions, employed atmospheric dispersion model and the critical loads database are conducted step-by-step, and the differences between the results are analysed. Results and discussion  For European average characterisation factors, the three characterisation methods of acidification produce results in which the contributions of NH₃, NO₂ and SO₂ to the acidification indicator do not differ much within each method when 1 kg of each acidifying substance is emitted. However, the NF methods cannot describe any spatial aspects of environmental problems. Both OT and AE methods show that the spatial aspects play an important role in the characterisation factors. The AE method results in greater differentiations between country-dependent characterisation factors than does the OT method. In addition, the results of the AE and OT methods differ from each other for individual countries. A major shortcoming of the OT approach is that it does not consider the sensitivity of the ecosystems onto which the pollutants are deposited, whereas the AE approach does. In the case of the AE method, a new atmospheric dispersion model, new information on emissions and critical loads have a different influence on the characterisation factors, depending on the country. The results of statistics show that the change in the atmospheric dispersion model has a greatest influence on the results, since ecosystem-specific depositions are taken into account for the first time. Conclusions and recommendations  The simple NF methods can be used in a first approximation to assess the impacts of acidification and terrestrial eutrophication in cases where we do not know where the emissions occur. The OT approach is a more advanced method compared with the NF method, but its capability to describe spatial aspects is limited. The AE factors are truly impact-oriented characterisation factors and the information used here represents the current best knowledge about the assessment practice of acidification and terrestrial eutrophication in Europe. The key message of this study is that there is no shortcut to achieving advanced characterisation of acidification and terrestrial eutrophication: an advanced methodology cannot develop without atmospheric dispersion models and information on ecosystem sensitivity.     

The Casal de' Pazzi archaic parietal: comparative analysis of new fossil evidence from the late Middle Pleistocene of Rome

A fossilized fragment of human parietal bone has been recently recovered from the lowest layer of the Casal de' Pazzi fluvial deposit (stratigraphically dated at about 200–250 ky BP). The fossil presents characters-i.e., thickness, degree and development of curvature, type of endocranial vascularization-which distinguish it from the corresponding cranial regions of both Homo erectus and anatomically modern Homo sapiens. While a morphological orientation towards Neanderthal characters can be considered, the affinities of the Casal de' Pazzi parietal are primarily with other late Middle Pleistocene specimens. The authors conclude that the Casal de' Pazzi human find can be assigned to the “archaic Homo sapiens” group falling within the European pre-Neanderthal range. Its particular morphology constitutes new evidence of human evolution from the geographical area of Rome.