On the Evolution of the Palynoflora and Climatic Environment During Late Pleistocence in Tengger Desert,China

Palynoflora studies on the Duantouliang section, located at 39º40'N, 103º55'E in Northwestern Tengger Desert, China showed that, based on the spore-pollen assemblages, the major vegetation and climatic environment between 42 000 to 23 000 a BP could be divided into the following different periods: Ⅰ. From 42 000 to 38 000 a BP, the spore-pollen assemblages displayed that the mixed conifer/deciduous broad-leaved forests developed on the mountain and its foothill regions where the bare Gobi-desert are at present; At that time, Populus and Salix forests and grassland surrounded the Paleaolake, the climate condition was much warmer and humid than today; Ⅱ. From 38 000 to 37 000 a BP, the climate was warm and moist, it was the most suitable period for the plant growth, the studied area was dominated by the temperate and warm-temperate mixed broad-leaf deciduous and needleleaf forest, there was meadow spreaded on the river sides and lake beaches; Ⅲ. From 31 000 to 30000 a BP, the needleleaf forests and cold-temperate Salix oritrepha shrubs were flourishing, and the climate at that time was relatively cold;Ⅳ. From 30000 to 28000 a BP, the temperatures began increasing, the high lake levels was formed during this time, and the vegetations were meadows and swamps; V. From 28 000 to 23 000 a BP, temperate Cupressaceae and Betula mixed conifer/deciduous forests grew on mountain and foothill region, grassland developed on plain areas, Salix was on lake and river sides. This indicates a warm and moist climate condition but it was drier than the earliest period.     

Holocene climate and environmental history of Brurskardstjørni,a lake in the catchment of Øvre Heimdalsvatn,south-central Norway

The Holocene lake history, vegetation history and climate history of Brurskardstjørni, an alpine lake in the Jotunheimen Mountains of south-central Norway, are reconstructed. The reconstructions are based on fossil pollen, plant macrofossils, diatoms, chironomids and sediment characteristics. Subsequent to deglaciation, the lake was formed at about 11,000 cal years BP. A diverse chironomid assemblage quickly colonised the lake, whereas the first diatoms were found about 400 years later. At that time, the lake water was turbid with a high pH. The surrounding soils were immature and unstable and dominated by open pioneer vegetation. Compared to the present, summer temperatures were warmer and there was less winter precipitation. From about 10,000 cal years BP, local organic production increased rapidly and from about 9,500 cal years BP a few macrofossils and a high pollen influx of birch suggest that the tree-line was close to the lake. Pine most likely reached its highest tree-line altitude around 9,000 cal years BP and has receded since that time. From about 5,000 cal years BP, the total amount of trees and shrubs decreased and the landscape became more open, probably due to decreasing temperatures and increasing effective moisture lowering the birch tree-line. Coinciding with a cooling during the last 3,000 years, lake-water pH decreased. There is large incongruence between the Holocene July temperatures inferred from pollen and chironomids. The biological proxies responded to a combined effect of environmental change and biotic interactions. This response is interpreted with reference to taxon–environment relationships in the modern calibration data sets and with reference to the latent structure and ecological demands of the fossil assemblages.     

From ice age to modern: a record of landscape change in an Andean cloud forest

Aim To investigate the palaeoecological changes associated with the last ice age, subsequent deglaciation and human occupation of the central Andes. Location Lake Pacucha, Peruvian Andes (13°36′26″ S, 73°19′42″ W; 3095 m elevation). Methods Vegetation assemblages were reconstructed for the last 24 cal. kyr bp (thousand calibrated ¹⁴C years before present), based on pollen analysis of sediments from Lake Pacucha. An age model was established using ¹⁴C accelerator mass spectrometry dates on bulk sediment. Fossil pollen and sedimentological analyses followed standard methodologies. Results Puna brava replaced the Andean forest at the elevation of Lake Pacucha at the Last Glacial Maximum (LGM). Deglaciation proceeded rapidly after 16 cal. kyr bp , and near‐modern vegetation was established by c. 14 cal. kyr bp . The deglacial was marked by the range expansion of forest taxa as grassland taxa receded in importance. The mid‐Holocene was marked by a lowered lake level but relatively unchanged vegetation. Quinoa and maize pollen were found in the latter half of the Holocene. Main conclusions Temperatures were about 7–8 °C colder than present at this site during the LGM. The pattern of vegetation change was suggestive of microrefugial expansion rather than simple upslope migration. The mid‐Holocene droughts were interrupted by rainfall events sufficiently frequent to allow vegetation to survive largely unchanged, despite lowering of the lake level. Human activity at the lake included a 5500‐year history of quinoa cultivation and 3000 years of maize cultivation.     

New High Resolution Pollen Records from Two Lakes in Xizang (Tibet)

Studies on the pollen from cores of Lake Hidden (29~48.77' N, 92~22.37' E) and Ren Co (30%3.97'N, 96~40.97'E) in Southeast Xizang (Tibet) showed climatic changes and the evolution of vegetation since the last Glacial Maximum (LGM). Before 16 ka BP pollen assemblages were dominated by Chenopodiaceae (20% -50% ) and Artemidis (10% ~ 30% ) and pollen influx values were very low from Lake Ren Co in Basu county. The vegetation around the lake was probably a desert-steppe during the LGM. The data also suggest that the climate in the Basu area was cold and dry during the LGM and the last glacial time. The mean annual temperature was probably 4 ~ 6 ℃ colder than the present, and the mean annual precipitation was only 250 mm, about 40% of the present. The Southwest monsoon became stronger from 12 ka BP to 6 ka BP reaching its acme by 7 ka BP, but weakened gradually from 5 ka BP to the present. The paleovegetation was dominantly forest or forest-meadow around Lake Hidden and Ren Co during the 9.2 ~ 5 ka BP. The mean temperature in January was probably 2 ~ 3 ℃ higher than the present, and the mean annual precipitation was 100 mm more than the present. The timing of vegetational and climatic changes based on pollen records generally supports the results of global climatic-model experiments that predict a strengthened monsoon system during the early to middle Holocene followed by a weakened monsoon system.     

云贵高原抚仙湖近13300年的花粉/炭屑记录*

抚仙湖是云贵高原著名的断陷深水湖,其沉积物蕴藏着流域地质历史时期丰富的环境信息。对钻取自该湖的900cm 湖泊沉积物岩芯进行花粉/炭屑分析及花粉数据的主成分分析表明,抚仙湖流域的植被、气候与火灾在过去的13 300年经历了5个阶段的变化：(1)13 300—10 400cal．a BP,植被以松林为主,伴有山地暗针叶林和常绿阔叶林,表明该时期气候较为冷湿,森林火灾多发,在后期随着温度和湿度的降低,森林火灾愈加频繁。(2)10 400—5 700cal．a BP,松林收缩,常绿阔叶林扩张,出现一定数量的落叶阔叶林,显示该时期气候偏暖偏干;此阶段早期随着气候变暖变干森林火灾的发生延续上阶段高发的状态,直到9 500cal．a BP后随着湿度的增加森林火灾明显减少。(3)5 700—1 800cal．a BP,松林变化较小,常绿/落叶阔叶林比重增大,首次出现了暖热性的枫香林,显示该时期暖湿的气候特征,火灾发生频率低。(4)1 800—500cal．a BP,松林扩张,阔叶林收缩,本阶段后期草本植被比重开始增加,显示该时期气候相对冷干,森林火灾发生频率较高。(5)500cal．a BP至今,松林收缩,落叶阔叶树种增多,草本植物花粉明显增多,显示该时期气候温凉偏干,森林火灾发生频率降低。     

Charcoal analysis, a method to study vegetation and climate of the Holocene: The case of Lago di Pergusa (Sicily, Italy)

The direct comparison between microcharcoal and pollen data from the Holocene sediment core of Lago di Pergusa (central Sicily, Italy) led us to investigate the linkages between fire, vegetation, and climate in the Mediterranean Basin. The role of human populations in shaping the environment of the last millennia was closely examined as well. Pollen and charcoal were extracted using a standard pollen methodology and both identified from the same pollen slides. At Lago di Pergusa the importance of fire in maintaining and favouring a Mediterranean vegetation, a basic concept in ecology, seems to be inconsistent, as important fires took place in the open environments with xeric vegetation at the beginning of the Holocene well before the expansion of evergreen vegetation. On the contrary some big fires seem to have happened in the period (around 8000 years BP) in which the greatest wetness, the thickest forest canopy, and the most important expansion of submontane vegetation of the record are found. A probable explanation of this phenomenon can be found in the precipitation regime, in the increased available biomass and in the resulting increase in combustible material, indicating also that the highest humidity achieved in central Sicily during the Holocene was not strong enough to prevent fires. During this period drops in arboreal pollen concentration match in minima in the charcoal curves. Fires were practically absent during the Neolithic and Eneolithic periods and not responsible for the general trend towards aridification identified in the pollen record, at times in which prehistoric populations are generally believed to have caused forest clearance by burning. Important and frequent burnings are documented for the Bronze and Iron Ages and probably not all of them were fires of living trees, but burning of wood for producing metals, cooking or heating. The occurrence of a fire around 3700 years BP temporarily caused a minor decrease of arboreal pollen concentration. In the open landscapes of the two last millennia an almost continuous sign of regional burnings is recognized, confirming the historical evidence of an intense land use in central Sicily at least since Roman times.     

科尔沁沙地麦里地区晚全新世植被演化

根据一个泥炭剖面高分辨率花粉资料序列,初步重建了科尔沁沙地麦里地区晚全新世的植被演化过程。在3100 a BP以前,该地以固定沙丘上生长的蒙古栎(Quercus mongolica Fisch.)等阔叶疏林为主;以后随着半固定沙丘和流动沙丘增多,蒿属(Artemisia L.)和藜科等草本植物逐渐增加,而固定沙丘上的阔叶树则趋于减少。在1000～660 a BP期间,麦里地区几乎所有沙丘植物都经历了繁盛生长,蒙古栎种群的扩大尤其显著。3100 a BP以来植被和沙丘的演化可能同人类活动影响有直接关系,而1000～660 a BP期间的植物繁盛则由生长季雨量增加引起。花粉统计分析还表明,在森林草原等生态过渡带,花粉浓度和花粉沉积率比花粉百分比更好地指示了植被演化过程。     

Late Quaternary vegetation and climatic history of eastern Nepal

The late Quaternary vegetation and climatic history at high altitude in eastern Nepal was studied through stratigraphy, radiocarbon dating and pollen analysis. The Thulo Pokhari lake (27°41′ N, 87°43′ E, ca. 3980 m a.s.l.) is just above the subalpine Abies spectabilis–Betula utilis forest zone and surrounded by Rhododendron scrub. The climate in the studied area is very humid under the influence of the monsoon. Silty sediments suggesting lake conditions prevailed at the lower part of the sediment samples, whereas the upper part consisted mainly of herbaceous peat indicating a mire condition. The pollen record extends back to ca. 11000 yr bp . Although the absence of major changes between the pollen assemblages of the Pleistocene and those in the Holocene was attributed to the situation of the sampling plot above the forest limit, the results suggest the following history. (1) The abundance of Pinus pollen around 11000 yr bp represents expansion of the area unoccupied by late successional trees and/or a drier condition. Significant occurrence of Picea pollen is noteworthy, since this genus is absent in eastern Nepal at present in spite of its wide distribution throughout the neighbouring areas of the Himalayas. (2) The increase in abundance of Quercus pollen and the decline in Pinus pollen in the latest Pleistocene represent climatic amelioration favourable to late successional mesophytes. (3) The climate was most moist and mild in the SP-III, although the absolute age of the pollen zone is not distinct. The percentages of broadleaved trees such as Alnus, Betula, Carpinus and Corylus were increased at the expense of Quercus and coniferous pollen types. Sedimentation was interrupted by lake level decline and/or glacial advance during the period from ca. 11000–1600 yr bp . (4) The increase in abundance of coniferous pollen types such as Pinus, Abies and Tsuga indicates that the climate became drier and cooler, and the present subalpine forest zone composed of A. spectabilis, T. dumosa and Betula utilis was established around 1600 yr bp . The formation of bog or alpine meadow vegetation during the period was shown by the ¹⁴C dates of the peat and the increase in Rosaceae and herbaceous pollen. (5) The increase in abundance of Ericaceae and Alnus pollen in the SP-V indicates that the present alpine Rhododendron scrub zone has been established since ca. 940 yr bp as a result of human influence on the original vegetation. This interpretation is supported by the decline in the proportion of arboreal pollen. Since the age of the initiation of human influence varies with the altitude and region, further study will be necessary to determine the accurate age.     

Vegetation changes in Empakaai Crater, northern Tanzania, at 14,800–9300 cal yr BP

Vegetation changes are documented from a well-dated pollen record from Lake Emakat, Empakaai Crater, northern Tanzania. This pollen record includes the time interval covering the Pleistocene/Holocene transition, analysed at a resolution interval averaging 200 yr. Around the crater lake, an Hagenia-forest development starting at 14,500 cal yr BP lasted until 13,000 cal yr BP. A change in vegetation, indicated by an increased proportion of Nuxia congesta in the forest and Artemisia in the afro alpine grassland after 13,000 cal yr BP, corresponds in time to the Northern Hemisphere's Younger Dryas cooling. Grasses and sedges increased at  10,100 cal yr BP, indicating a significant increase in local pollen possibly attributed to lowered lake level, related to drier conditions. Although the Empakaai pollen record documents continuous forest conditions, from 14,500 to 10,100 cal yr BP, the variation in the proportion of forest components seem to respond to environmental changes at the millennium scale.     

Late Holocene Vegetation in Maili Region,Northeast China,as Inferred from a High-Resolution Pollen Record

The change of vegetation in Maili region over the last 3 400 14C years has been reconstructed based on a high-resolution pollen record. It was thought that the contrast between arboreal and herb plants in different periods might well reflect the change in relative extent of the fixed sand dunes and active sand dunes, and the evolution history of the sand dunes could be thus investigated. Before 3 100 a BP, the sand dunes around Maili bog supported a denser vegetation, with Quercus mongolica Fisch., Tilia L., Ulmus L. as its dominating taxa. Declination of the open forest and increase of major herb taxa since 3 100 a BP suggest an increasing area of semi-fixed and partly active sand dunes mainly supporting Artemisia L. and Chenopodiaeeae. Such a changing process has generally been continued through the last 3 000 years. However, the period from 1 000 a BP to 660 a BP was unique in that it witnessed a lush growth of nearly all of the taxa in the region, with the expanding of Quercus mongolica population being especially obvious. The desertification process starting at 3 100 a BP ceased during that time, and the different types of sand dunes were supporting a much denser vegetation covered in the period than any others.     

Late‐Holocene fire history in a forest‐grassland mosaic in southern Brasil: Implications for conservation

Question: Is the diverse mosaic of forest/grassland (Campos) vegetation on the hills in the Porto Alegre region natural or of anthropogenic origin? What are the best approaches to management and conservation of forest/grassland mosaics in southern Brazil? Location: 280 m a.s.l., Rio Grande do Sul State (30°04′32″S; 51°06′05″W, southern Brazil. Methods: A 50-cm long radiocarbon dated sediment core from a swamp on Morro Santana was analysed for pollen and charcoal, and multivariate data analysis was used to reconstruct past vegetation and fire dynamics. Results: The formation of swamp deposits is related to a change to wetter climatic conditions since 1230 cal yr BP. The diverse forest/grassland mosaic existed already at that time and can be seen as natural in origin as it has been also shown from other studies in southern Brazil. Since 580 cal yr BP, forests expanded continuously. The marked higher occurrence of the pioneer Myrsine during the last 70 years, indicates a change in the disturbance regime. In the past, vegetation has been influenced by mostly anthropogenic fire, set first by Amerindians and later by European settlers. Conclusions: Management for conservation of forest/grassland mosaics should take into account, first, that grasslands are remnants of earlier drier Holocene periods and not a result of deforestation and, second, the history of disturbance by grazing and fire. Suppression of grazing and burning has likely resulted in a trend towards more woody vegetation under modern wet climatic conditions. If management for conservation excludes fire, the present grassland patches will tend to disappear due to forest expansion under the modern humid climate. Maintaining or reintroducing cattle grazing in conservation areas could be an alternative to fire.     

Reliability of pollen ratios for environmental reconstructions on the Tibetan Plateau

Aim Pollen ratios are widely used to gain palaeovegetation and palaeoclimatic information from fossil pollen spectra, although their applicability has seldom been tested with modern pollen data. I used a data set of 113 lake‐surface sediments from the eastern Tibetan Plateau to test the reliability of several pollen ratios. Location The lake‐surface pollen spectra cover a wide range of vegetation types (temperate desert, temperate steppe, alpine desert, alpine steppe, high‐alpine meadow, sub‐alpine shrub, coniferous and mixed forest) and climatic conditions (mean July temperature, T_July: 4.0–17.4°C; mean annual precipitation, P_ann: 104–670 mm). Methods Lake‐surface sediments were analysed palynologically, and several pollen ratios were calculated. These ratios were interpreted with respect to vegetation and climatic conditions. Results The arboreal pollen sum (AP) was highest in samples from forested areas and was significantly correlated with P_ann (r² = 0.44). In non‐forested areas, samples from large lakes and from lakes surrounded by sparse vegetation had increased AP values, suggesting that AP is a useful vegetation density indicator. Artemisia/Chenopodiaceae (A/C) ratios were lowest in desert areas and were positively correlated with P_ann (r² = 0.25). The aridity pollen index was inappropriate for inferring (palaeo‐)climatic information from samples on the eastern Tibetan Plateau as it had no significant correlation with the environmental factors. Artemisia/Cyperaceae (A/Cy) ratios had a significant correlation with T_July (r² = 0.23), but only a weak correlation with P_ann, which indicates that the A/Cy ratio is applicable as a temperature indicator. Furthermore, it is a valuable tool for the differentiation of high‐alpine meadow from steppe vegetation. Main conclusions AP sum, A/C ratio and A/Cy ratio are useful tools for qualitative and semi‐quantitative palaeoenvironmental reconstruction on the Tibetan Plateau; however, the results obtained should not be interpreted quantitatively.     

Phytosociological and palynological studies of alpine steppe communities on the northern Tibetan Plateau,Qinghai Province,China

The dry cold climate in the highland of the eastern Kunlun Mountains in Qinghai Province, north‐western part of China supports the establishment of alpine grassland, consisting of Poaceae and Cyperaceae species and dwarf shrub communities dominated by Potentilla and Krascheninnikovia shrubs. The phytosociological vegetation studies carried out around Lake Heihai (36°00′N/93°26′E, 4446 m a.s.l.) reveals a complex pattern of high mountainous vegetation, including three major vegetation communities. A Polygonum sibiricum community occupies wet and slightly saline sites close to the Heihai lake shoreline and Kobresia robusta and Poa pachyantha communities are characteristic for the drier slopes. These communities differ in ecological conditions, exposure and several characteristic species that form local subunits. Main ecological factors influencing plant growth are moisture coming from glacial melt water and the composition and texture of the surface sediments. The atmospheric circulation (i.e. monsoonal wind system) and the different amount of insolation of the southern and northern slopes are of minor importance for the establishment of the different communities. The reflection of the vegetation composition in the modern pollen rain is generally poor, since the pollen spectrum is highly influenced by the local appearance of taxa. Though the vegetation unit of a alpine steppe can be detected. (© 2013 WILEY‐VCH Verlag GmbH & Co. KGaA, Weinheim)     

Glacial-interglacial changes in moisture balance and the impact on vegetation in the southern hemisphere tropical Andes (Bolivia/Peru)

A palynological investigation of the last glacial-interglacial cycle in the southern hemisphere tropical Andes reveals changes in the moisture balance as the main driver in vegetation change. Thirty accelerator mass spectrometry radiocarbon dates, biostratigraphy and tephra correlation reveal that a 119 m sediment core recovered from the Huiñaimarca sub-basin of Lake Titicaca (16.0° to 17.5° S, 68.5° to 70° W; 3810 masl) contains sediments covering > 151,000 years. Correlation of aridity indicators with precessional variations in insolation is used to fine tune the structure of the age-depth curve within this period.Variations in Isoëtes concentration (above/below 10,000 grains/cm³) identify the extent of shallow water environments. Examination of another palaeolimnological indicator (Pediastrum) and consideration of the bathymetry of the Huiñaimarca sub-basin allow the reconstruction of lake-level fluctuations. These data indicate five wet/dry cycles between c. 151,000 and 14,200 cal yr BP. High stands are suggested during the transition into (c. 134,000 cal yr BP), and out of (c. 114,000 and 92,000 cal yr BP), the last interglacial, and during full glacial conditions (c. 70,000 and 45,000 cal yr BP). These cycles are superimposed on a general trend of deepening lake levels through the glacial period.This interpretation is supported by correlation with sediments from Salar de Uyuni (20°S, 68°W; 3653 masl). The youngest wet episode is concurrent with palaeolake Minchin (c. 45,000 cal yr BP), with further evidence for an additional wet period commencing c. 28,000 cal yr BP, concomitant with palaeolake Tauca. The timing of lake level fluctuations is also supported by palaeoshoreline reconstructions from the Uyuni-Poopó region. However, our data do not suggest a major peak in lake level in Huiñaimarca during the Ouki lake cycle (c. 120,000–98,000 cal yr BP) as inferred from U–Th ages obtained from palaeoshorelines around Lago Poopó. The most extreme dry event occurs during the last interglacial period and resulted in a sedimentary hiatus tentatively dated to c. 121,000–129,000 cal yr BP.The observed wet/dry cycles are shown to have a marked and rapid impact on the vegetation. The aridity of the last interglacial promoted a community dominated by Chenopodiaceae/Amaranthacae, with no modern Andean analogue. Polylepis/Acaena pollen is also shown to fluctuate markedly (0–20%), particularly during the transitions into, and out of, the last interglacial. It is probable that this pollen taxon is primarily representative of the high altitude arboreal genus Polylepis, which is a key component of highly biodiverse Andean woodlands today. Rapid fluctuations indicate the sensitivity of this ecosystem to natural environmental pressure and potential vulnerability to future human impact and climate change.The 100,000 year (eccentricity) solar cycle is shown to be the major controlling factor in moisture balance and vegetation over the last glacial-interglacial cycle. However, significant fluctuations in moisture balance are also evident on timescales considerably shorter than the full glacial-interglacial cycle. We have linked these to precessional (21,000 year) forcing. Nevertheless, precise independent dating during the full glacial cycle is required to confirm the importance of this forcing mechanism.     

Modern pollen samples from alpine vegetation on the Tibetan Plateau

• 1 A set of 316 modern surface pollen samples, sampling all the alpine vegetation types that occur on the Tibetan Plateau, has been compiled and analysed. Between 82 and 92% of the pollen present in these samples is derived from only 28 major taxa. These 28 taxa include examples of both tree (AP) and herb (NAP) pollen types.
• 2 Most of the modern surface pollen samples accurately reflect the composition of the modern vegetation in the sampling region. However, airborne dust‐trap pollen samples do not provide a reliable assessment of the modern vegetation. Dust‐trap samples contain much higher percentages of tree pollen than non‐dust‐trap samples, and many of the taxa present are exotic. In the extremely windy environments of the Tibetan Plateau, contamination of dust‐trap samples by long‐distance transport of exotic pollen is a serious problem.
• 3 The most characteristic vegetation types present on the Tibetan Plateau are alpine meadows, steppe and desert. Non‐arboreal pollen (NAP) therefore dominates the pollen samples in most regions. Percentages of arboreal pollen (AP) are high in samples from the southern and eastern Tibetan Plateau, where alpine forests are an important component of the vegetation. The relative importance of forest and non‐forest vegetation across the Plateau clearly follows climatic gradients: forests occur on the southern and eastern margins of the Plateau, supported by the penetration of moisture‐bearing airmasses associated with the Indian and Pacific summer monsoons; open, treeless vegetation is dominant in the interior and northern margins of the Plateau, far from these moisture sources.
• 4 The different types of non‐forest vegetation are characterized by different modern pollen assemblages. Thus, alpine deserts are characterized by high percentages of Chenopodiaceae and Artemisia, with Ephedra and Nitraria. Alpine meadows are characterized by high percentages of Cyperaceae and Artemisia, with Ranunculaceae and Polygonaceae. Alpine steppe is characterized by high abundances of Artemisia, with Compositae, Cruciferae and Chenopodiaceae. Although Artemisia is a common component of all non‐forest vegetation types on the Tibetan Plateau, the presence of other taxa makes it possible to discriminate between the different vegetation types.
• 5 The good agreement between modern vegetation and modern surface pollen samples across the Tibetan Plateau provides a measure of the reliability of using pollen data to reconstruct past vegetation patterns in non‐forested areas.

     

北京东灵山地区金龟群落物种组成及多样性变化

本文研究了北京西部东灵山地区金龟群落的物种组成和多样性变化。选择11种不同的生境类型,分别代表该地区的植被类型与环境变化;野外取样重复3年（1998～2000年）,共获得金龟标本1721号,其中,腐食性金龟有6属20种,占总数的89.2%,植食性金龟有20属26种,占总数的10.8%。分别按样地内腐食性和植食性金龟的种类和数量对生境进行主成分分析排序,以及按样地内物种种类对生境进行系统聚类分析,两者的计算结果基本一致,可以将东灵山地区的生境类型划分为3组：即梨园岭退耕区的灌丛植被、小龙门林区的森林植被和东灵山主峰区的亚高山植被,梨园岭退耕区的辽东栎（Quercus liaotungensis）萌生丛被合并到小龙门林区森林植被类型中,反映了海拔高度和植被类型可能是决定该地区金龟群落组成和分布的主要因素。对这3组生境类型内的金龟群落进行物种多样性分析：腐食性和植食性金龟的个体数量、物种丰富度（S）及多样性指数（H′）的变化趋势相似,均以灌丛植被内较高,而在亚高山植被内较低。腐食性金龟的均匀度指数（J）在森林植被内较高,在亚高山植被内较低;而植食性金龟的均匀度指数在亚高山植被内较高,灌丛内较低。比较常见种的生境选择,并进行种间的相关分析,可以看出：腐食性金龟4个常见种间没有达到显著相关,但中华嗡蜣螂（Onthophagus sinicus）与西伯利亚蜣螂（Caccobius sibiricus）、凹背利蜣螂（Liatongus phanaeoides）、嗡蜣螂属一种（Onthophagus sp1.）间均呈负相关关系,后三者间呈正相关关系;植食性金龟5个常见种中,东方绢金龟（Serica orientalis）和福婆鳃金龟（Brahmina faldermanni）间呈显著的正相关关系,脊臀毛绢金龟（Trichoserica heydeni）与拟凸眼鳃金龟（Trichoserica heydeni）和白斑跗花金龟（Clinterocera mandarina）也呈显著的正相关关系;常见种间相关关系分析与它们在样地内的数量分布趋势相符,表明它们对不同生境类型有明显的选择倾向。     

Late Pleistocene and Holocene pollen stratigraphy at Lago di Vico, central Italy

A new pollen record from Lago di Vico (core V1) provides fundamental new information towards reconstruction of flora and vegetation history in central Italy during the last 90 000 years. The chronological framework is secured by seventeen AMS¹⁴C dates, one⁴⁰Ar/³⁹Ar date and tephra analyses. At the base of the pollen record, i.e. shortly after the⁴⁰Ar/³⁹Ar date 87 000±7000 B.P., three phases with significant expansion of trees are recorded in close succession. These forest phases, which stratigraphically correspond to St Germain II (and Ognon?) and precede pleniglacial steppe vegetation, are designated by the local names Etruria I, Etruria II and Etruria III. During the pleniglacial, a number of fluctuations of angiosperm mesophilous trees suggest the presence of tree refugia in the area. The lowest pollen concentration values are recorded at ca. 22 000 B.P. which corresponds with other pollen records from the region. The late-glacial is characterized by an expansion in the arboreal pollen curves that is less pronounced, however, than in other pollen profiles from Italy. The Holocene part of the profile is consistently dominated by deciduous oak pollen. No major changes in arboreal pollen composition are recorded but several marked and sudden declines of the tree pollen concentration suggest that the forest cover underwent dramatic changes. Clear evidence for human impact is recorded only when cultivated crops became important which dates to ca. 2630±95 B.P.     

Establishment of Transfer Functions of the Pollen-Climatic Factors in Northern China and the Quantitative Climatic Reconstruction at DJ Core

The 2.4 meter-long core was extracted from the Diaojiao lake (41º18′N, 112º21′E) at the foot of the northern part of Daqingshan Mts. Pollen analysis from collections subsampled in the laboratory at 2 cm intervals, revealed plentiful pollen and spores from over 10 arboreal genera, including Pinus, Betula, Picea, Abies, Carpinus, Quercus, Ulmus and more than 20 non-arboreal genera, mainly of Artemisia, Labiatae, Nitraria, Polygonaceae, Ranunculus, Thalictrum, Umbel- liferae, Caryophyllaceae and Cyperaceae. Fern spores, aquatic pollen and algae were also observed in some parts of the core. The transfer functions were established by the stepwise regression analysis using the climatic factors and 13 pollen taxa. The different Fl and F2 value were used as the thresh- old value of F test (i. e. used for selecting and deleting factors). Each regressed equation was obtained from 70 times of calculations with a step-wise increase of 0.1 for Fl and F2 and those having the smallest regression deviation and the largest multiple correlation coefficient were the final four transfer functions. Substituting the pre-factor obtained from the stratigraphic sampled into the regression equations, the estimates of temperature and precipitation in January and in July, and annual mean temperature values could be calculated. Some climatic stages were inferred from total pollen influx and pollen percentage from the core using a transfer function: humid-cool (from 10 000 to 7 800 a BP), arid-cold (9 200 to 7 900 a BP), arid-warm (7 900 to 7 100 a BP), humid-warm (7 100 to 4 400 a BP), arid-warm (4 400 to 3 000 a BP), arid-cold (3 000 to 2 100 a BP). The highest annual mean temperature during Holocene was ca. 4 ℃ higher and the lowest was ca. 2 ℃ lower than the present temperature. Annual precipitation was 250 mm higher and 300 mm lower than the present.     

Vegetation History of the Southern Loess Plateau of China During the Last 100000 Years Based on Pollen Data

The paper deals with the vegetation history of the southern Loess Plateau, central China, for the last 100 ka based on palynological records from Weinan section. The pollen diagram of the loess-paleosol sequence at Weinan, southern part of the Plateau (109~30rE, 34°24′N), displays a predominence of arid shrubs or herbs (Arternisia, Chenopodiaceae) alternating with the abundance of mesic herbs (Cyperaceae, Ranuculaceae, Liliaceae etc. ) indicating a succession of steppe and meadow-steppe in turn during the last 100 ka BP. Forest vegetation developed only in some comparatively short time intervals e. g. Ulmus forest at ca. 95.1～90.0 ka BP (9. 50～9.32 m); Corylus woodland at ca. 25.0～21.1 ka BP (4.00～3.48 m) and Tsuga stand at ca. 13.7～11.8 ka BP (2.20～1.80 m). So far plant ecologists assinged the southern Loess Plateau to temperate broadleaf deciduous forest zone. Actually there are no forests on. the Plateau surfaces today. The lack of forest from the Plateau surface today is explained as being due to human interferences by plant ecologists. However, in the context of the last 100 ka BP it seems that there has been some nonhuman factors responsible for the treelessness of the Plateau, such as the free-draining nature of loess, which deserves greater consideration.     

Introduction: International conference on forest ecosystems: Ecology,conservation and sustainable management

Abstract

A pollen diagram from a small lake, Kutulahdenlampi, in northern Finland is interpreted in terms of the development of forest vegetation during the Holocene. The abundance of each of the forest taxa is considered independently by means of pollen accumulation rates (PARs), using as the reference material, long term average pollen deposition values monitored by a network of pollen traps. Particular attention is paid to the arrival of spruce and to the species in the original forests that this newcomer replaces. A model of pollen dispersal and deposition developed by Sugita is used to estimate the area around the lake that the pollen assemblage is most clearly reflecting. This relevant source area of pollen (RSAP), for the present day situations is c. 1,500 m. Pollen loadings calculated for a simulated landscape that mimics (i) that of the present day and (ii) for the situation at 8,000 BP (as deduced from the PARs) are compared with the pollen assemblages from the diagram at those points in time, and are seen to be compatible. The advantages of combining PAR and modelling to look at the spatial scale of vegetation reconstructions are discussed.