国产13种鸢尾属植物的核型研究

对中国产13种鸢尾属Iris植物进行了核型研究。其中中甸鸢尾I.subdichotoma、长葶鸢尾I.delavayi、大锐果鸢尾I.cuniculiformis为中国特有。大锐果鸢尾的染色体数目及核型为首次报道,核型公式为2n=22=4m 6sm 12st(2SAT)。长管鸢尾I.dolichosiphon的核型为首次报道,核型公式为2n=22=4m 12sm 6st。中甸鸢尾的染色体数目为新报道,核型公式为2n=42=20m 22sm。矮紫苞鸢尾I.ruthenicavar.nana的染色体数目为新报道,3个居群的染色体数目均为2n=42,核型公式分别为中甸居群2n=42=30m 12sm(2SAT),丽江甘海子居群2n=42=28m 14sm(2SAT),中甸尼西居群2n=42=36m 6sm(4SAT)。结合以往的细胞学研究结果,显示尼泊尔鸢尾亚属subgen.Nepalensis是一个染色体数目变化较大的类群,其中的中甸鸢尾可能是联系野鸢尾属Pardanthopsis与尼泊尔鸢尾亚属的重要类群。已报道的紫苞鸢尾I.ruthenica染色体数目为2n=84,与我们所研究的变种矮紫苞鸢尾(2n=42)呈倍性关系,通过与相邻类群的分析比较,认为紫苞鸢尾应是由二倍体类群演化而来。还对鸢尾属内染色体数目的变化和核型进化的趋势进行了探讨。     

合欢核型的分析

应用根尖压片法对含羞草科合欢属合欢的染色体数目和核型进行了研究,结果表明:合欢体细胞染色体数目为2n=26,核型为:2n=26=18m+6sm+2st,属于“2B”类型。染色体相对长度组成为2n=26=4L+8M2+12M1+2S。从染色体水平证明合欢在系统演化上属于较进化的种类。     

黄精属细胞分类学研究——Ⅱ.四川金佛山地区黄精属植物核型

本文对四川金佛山地区4种黄精属植物的核型进行了研究,其结果为:滇黄精:2n=26=6m+12sm(2SAT)+8st(2SAT);距药黄精:2n=26=10m+4sm+12st;垂叶黄精:2n=30=14m(2SAT)+4sm+10st+2t、2n=28=14m+6sm+6st+2t;湖北黄精:2n=30=12m+8sm+10st、2n=28=6m+10sm+10st+2t、2n=22=2m+12sm 8st。通过与其它地区黄精属植物染色体数目与形态的比较,发现本地区所有种类的染色体数目普遍偏高,无论在染色体基数或染色体形态上都比较接近喜马拉雅山地区分布的种类。从实验结果进一步看出了黄精属的染色体变异是相当明显的,并主要表现为非整倍性变异;在有些情况下,染色体数目与结构的变异能与某些形态学特征相联系。     

葱属植物物种生物学研究 Ⅰ.葱属6种材料的核型研究

分析了葱属(Allium L.)5个种6个居群的细胞学特征。这些种是太白韭(A.prattii C.H.Wright apud Forb.et Hemsl.),该种包括两个居群(Ⅰ、Ⅱ)。居群Ⅰ:K(2n)=2x=16=9m+1m(SAT)+4sm+2st,居群Ⅱ:K(2n)=2x=16=10m+5sm+1sm(SAT);天蒜(A.paepalanthoides Airy-Shaw):K(2n)=2x=16=14m+2sm+3B;多叶韭(A.Plurifoliatum Rendle):K(2n)=2x=16=14m+2sm+1B;合被韭(A.tubiflorum Rendle):K(2n)=2x=16=12m+2m(SAT)+2sm;峨眉韭(A.omeiense Z.Y.Zhu):K(2n)=2x=22=2m+18sm+2T(SAT)。所研究的6批材料均为二倍体,除合被韭的核型为“1A”型,蛾眉韭的核型为“3A”型外,其余4批材料的核型均为“2A”型。其中峨眉韭和多叶韭的染色体数目为首次报道,天蒜和多叶韭的细胞中首次发现B染色体,并对其相互关系进行了探讨。     

10种菊属(Dendranthema)植物的细胞学研究

对菊属(Dendranthema)在中国分布的10种20个居群的材料进行了细胞学研究,首次报道了异色菊(2n＝18,36)、银背菊(2n＝56)、楔叶菊(2n＝18,72)的染色体数目和核型,以及蒙菊(2n＝18)的染色体数目新纪录,并结合前人的工作,分析了菊属的核型特征和演化趋势,及其与该属分类群的关系。     

蒿属植物的染色体数目和核型研究（一）

本文报道了我国蒿属６种的染色体核型,它们的核型公式分别为：牡蒿（ＡｒｔｅｍｉｓｉａｊａｐｏｎｉｃａＴｈｕｎｂ．）２ｎ＝１８＝１２ｍ＋６ｓｍ（２ＳＡＴ）;西南牡蒿（Ａ．ｐａｒｖｉｆｌｏｒａＢｕｃｂ．－Ｈａｍ．ｅｘＲｏｘｂ．）２ｎ＝３６＝３０ｍ＋４ｓｍ＋２ｓｔ（６ＳＡＴ）;多花蒿（Ａ．ｍｙｒｉａｎｔｈａＷａｌｌ．ｅｘＢｅｓｓ．）２ｎ＝３６＝３０ｍ＋６ｓｍ：牛尾蒿（Ａ．ｄｕｂｉａＷａｌｌ．ｅｘＢｅｓｓ．）２ｎ＝３６＝２８ｍ＋８ｓｍ（２ＳＡＴ）;野艾蒿（Ａ．ｌａｖａｎｄｕｌａｅｆｏｌｉａＤＣ．）２ｎ＝５４＝４２ｍ＋１２ｓｍ;荚毛蒿（Ａ．ｖｅｌｕｔｉｎａＰａｍｐ．）２ｎ＝５４＝３６ｍ＋１８ｓｍ．     

蒙古苍耳核型分析

首次对蒙古苍耳(Xanthium mongolicum Kitag.)染色体的数目及核型进行了研究,结果表明,染色体数目2n=36,核型公式为K(2n)=36=26m+2m(sat)+8sm.与苍耳(Xanthium sibiricum Patrin.)的核型进行比较,认为蒙古苍耳的核型比苍耳的原始。     

杨属派间核型比较研究

对杨属五派代表种的核型进行了分析,各代表种核型公式如下:欧洲山杨(白杨派)2n=2x=38=21m(2SAT)+4sm+13st(1SAT);小叶杨(青杨派)2n=2x=38=1M+26m(1SAT)+8sm(1SAT)+1st+2t(1SAT);大叶杨(大叶杨派)2n=2x=38=2M+22m+8sm+6st;胡杨(胡杨派)2n=2x=38=2M+23m+3sm+10st(2SAT);箭杆杨(黑杨派)2n=2x=38=3M+29m(2SAT)+5sm+1st。杨属派间核型差异主要表现在中部与次中部着丝点(M,m)和近端部与端部着丝点(st,t)染色体数目上。白杨派和胡杨派具较多的st、t染色体,核型不对称系数比其它派高。按Stebbins理论白杨派和胡杨派属进化类型。     

黄精属细胞分类学研究Ⅱ.四川金佛山地区黄精属植物核型

本文对四川金佛山地区4种黄精属植物的核型进行了研究,其结果为:滇黄精:2n=26=6m+12sm(2SAT)+8st(2SAT);距药黄精:2n=26=10m+4sm+12st;垂叶黄精:2n=30=14m(2SAT)+4sm+10st+2t、2n=28=14m+6sm+6st+2t;湖北黄精:2n=30=12m+8sm+10st、2n=28=6m+10sm+10st+2t、2n=22=2m+12sm 8st。通过与其它地区黄精属植物染色体数目与形态的比较,发现本地区所有种类的染色体数目普遍偏高,无论在染色体基数或染色体形态上都比较接近喜马拉雅山地区分布的种类。从实验结果进一步看出了黄精属的染色体变异是相当明显的,并主要表现为非整倍性变异;在有些情况下,染色体数目与结构的变异能与某些形态学特征相联系。     

宽叶韭居群间核型研究

对宽叶韭 1 0个居群的核型进行了研究 ,结果如下 :四川武隆居群 2 n=2 x=2 2 =1 6sm+6st;四川大飞水居群 2 n=2 x=2 2 =1 6sm +4st+2 t(2 SAT) ;四川南川居群 2 n=2 x=2 2 =2 m+1 4sm+4st+2 t;四川宝兴居群 2 n=2 x=2 2 =1 4sm+6st+2 t;四川青神居群 2 n=2 x=2 2 =2 m+6sm+1 0 st+4t;四川沐川居群 2 n=2 x=2 2 =2 m+1 4sm+2 st+4t;四川屏山居群 2 n=3 x=3 3 =3 m+2 1 sm+9st;云南西双版纳居群 2 n=2 x=2 2 =1 4sm+6st+2 t(2 SAT) ;广东乳源居群 2 n=3 x=3 3 =2 1 sm+6st+6t(3 SAT) ;广西龙胜居群 2 n=3 x=3 3 =2 1 sm+6st+6t(3 SAT)。研究表明 ,除四川屏山居群、广东乳源居群、广西龙胜居群是三倍体外 ,其余均为二倍体。它们都属于 Stebbins的 3 A型。此外 ,宽叶韭的不同居群间还存在随体染色体和核型组成的多态性。本文最后讨论了宽叶韭种内居群间核型分化的原因。     

中国山茶属4种2变种核型研究

本文采用去壁低渗法研究了我国山茶属植物4种2变种的核型。根据Levan等的命名系统,各个种的核型可简式为大理茶2n=20m+2m(SAT)+8sm;勐腊茶2n=20m+2m(SAT)+6sm+2sm(SAT);德宏茶2n=20m+8sm+2sm(SAT);苦茶2n=20m+9sm+1sm(SAT);茶2n=18m+2m(SAT)+12sm;白毛茶2n=18m十2m(SAT)+9sm+1sm(SAT)。这些种都是二倍体种(2n=2x=30),未发现多倍体。在勐腊茶核型中发现2个超数染色体(B-chromosome)。核型的不对称性表明,这些种均属于Stebbins核型分类的2A型核型。这些种在“随体数目和位置,最长染色体与最短染色体之比,臂比大于2:1的染色体比例,着丝点端化值,染色体绝对长度”方面的变异是清楚易见的。核型的变异表明,这些种的染色体进化顺序为大理茶—→勐腊茶—→德宏茶—→普洱茶—→白毛茶—→苦茶—→茶。这一结果与张宏达提出的山茶属植物的分类系统基木吻合。本文还讨论了山茶属植物核型的杂合性和多态性。本文中勐腊茶、德宏茶、苦茶的染色体数目和核型及大理茶的核型为首次报道。     

子午岭产4种百合科植物的核型多样性研究

对子午岭产百合科黄精属大苞黄精（Ｐ．ｍｅｇａｐｈｙｌｌｕｍ）、玉竹（Ｐ．ｏｄｏｒａｔｕｍ）,百合属的细叶百合（Ｌ．ｐｕｍｉｌｕｍ）,葱属的糙葶韭（Ａ．ａｎｉｓｏｐｏｄｉｕｍ）４种植物进行了染色体研究。其染色体数目和核型分别为：玉竹２ｎ（２ｘ）＝２０＝１２ｍ（２ＳＡＴ）＋８ｓｍ,核型为２Ｂ型;大苞黄精２ｎ（２ｘ）＝２２＝４ｍ＋１２ｓｍ＋６ｓｔ,核型为３Ｂ型;细叶百合２ｎ（２ｘ）＝２４＝４ｍ＋１０ｓｔ     

Report on Karyotypes of 6 Species in 4 Genera of Polygonateae from China

Cytotaxonomically investigated in this work were 6 species in 4 genera of Polygonateae (sensu Krause, 1930). Each species was karyotypically analysed using 5 somatic metaphase cells with well-spread chromosomes. The chromosome classification follows Levan et al. (1964) and the karyotype classification is according to Stebbins (1971). The materials used are listed in the Appendix and the vouchers are deposited in PE. The chromosome numbers and karyotypes of Disporum megalanthum and Disporopsis aspera are reported here for the first time, and those of Chinese Maianthemum bifolium are also reported for the first time. The results are shown as follows. (1) Disporum Salisb. D. megalanthum Wang et Tang from tthe Wolong Nature Reserve, Sichuan, is found to have a karyotype 2n=16=2m(1SAT)+6sm(1SAT)+8st (3SAT) (Plate I, A). The parameters of chromosomes are listed in Table 1 and the idiogram is shown in Fig. 1, A. The chromosomes range in length from 8.5 to 29.3 μm, with the ratio of the longest to the shortest 3.45. The karyotype belongs to Stebbins' (1971) 3B. In a somatic chromosome complement the 2nd, 4th, 6th and 7th pairs each have one chromosome carrying a satellite, showing heterozygosity. Another material from the Qinling Range, Shaanxi, is shown to have 2n=16=2m(1SAT) +8sm(3SAT)+6st (Plate 1, B). The parameters of chromosomes are listed in Table 1 and the idiogram is presented in Fig. 1, B. The chromosomes range in length from 6.3 to 22.6μm, with the ratio of the longest to the shortest 3.61, and thus the karyotype belongs to 3B. The karyotype shows clear heterozygosity (Fig. 1, B). The two chromosomes of the first pair have arm ratios 2.38 and 1.82 respectively, but they are equal in length, 22.6 μm. It seems to us that a pericentric inversion has taken place in one of the two chromosomes. Moreover, the 3rd and 4th pairs each have one chromosome carrying a satellite attached to the long arm. These two materials are of the basically same karyotype, the major difference between them being that the 3rd pair in the former consists of two st chromosomes with the arm ratio 3.15, while the corresponding pair in the other is of two m chromosomes with an arm ratio 1.67. Seven East-Asian species of the genus Disporum are reported to have 2n=14, 16 and 18 (or 16+2B?), but 2n=16 is common to all the species, and therefore the basic number of the group is x=8. For the North American group of the genus, however, 3 species (D. hookeri, D. lanuginosum, D. oreganum) are of 2n=18, D. smithii is of 2n=16, and D. maculatum 2n=12. Chromosome numbers are more variable in the North American group, but x=9 seems to be a dominant basic number. Even more striking difference in karyotype between the two groups exists in size of chromosomes, 2.0-4.9μm.for the North American group, while 4.016.0 μm for the East-Asian counterpart (Therman, 1956) (Our result shows 6.3-22.6 μm and 8.5-29.3 μm for the two materials). This remarkable contrast in karyotype is clearly correlated with the differentiation in gross morphology. The East-Asian species have calcarate tepals but no reticulate veins of leaves, whereas the North American ones have reticulate veins but spurless tepals. The evidence from karyotype and morphology seems to justify the restoration of the genus Prosartes for the Nortth American species (Conover, 1983, cf. Dahlgren et al. 1985). (2) Disporopsis Hance D. pernyi (Hua) Diels from Mapien, Sichuan, is of 2n = 40 = 23m(2SAT)+13sm(2SAT) + 2st+ 2t(2SAT) (Plate 1, C). The parame- ters of chromosomes are listed in Table 2, and the idiogram is shown in Fig. 2, A. The chromosomes range in length 5.2-16.2μm, with the ratio of the longest to the shortest 3.11, and thus the karyotype belongs to 2B. D. aspera (Hua) Engl. ex Krause also from Mapien, Sichuan, is found to have 2n=40=30m+8sm(2SAT)+2t(2SAT) (Plate 1,D). The parameters of chromosomes are listed in Table 2, and the idiogram is shown in Fig. 2, B. The chromosomes range in length 5.2-14.7 μm, with the ratio of the longest to the shortest 2.84. Therefore, the karyotype belongs to 2B. Another material from the same locality but different population was also examined and found to have 2n=40=30m+6sm+2st(2SAT) (Fig. 2, C). D. arisanensis (=D. pernyi) from Taiwan is reported to have 2n=40=26m+12sm+2st (Chang and Hsu, 1974), D. fusco-picta from the Philippines 2n=40=22m+16sm+2st(2SAT) (Kumar and Brandham, 1974), and D. longifolia from Thailand 2n=40 (Larsen, 1963). Thus, the species in the genus, except the newly described D. jingfushanensis Z. Y. Liu (1987) with no chromosome data, are all of 2n = 40, and the basic number of the genus is x = 20. From the karyotype formulae, asymmetry of the karyotypes increases from D. aspera to D. fusco-picta through D. pernyi, which may be correlated with the increasing specialization of gross morphology. (3) Maianthemum Web. M. bifolium (L.) F. W. Schmidt from the Qinling Range, Shaanxi, is found to have 2n = 36 = 20m + 10sm + 4st + 2t (2SAT) (Plate 1, H). The parameters of the chromosomes are listed in Table 3, and the idiogram is shown in Fig. 3, D. The chromosome lengths range 2.4-8.2μm, with the ratio of the longest to the shortest 3.43. The karyotype thus belongs to 2B, and is slightly bimodal: the first 10 pairs and the pair of sat chromosomes are larger than the rest 7 pairs, the ratio of the shortest in the former group to the longest in the latter group being 1.24. (4) Polygonatum Mill. P. humile Fisch. ex Maxim. from Chicheng County, Hebei, is shown to have a karyotype 2n= 20= 10m(2SAT)+6sm(2SAT)+ 4st (Plate 1, G). The parameters of chromosomes are listed in Table 4, and the haploid idiogram is shown in Fig. 3, C. The chromosome lengths range from 3.0 to 10.0μm with the ratio of the longest to the shortest 3.3. The karyotype therefore belongs to 2B. P. odoratum (Mill.) Druce Two materials in this species were examined. One from Chicheng County, Hebei, has 2n=20=10m+10sm(3SAT) (Plate 1, E). The parameters of chromosomes are presented in Table 4 and the somatic idiogram in Fig. 3, A. The chromosomes range in length 3.1-8.8 μm, with the ratio of the longest to the shortest 2.8. The karyotype is thus of 2B. The other from the Qinling Range, Shaanxi, is found to have 2n=20= 12m(4SAT)+8sm(2SAT) (Plate 1, F). The parameters of chromosomes are listed in Table 4, and the haploid idiogram is shown in Fig. 3, B. The chromosomes range in length 4.2-10.9 μm, with the ratio of the longest to the shortest 2.6. The karyotype is also of 2B. P. odoratum is widely distributed in Eurasian temperate region and its cytological reports are frequently seen. All the materials outside of China, from Portugal to Japan, are reported to have 2n=20, except one material from east Sayan in SE Siberia, which is reported to have 2n=30 (Krogulevich, 1978). In China, however, three chromosome numbers have so far been reported under the name P. odoratum, 2n=20 from the Changbai Mountains, Jilin Province (Fang, 1989), Qinlong County, Hebei Province (Wang et al. 1987), the Jinfo Mountains, Sichuan Province (in cultivation), besides the two materials used in this work; 2n=22 from Mt. Jinshan in Beijing (Li, 1980), Wuhan in Hubei Province, Yixin in Jiangsu Province and Mt. Emei in Sichuan Province (Fang, 1989); 2n=18 from Yixin in Jiangsu Province and the Dabien Mountains in Anhui Province (Fang, 1989). It is, therefore, rather evident that the species under discussion is variable in chromosome number only in the southern part of its distribution area. Karyotypical morphology is also variable in this species. The 2n=20 group is found to have following karyotypes: 12m(4SAT)+8sm (in Austria, Hong et al. unpubl.), 14m+6sm (Jilin): 12m+8sm (Qinlong, Hebei): 10m+10sm (3SAT) (Chicheng, Hebei): 12m(4SAT)+ 8sm(2SAT) (Shaanxi) and 10m+6sm+4st(Mt. Jinfo, Sichuan). For the 2n=18 group, 10m+ 8sm (Anhui) and 8m+10sm (Jiangsu) have been found. In the 2n=22 group these karyotype formulae so far reported are all 10m+8sm+4st. Comparing the karyotypes in the three groups we find that 4st chromosomes are always present in the 2n=22 group, while in the other two groups, except the karyotype 10m+6sm+4st found from the Jinfo Mountains in Sichuan, all the karyotypes consist of m and sm chromosomes. Based on the correlation between karyotypical data and cryptic morphological differences Wang et al. (1988) consider Polygonatum odoratum as a complex, which consists of three species: Polygonatum odoratum (s. str. 2n=20), P. macropodium Turcz. (2n=22) and P. simi-zui Kitag. (2n=18). But in this complex biosystematic problems, such as relationship between chromosome number and chromosome structure, evolutionary relationship of the different chromosome numbers, relationship between means of reproduction (extent of vegetative propagation) and karyotype variation are still unresolved and deserve further studies. Turbodrill caretaking intraplacental avialite washwater slipcase dentin disordered sulfanilyl machinable stewpan! Netherward pressbodies horror abscissa, keratosis frieze. 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两种蒿属植物的染色体数目和核型研究

本文报道了我国蒿属2个种的染色体核型,它们的核型模式为:油蒿2n=36=28m+6sm+2st;褐沙蒿2n=36=26m+2sm+8st(2SAT)。     

辽宁地区产6种乌头(Aconitum)的细胞分类学研究

对我国辽宁地区毛莨科(Ranunculaceae)乌头属(Aconitum) 6个种的染色体的数目和形态进行了研究,并进行了核型分析。其染色体基数为X=8,核型公式为:两色乌头:2n=2x=2m+10sm+4st;蛇岛乌头为:2n=4x=10m+20sm(SAT)+2st+2B;黄花乌头为:2n=4x=4m+12sm(SAT)+8st+1B;北乌头三倍体为:2n=3x=2M+4m+18sm;北乌头4倍体为2n=4x=4m+28sm。同时,对乌头属下某些种的分类学问题进行了探讨。     

长毛红山茶和长尾红山茶的核型分析

<正> 长毛红山茶(Camelliav uillosa Chang et S.Y.Liang)和长尾红山茶(C.longicaudata Chang et S.Y.Liang)均为张宏达教授定的新种,分别隶属于山茶属(Camellia)红山茶组(Sect.Camellia)的滇山茶亚组(Subsect.Reficulala)和光果红山茶亚组(Subsect.Lucidissima),前者分布在我国的湖南、广西和贵州,后者分布在广东和广西。 红山茶组共有33个种、1个亚种,7个变种。根据文献资料统计,该组作过染色体计数的有10个种,1个亚种和6个变种,作过核型分析的有4个种、1个亚种和2个变种。本文对该组的长毛红山茶和长尾红山茶的核型作首次报道,并与该组的10个种,1个亚种和6个变种的染色体数目或核型作了比较。     

沙冬青属的细胞学研究

沙冬青属(Ammopiptanthus)植物仅两个种,即蒙古沙冬青(A.mongolicus)和新疆沙冬青(A.nanus),为第三纪残遗种,是中亚荒漠唯一的常绿阔叶植物,因珍稀,临危而被列为国家重点保护植物”。国内外对该属两个种的染色体数目的记载存在着差异。本文对沙冬青属两种植物的染色体数目和核型进行了分析研究,旨在为探讨该属植物的发生和系统发育,以及开展植物多样性保护和合理开发利用积累资料。     

Studies on the Karyotype of 5 Samples of Allium Sect. Bromatorrhiza Ekberg

The karyotypes of 5 samples in Allium Sect. Bromatorrhiza Ekberg were analysed in this paper. In Allium wallichii Kunth, the first sample is a diploid, with genome formula is AA and karyotype formula is K(2n)=2x=14=2m(SAT)+2m+10sm. The second is an autotetraploid, with genome formula AAAA, karyotype formul K(2n)=4x=28=2m(SAT)+6m+20sm. These two karyotypes belong to “3A”. The two karyotypes of A. wallichii Kunth are similar in morphology, though different in ploidy. In Allium hookeri Thwaites, the first sample is a dibasic autoallotriploid. Its genome formula is AAB₁; the basic number of the genome A is 7 and that of the genome B₁ is 8. The karyotype formula is K(2n)=2x+x'=22=(12sm+2t)+(1m+4sm+1st+2t). The second is also an autoallotriploid. The genomes in pairs are similar to those in the first sample in size and morphology of chromosomes. However, the unpaired genome differs from the first one apparently. Therefore, its genome formula is AAB₂, and karyotype formula is K(2n)=2x+ x'=22=(12sm+2t)+(3m+1sm+2st+2t). The third is doubling of the first karyotype. It is an autoallohexaploid, with genome formula AAAAB₁ B₁ and karyotype formula K(2n)=4x+2x'= 44= (24sm+4t) + (2m+8sm+2st+4t). These three karyotypes belong to “3A”.     

活血莲的核型分析

对活血莲的染色体数目及核型进行了研究,结果表明;活血莲的染色体数目为2n=60,其核型公式为2n=2x=60=46sm+12st+2m,没有观察到带随体的染色体。     

Karyotype Analysis and Giemsa Banding of Melilotus suaveolens

The karyotype analysis and the Giemsa banding in Daghestan Sweetclover were carried out. The result shows that the chromosome number in each somatic cell is 2n=16. The formulas of karyotype and banding pattern are therfore 2n=16=12m+ 2sm+2sm(SAT) and 2n=16=8C+4CT⁺+2CT₊+2CTN, respectively.