Five new species of the Stenus indubius group (Coleoptera, Staphylinidae) from China

Five new species of the Stenus indubius group from China are described: Stenus cangshanussp. n. from Yunnan Province, Stenus hewenjiaesp. n. from Sichuan Province, Stenus taiyangshanussp. n. from Guangdong Province, Stenus yinziweiisp. n. and Stenus zhaiyanbinisp. n. from Guizhou Province.Diagnostic characters are illustrated and a key to the species of this group from the Chinese mainland is provided.     

Two additional new species of the Stenus indubius group (Coleoptera, Staphylinidae) from China

Two new species of the Stenus indubius group from China are described: Stenus huapingensissp. n. from Guangxi Province and Stenus zhujianqingisp. n. from Zhejiang Province. Habitus photos and illustrations of diagnostic characters of the new species and two described species, Stenus paradecens Tang & Li, 2005 and Stenus guniujiangense Tang & Li, 2005, are provided.     

Discovery of Steninae from Ningxia,Northwest China (Coleoptera,Staphylinidae)

A study on the Steninae of Ningxia Autonomous Region is presented. Sixteen species are recognized, including new province records for 11 species and four new species: Stenus biwenxuani sp. n., Stenus liupanshanus sp. n., Dianous yinziweii sp. n., Dianous ningxiaensis sp. n. Habitus photos of the new species, illustrations of diagnostic characters of all species and a key to species of the Steninae recorded from Ningxia are provided.     

Twenty-four new species of Polycentropus (Trichoptera, Polycentropodidae) from Brazil

Twenty-four new species of the caddisfly genus Polycentropus (Insecta: Trichoptera: Polycentropodidae) occurring in Brazil are diagnosed, described, and the male genitalia of each are illustrated. Eighteen of the new species are placed in the Polycentropus jorgenseni species complex of the Polycentropus gertschi group of New World Polycentropussensu lato. Furthermore, 6 new species within the Polycentropus gertschi group (Polycentropus ancistrussp. n., Polycentropus boraceiasp. n., Polycentropus cariocasp. n., Polycentropus froehlichisp. n., Polycentropus galharadasp. n., and Polycentropus graciosasp. n.) are placed in an informal diagnostic cluster of species with Polycentropus urubici Holzenthal and Almeida. Ten of the other Polycentropus gertschi group species form a second cluster of diagnostically similar species, the Polycentropus soniae cluster (Polycentropus caaetesp. n., Polycentropus carolaesp. n., Polycentropus cheliceratussp. n., Polycentropus fluminensissp. n., Polycentropus itatiaiasp. n., Polycentropus minerosp. n., Polycentropus santateresaesp. n., Polycentropus soniaesp. n., Polycentropus tripuisp. n., and Polycentropus virginiaesp. n.). Two of the remaining 8 new species are included in the Polycentropus jorgenseni species complex (Polycentropus cipoensissp. n. and Polycentropus verruculussp. n.), while the remaining 6 are unique and cannot be placed in one of the groups at this time (Polycentropus acinaciformissp. n., Polycentropus amphirhamphussp. n., Polycentropus cachoeirasp. n., Polycentropus inusitatussp. n., Polycentropus paprockiisp. n. and Polycentropus rosalysaesp. n.).     

Systematics, conservation and morphology of the spider genus Tayshaneta (Araneae, Leptonetidae) in Central Texas Caves

The spider genus Tayshaneta is revised based on results from a three gene phylogenetic analysis (Ledford et al. 2011) and a comprehensive morphological survey using scanning electron (SEM) and compound light microscopy. The morphology and relationships within Tayshaneta are discussed and five species-groups are supported by phylogenetic analyses: the anopica group, the coeca group, the myopica group, the microps group and the sandersi group. Short branch lengths within Tayshaneta contrast sharply with the remaining North American genera and are viewed as evidence for a relatively recent radiation of species. Variation in troglomorphic morphology is discussed and compared to patterns found in other Texas cave invertebrates. Several species previously known as single cave endemics have wider ranges than expected, suggesting that some caves are not isolated habitats but instead form part of interconnected karst networks. Distribution maps are compared with karst faunal regions (KFR's) in Central Texas and the implications for the conservation and recovery of Tayshaneta species are discussed. Ten new species are described: Tayshaneta archambaultisp. n., Tayshaneta emeraldaesp. n., Tayshaneta fawcettisp. n., Tayshaneta grubbsisp. n., Tayshaneta madlasp. n., Tayshaneta oconnoraesp. n., Tayshaneta sandersisp. n., Tayshaneta sprouseisp. n., Tayshaneta vidriosp. n. and Tayshaneta whiteisp. n. The males for three species, Tayshaneta anopica (Gertsch, 1974), Tayshaneta devia (Gertsch, 1974) and Tayshaneta microps (Gertsch, 1974) are described for the first time. Tayshaneta furtiva (Gertsch, 1974) and Tayshaneta uvaldea (Gertsch, 1974) are declared nomina dubia as the female holotypes are not diagnosable and efforts to locate specimens at the type localities were unsuccessful. All Tayshaneta species are thoroughly illustrated, diagnosed and keyed. Distribution maps are also provided highlighting areas of taxonomic ambiguity in need of additional sampling.     

A taxonomic revision of the Neoserica (sensu lato) pilosula group (Coleoptera,Scarabaeidae, Sericini)

Nine new species of the Neoserica (sensu lato) pilosula Moser, 1915, group are described from China: Neoserica curvipenis sp. n., N. emeishanensis sp. n., N. lincangensis sp. n., N. ludingensis sp. n., N. lushuiana sp. n., N. rangshuiensis sp. n., N. shennongjiaensis sp. n., N. tianeana sp. n., and N. weibaoshanica sp. n. The lectotype of Neoserica pilosula Moser, 1915, is designated. Habitus and male genitalia are illustrated, a key to the species of the group and a map of species distribution are given.     

Morphology and infraciliature of a new marine ciliate Euplotidium smalli n. sp. with description of a new genus,Paraeuplotidium n. g. (Ciliophora,Euplotida)

The morphology and infraciliature of a new hypotrichous ciliate, Euplotidium smalli n. sp., isolated from eutrophic coastal water in Korea, were observed in living cells and investigated using the protargol impregnation technique. This new ciliate bears 13-14 frontoventral cirri, 7 transverse cirri, and 5-6 dorsal kineties. Neither left marginal cirrus nor caudal cirrus is present. The new species differs from the related species, Euplotidium agitatum Noland, 1937 in the different number of frontoventral and transverse cirri and different body shape. With the exception of Euplotidium agitatum, the known species of the genus Euplotidium Noland, 1937 with the presence of left marginal cirrus are assigned to a new genus, Paraeuplotidium n. g. Diagnosis of Paraeuplotidium is: Gastrocirrhidae with funnel-shaped buccal cavity; with frontoventral and transverse cirri; left marginal cirrus present. Paraeuplotidium itoi (Ito, 1958) n. comb. is designated here the type species. Four additional species are included: Paraeuplotidium psammophilus (Vacelet, 1961) n. comb., Paraeuplotidium arenarium (Magagnini & Nobili, 1964) n. comb., Paraeuplotidium helgae (Hartwig, 1980) n. comb., and Paraeuplotidium prosaltans (Tuffrau, 1985) n. comb. An improved generic diagnosis of Euplotidium is suggested based on morphology and infraciliature characters: marine hypotrichs with a funnel-shaped buccal cavity; with frontoventral and transverse cirri; neither left marginal cirrus nor caudal cirri present.     

Two new species of Quedius Stephens, subgenus Raphirus Stephens from Yunnan, Southwest China (Coleoptera, Staphylinidae, Staphylinini)

Two new species of the genus Quedius Stephens collected from Nabanhe Nature Reserve, Yunnan Province, Quedius nabanhensissp. n. and Quedius maoxingisp. n., are described and illustrated. Keys to the multipunctatus group and intricatus group of Quedius species of Yunnan are provided. A map of the collecting sites is given.     

A taxonomic review of the Neoserica (sensu lato) abnormis group (Coleoptera,Scarabaeidae, Sericini)

The present paper revises the species belonging to the Neoserica (sensu lato) abnormis group, so far known only with two nominal species. Twenty new species are herein described from Indochina and southern China: N. abnormoides sp. n. (Vietnam, China), N. allolaotica sp. n., N. namthaensis sp. n., N. simplicissima sp. n. (Laos), N. thailandensis sp. n. (Thailand), N. alloputaoana sp. n., N. kanphantensis sp. n., N. natmatoungensis sp. n., N. putaoana sp. n., N. taunggyiana sp. n. (Myanmar), N. lamellosa sp. n., N. tonkinea sp. n. (Vietnam), N. bairailingshanica sp. n., N. euyunnanica sp. n., N. huangi sp. n., N. jiangxiensis sp. n., N. trifida sp. n., N. yaoi sp. n., N. yingjiangensis sp. n. (China), N. cardamomensis sp. n. (Indochina and southern China). One new combination is established: Neoserica ponderosa Arrow, 1946, comb. n. The lectotypes of Neoserica abnormis Moser, 1908 and the taxonomically uncertain N. inclinata Brenske, 1898, which very likely also belongs to this species group, are designated herein. A key to the species and to species groups is given, the genitalia of all species including their habitus are illustrated. Maps of species distribution are included.     

Review of the Southeast Asian species of the Aenictus javanus and Aenictus philippinensis species groups (Hymenoptera, Formicidae, Aenictinae)

The Southeast Asian species of the Aenictus javanus and Aenictus philippinensis groups are revised. Six species (four named and two new species) of the Aenictus javanus group occurring in this area are: Aenictus doydeei Jaitrong & Yamane, 2011, Aenictus duengkaei Jaitrong & Yamane, sp. n., Aenictus javanus Emery, 1896, Aenictus longinodus Jaitrong & Yamane, sp. n., Aenictus nishimurai Terayama & Kubota, 1993, and Aenictus piercei Wheeler & Chapman, 1930. Four species (three named and one new species) are recognized in the Aenictus philippinensis group: Aenictus pangantihoni Zettel & Sorger, 2010, Aenictus philippinensis Chapman, 1963, Aenictus punctatus Jaitrong & Yamane, sp. n., and Aenictus rabori Chapman, 1963. Aenictus piercei is removed from the members of the Aenictus piercei group sensu Jaitrong and Yamane (2011) and transferred to the Aenictus javanus group. Lectotypes and paralectotypes are designated for Aenictus piercei and Aenictus rabori. Size variation occurs among individuals from single colonies of the Aenictus javanus group, while the workers in the Aenictus philippinensis group are clearly monomorphic.     

A taxonomic review of the Neoserica (sensu lato) septemlamellata group (Coleoptera,Scarabaeidae, Sericini)

In the present paper the species belonging to the Neoserica (sensu lato) septemlamellata group, that included so far only four known species, are revised. Here we describe eleven new species originating mainly from Indochina and Southern China: N. daweishanica sp. n., N. gaoligongshanica sp. n., N. guangpingensis sp. n., N. igori sp. n., N. jiulongensis sp. n., N. plurilamellata sp. n., N. weishanica sp. n., N. yanzigouensis sp. n. (China) N. sapaensis sp. n. (China, Vietnam), N. bansongchana sp. n., N. takakuwai sp. n. (Laos). The lectotypes of Neoserica septemlamellata Brenske, 1898 and N. septemfoliata Moser, 1915 are designated. Keys to the species and species groups are given, the genitalia of all species and their habitus are illustrated and distribution maps are included.     

A revision of the spider genus Selenops Latreille, 1819 (Arachnida, Araneae, Selenopidae) in North America, Central America and the Caribbean

The spider genus Selenops Latreille, 1819 occurs in both the Old World and New World tropics and subtropics and contains nearly half of the species in the family Selenopidae Simon, 1897. In this paper the members of the genus Selenops found in North America, Central America, and on islands of the Caribbean are revised, excluding Cuban endemics. No taxonomic changes are currently made to the species from the southwestern United States. In total, 21 new species are described, including Selenops arikoksp. n., Selenops chamelasp. n., Selenops amonasp. n., Selenops bawekasp. n., Selenops bocacanadensissp. n., Selenops enriquillosp. n, Selenops ixchelsp. n., Selenops huetocatlsp. n., Selenops kalinagosp. n., Selenops oviedosp. n., Selenops morrosp. n., Selenops deniasp. n., Selenops duansp. n., Selenops malinalxochitlsp. n., Selenops oricuajosp. n., Selenops petenajtoysp. n., Selenops guerrerosp. n., Selenops makimakisp. n., Selenops souligasp. n., Selenops wilmotorumsp. n., and Selenops wilsonisp. n. Six species names were synonymized: Selenops lunatus Muma, 1953 syn. n. =Selenops candidus Muma, 1953; Selenops tehuacanus Muma 1953 syn. n., Selenops galapagoensis Banks, 1902 syn. n. and Selenops vagabundus Kraus, 1955 syn. n. = Selenops mexicanus Keyserling, 1880; Selenops santibanezi Valdez-Mondragón, 2010 syn. n. = Selenops nigromaculatus Keyserling, 1880; and Selenops salvadoranus Chamberlin, 1925 syn. n. = Selenops bifurcatus Banks, 1909. Lectotypes are designated for the following three species: Selenops marginalis F. O. Pickard-Cambridge, 1900 (♂), Selenops morosus Banks, 1898 (♂), and Selenops mexicanus Keyserling, 1880 (♀). The female neotype is designated for Selenops aissus Walckenaer, 1837. The males of Selenops bani Alayón-García, 1992 and Selenops marcanoi Alayón-García, 1992 are described for the first time, and the females of Selenops phaselus Muma, 1953 and Selenops geraldinae Corronca, 1996 are described for the first time. Almost all species are redescribed, barring Cuban endemics and a few species recently described. New illustrations are provided, including those of the internal female copulatory organs, many of which are illustrated for the first time. A key to species is also provided as are new distributional records.     

Review of the genus Hartemita Cameron, 1910 (Hymenoptera, Braconidae, Cardiochilinae), with the description of six new species from Vietnam

The Oriental and East Palaearctic genus Hartemita Cameron, 1910 (Braconidae: Cardiochilinae) is recorded for the first time from Vietnam. Sixteen species of the genus Hartemita are currently recognized from Oriental and East Palaearctic regions. One species is newly recorded for Vietnam, Hartemita singaporensis (Mao, 1945)and six new species from Vietnam are described and illustrated: Hartemita coffeanasp. n., Hartemita daklakasp. n., Hartemita khuatbaolinhaesp. n., Hartemita similissp. n., Hartemita maculatasp. n. and Hartemita vietnamicasp. n. A key to species of the genus Hartemita Cameron is included.     

New earthworm species of the genus Amynthas Kinberg, 1867 from Thailand (Clitellata, Oligochaeta, Megascolecidae)

Four new species of terrestrial earthworms from the zebrus-group in the genus Amynthas Kinberg, 1867, are described from Nan province, north Thailand: Amynthas phatubensissp. n., from Tham Pha Tub Arboretum, Amynthas tontongsp. n., from Tontong Waterfall, Amynthas borealissp. n., from Chaloemprakiat district, and Amynthas srinansp. n., from Srinan National Park.After comparing with the two closely related Laos species Amynthas chandyi Hong, 2008 and Amynthas namphouinensis Hong, 2008, the four new species show clear morphological differences, and also it is confirmed that there are no previous records of the species described here. Amynthas phatubensissp. n. is the largest (longest) sized of these earthworms and is the only species that lives in limestone habitats. The genital characters are different among them and also from the two Laotian species. Molecular systematics would be a good method for further analysis of the diversity and species boundaries in SE Asian Amynthas.     

La famille des Cicerinidae (Turbellaria, Kalyptorhynchia)

The present study deals with 12 species of Cicerinidae. Further details are given for 6 species previously described: Toia ycia Marcus, Cicerina eucentrota Ax, Nan-norhynchides vividus Brunet, JV. harparius Brunet, N. corneus Brunet, Didiadema picardi Brunet. Two new species are described, Toia calceformis sp.n., different from T. ycia by half-boot shape of the cirrus, and Paracicerina globulosa sp.n., characterized by very short (4 μm) cuticular spermatic ducts and a spherical copulatory bulb provided with a spiny cirrus (35–40 μm long). Histological studies of Ptyalorhynchus coecus Ax confirm the generic separation of Ptyalorhynchus and Paracicerina; the main difference is the glandular girdle structure of the proboscis in Ptyalorhynchus which is split into two areas (inner and outer one) by basal circular muscles; the intensively stained secretion of the inner area is localized in 15 membranous canals. Pocillorhynchus gen.n., with two species, P. agilis sp.n. and P. minutus sp.n., is closely related to the genera Toia and Nannorhynchides , but differs from them by the following characters: proboscis with broad glandular girdle, devoid of circular muscle bulb, cuticular spermatic ducts long and thin (95–125 μm). From the comparison of their morphological and anatomical features, two groups of genera are distinguished in Cicerinidae; first group including Toia - Nannorhynchides - Pocillorhynchus , second group including Cicerina - Paracicerina - Etmorhynchus - Ptyalorhynchus - Zonorhynchus - Xenocicerina - Didiadema. Special attention is paid to proboscis structure. The study ends with an identification key for all known Cicerinidae.     

Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)

The large genus Orthomorpha is rediagnosed and is shown to currently comprise 51 identifiable species ranging from northern Myanmar and Thailand in the Northwest to Lombok Island, Indonesia in the Southeast. Of them, 20 species have been revised and/or abundantly illustrated, based on a restudy of mostly type material; further 12 species are described as new: Orthomorpha atypicasp. n., Orthomorpha communissp. n., Orthomorpha isarankuraisp. n., Orthomorpha picturatasp. n., Orthomorpha similanensissp. n., Orthomorpha suberectasp. n., Orthomorpha tuberculiferasp. n.,Orthomorpha subtuberculiferasp. n. and Orthomorpha latitergasp. n., all from Thailand, as well as Orthomorpha elevatasp. n.,Orthomorpha spiniformissp. n. and Orthomorpha subelevatasp. n., from northern Malaysia. The type-species Orthomorpha beaumontii (Le Guillou, 1841) is redescribed in due detail from male material as well, actually being a senior subjective synonym of Orthomorpha spinala (Attems, 1932), syn. n. Two additional new synonymies are proposed: Orthomorpha rotundicollis (Attems, 1937) = Orthomorpha tuberculata (Attems, 1937), syn. n., and Orthomorpha butteli Carl, 1922 = Orthomorpha consocius Chamberlin, 1945, syn. n., the valid names to the left. All species have been keyed and all new and some especially widespread species have been mapped. Further six species, including two revised from type material, are still to be considered dubious, mostly because their paraterga appear to be too narrow to represent Orthomorpha species. A new genus, Orthomorphoidesgen. n., diagnosed versus Orthomorpha through only moderately well developed paraterga, coupled with a poorly bi- or trifid gonopod tip, with at least some of its apical prongs being short spines, is erected for two species: Orthomorpha setosus (Attems, 1937), the type-species, which is also revised from type material, and Orthomorpha exaratus (Attems, 1953), both comb. n. ex Orthomorpha.     

Molecular phylogeny of the Trechus brucki group, with description of two new species from the Pyreneo-Cantabrian area (France, Spain) (Coleoptera, Carabidae, Trechinae)

A molecular phylogeny of the species from the Trechus brucki clade (previously Trechus uhagoni group)based on fragments of four mitochondrial genes and one nuclear gene is given. We describe Trechus (Trechus) bouillonisp. n. from the western pre-Pyrenees: Sierras de Urbasa-Andía, Navarra, Spain. The species was collected in mesovoid shallow substratum (mss), a subterranean environment. Molecular as well as morphological evidences demonstrate that the new species belongs to the Trechus brucki clade. A narrow endemic species of high altitude in western French Pyrenees merged with Trechus brucki Fairmaire, 1862a, Trechus bruckoidessp. n., is described. A lectotype is designated for Trechus brucki and Trechus planiusculus Fairmaire, 1862b (junior synonym of Trechus brucki). The species group is redefined based on molecular and morphological characters, and renamed as the brucki group, as Trechus brucki was the first described species of the clade. A unique synapomorphy of the male genitalia, a characteristic secondary sclerotization of the sperm duct, which is shared by all the species of the brucki group sensu novo, is described and illustrated. The Trechus brucki group sensu novo is composed of Trechus beusti (Schaufuss, 1863), Trechus bouillonisp. n., Trechus brucki, Trechus bruckoidessp. n., Trechus grenieri Pandellé, 1867, T. uhagoni uhagoni Crotch, 1869, T. uhagoni ruteri Colas, 1935 and Trechus pieltaini Jeannel, 1920. We discuss the taxonomy of the group and provide illustrations of structures showing the differences between the species, along with distribution data and biogeographical comments.     

New Palaearctic species of the tribe Thalassaphorurini Pomorski, 1998 (Collembola, Onychiuridae)

The paper is devoted to a taxonomic revision of the genus Sensillonychiurus Pomorski & Sveenkova, 2006. Five new species of this genus, i.e. Sensillonychiurus mirussp. n., Sensillonychiurus taimyrensissp. n., Sensillonychiurus vegaesp. n., Sensillonychiurus vitimicussp. n., and Sensillonychiurus amuricussp. n., as well as three new species of the related genus Allonychiurus Yoshii, 1995, i.e. Allonychiurus subvolinensissp. n., Allonychiurus elikoniussp. n., and Allonychiurus unisetosussp. n. are being described from various regions of Eurasia. The diagnoses of both genera are amended to include described species. Two genera, Tantulonychiurus Pomorski, 1996 and Thibaudichiurus Weiner, 1996, are treated as junior synonyms of the genus Allonychiurus. Agraphorura eisi (Rusek, 1976) is transferred to Sensillonychiurus; Tantulonychiurus volinensis (Szeptycki, 1964) and Tantulonychiurus asiaticus Babenko, 2007 to Allonychiurus. A review of morphological peculiarities of Sensillonychiurus is performed, comparisons with the other genera of Thalassaphorurini given, and a key to the known species provided.