Climatic impacts on the vegetation of eastern North America during the past 2000 years

Pollen diagrams from seven lakes with annually laminated sediments sampled at 40-year intervals are analyzed to isolate the climatic effects from other effects on the long-term dynamics of vegetation during the past 1000–2000 years along a transect from Maine to Minnesota. Principal components analysis is used to reduce the dimensionality of the pollen data. The pollen records from all lakes show long-term trends, medium frequency oscillations, and higher frequency fluctuations. The long-term trend is associated with the neoglacial expansion of the boreal forest. The mechanism causing this replacement is a change in frequency of air masses in the area. The medium-frequency oscillations are also associated with climate changes, the most recent of which is the ‘Little Ice Age’. The climate-related mechanism causing the medium-frequency changes may be changes in disturbance frequency. The higher frequency fluctuations may also be related to disturbance. This analysis of pollen diagrams into time scales of variation has enabled the separation of climate from other factors affecting vegetation dynamics. By comparing the principal components across a transect of sites it proved possible to interpret the climatic effects on vegetation at most sites and not only at range boundaries and ‘sensitive’ sites.     

Seed bank and vegetation composition of forest stands of varying age in central Belgium: consequences for regeneration of ancient forest vegetation

Vegetation composition differs significantly between ancientand recent forest, due to slow colonization capacity of typical forest speciesand the higher abundance of early successional species in recent forest.However, little is known about differences in persistent seed bank compositionbetween ancient and recent forest and about the interaction between seed bankand vegetation in relation with forest age. We surveyed the seed bank and theunderstorey vegetationcomposition in transects from ancient to recent forest. Seed bank and fieldlayer vegetation characteristics and similarity between seed bank andvegetationwere analysed in relation to recent forest age and distance to the ancientforest. A total of 39 species and 14,911 seedlings germinated, whichcorresponds with a seed density of 12,426 seeds/m².Total seed density is significantly higher in the youngest recent forest parcel(55 years). Also the seed bank composition in the youngest forest parceldifferssignificantly from the other parcels. After a longer period of reforestation,the seed bank approaches that of the ancient forest, suggesting seed bankdepletion, although the seed bank is permanently replenished to some extent byseed bank forming species from local disturbances. Seed bank composition doesnot change significantly with distance to the ancient forest. Similaritybetween seed bank and vegetation composition, nomatter the forest age, is very low, but decreases with increasing forest age.The most frequent species in the vegetation are absent in the seed bank andvice versa. The contribution of forest species is highin the vegetation and they almost not occur in the seed bank, while species offorest edges and clearings, and species of disturbed environments are morefrequent in the seed bank. The seed bank is mainly composed of earlysuccessional species of former forest stages or species which temporary occurinsmall-scale disturbances. The seed bank may enhance the negative effects ofearlysuccessional, mainly competitive species to the forest species richness in therecent forest. In this respect, forest management should minimise forestdisturbances, to prevent germination of competitive species form buriedseeds.     

The Holdridge life zones of the conterminous United States in relation to ecosystem mapping

Aim Our main goals were to develop a map of the life zones for the conterminous United States, based on the Holdridge Life Zone system, as a tool for ecosystem mapping, and to compare the map of Holdridge life zones with other global vegetation classification and mapping efforts. Location The area of interest is the forty-eight contiguous states of the United States. Methods We wrote a PERL program for determining life zones from climatic data and linked it to the image processing workbench (IPW). The inputs were annual precipitation (Pann), biotemperature (T_bio), sea-level biotemperature (T₀bio), and the frost line. The spatial resolution chosen for this study (2.5 arc-minute for classification, 4-km for mapping) was driven by the availability of current state-of-the-art, accurate and reliable precipitation data. We used the Precipitation-elevation Regressions on Independent Slopes Model, or PRISM, output for the contiguous United States downloaded from the Internet. The accepted standard data for air temperature surfaces were obtained from the Vegetation/Ecosystem Modelling and Analysis Project (VEMAP). This data set along with station data obtained from the National Climatic Data Center for the US, were used to develop all temperature surfaces at the same resolution as the Pann. Results The US contains thirty-eight life zones (34% of the world's life zones and 85% of the temperate ones) including one boreal, twelve cool temperate, twenty warm temperate, four subtropical, and one tropical. Seventy-four percent of the US falls in the ‘basal belt’, 18% is montane, 8% is subalpine, 1% is alpine, and < 0.1% is nival. The US ranges from superarid to superhumid, and the humid province is the largest (45% of the US). The most extensive life zone is the warm temperate moist forest, which covers 23% of the country. We compared the Holdridge life zone map with output from the BIOME model, Bailey's ecoregions, Küchler potential vegetation, and land cover, all aggregated to four cover classes. Despite differences in the goals and methods for all these classification systems, there was a very good to excellent agreement among them for forests but poor for grasslands, shrublands, and nonvegetated lands. Main conclusions We consider the life zone approach to have many strengths for ecosystem mapping because it is based on climatic driving factors of ecosystem processes and recognizes ecophysiological responses of plants; it is hierarchical and allows for the use of other mapping criteria at the association and successional levels of analysis; it can be expanded or contracted without losing functional continuity among levels of ecological complexity; it is a relatively simple system based on few empirical data; and it uses objective mapping criteria.     

Holocene climate and vegetation in the Milford drainage basin,Maine, U.S.A., and their implications for human history

A 9200 ¹⁴C year fossil pollen record from a small kettle lake in central Maine, northeast U.S.A., records the development of nearby upland vegetation throughout the Archaic, Ceramic, and Historic periods of human history. The Early Archaic period (9000 to 8000 B.P.) began as open woodland dominated by Picea, Populus, and Larix, which was replaced by Pinus forest. During the Middle Archaic (8000-6000 B.P.) Tsuga-dominated forest, which developed ca. 7400 B.P., was followed by Pinus forest (ca. 6400 B.P.). The Late Archaic (6000-3000 B.P.) was a period of great transition; Tsuga forest developed again ca. 5700 B.P., but was abruptly replaced by northern hardwood forest ca. 4700 B.P. That Late Archaic expansion of hardwoods would have provided better forage for beaver. Coincidentally, boreal wetland mammals such as beaver (Castor canadensis) and muskrat (Ondatra zibethicus) increase in faunal assemblages of local archaeological sites, while remains of anadromous fish decrease. We postulate that the apparent increase in human populations throughout the region during the Late Archaic may be attributed to an increase in the resource base within both upland and wetland areas resulting from the development of hardwood forest in response to climatic cooling.     

Cytopathology training in the United States – a personal view

In this review we present an outline of cytopathology training in the United States, for a non-US readership.     

Compensatory mechanisms of central U.S. hardwood forest communities in a changing environment

Although it is known that forest productivity is highly correlated with canopy leaf area, the influence of environmental gradients within a forest on leaf area-productivity relationships has not been well-documented, particularly for mixed-species forests. The present study was conducted to define the mechanisms by which Central Hardwood forest communities respond to changes in environment. Ecological Land Type Phases (ELTPs) of the Central Hardwood region, delineated by soils, topography, and indicator vegetation, provided a framework to examine the impacts of environmental gradients on leaf area-productivity relationships at a landscape level. Three ELTPs- (1) Quercus prinus / Carex picta-Vaccinium, Dry slope, (2) Quercus alba-Acer saccharum / Parthenocissus, Dry-Mesic Slope, and (3) Fagus-Acer saccharum / Arisaema, Mesic Slope- were selected and stratified into three slope positions viz. the upper edge of the community, the middle or center of the community on the slope, and the lower edge of the community. Circular plots of 0.02 ha in size were established for sampling. Standing stock (basal area, volume, and biomass), annual production, leaf area index, and growth efficiency were determined for each slope position and examined to determine the mechanisms and trends for community change across slope positions. Among the three communities studied, dry slopes compensated for drier conditions through both a shift in species composition and reduced annual production. Dry-mesic slopes exhibited adaptation only through changing species composition whereas mesic slopes compensated for drier conditions primarily through reduction in annual production. These results indicate that mixed-species forest communities adjust to environmental gradients through several mechanisms and that the relative importance of these mechanisms changes with community type.     

黄土高原森林植被对流域径流的调节作用

对黄土高原腹地子午岭典型森林流域与非森林流域年径流变化的对比分析结果表明,森林流域径流年内分配比非森林流域相对均匀,汛期（6～9月）总径流量减少了8.9mm（葫芦河比蒲河）和7.1mm（合水川比东川）,枯季径流与汛期降水及枯水季节降水的回归分析可见,森林植被能将雨季蓄积的部分降水转化为地下径流,增大枯水季节的径流量,但由于黄土区土层深度,植被蒸腾耗水强烈,森林植被对枯水期河川径流的调节作用十分有限,森林植被对10～12月径流总的补枯效应仅为1.69mm（葫芦河比蒲河）和0.5mm（合水川比东川）,对1～4月径流无调节作用,说明森林植被拦蓄的大部分降雨被植物吸收利用,消耗于蒸腾,从而揭示了森林植被对河川径流的削洪补枯效应及其机理。     

Law of the Sea Issues Between the United States and East Asian States

The United States is the sole superpower in the contemporary world and its role in the development of the law of the sea cannot be ignored. Although having not yet acceded to the U.N. Convention on the Law of the Sea, the United States has contributed to the development of the international law of the sea in numerous ways, including responding to the so-called excessive maritime claims in East Asia and creating new rules of maritime enforcement. This article assesses this recent U.S. practice.     

Spatial patterning in the canopy gap regime of a subalpine Abies-Picea forest in the northeastern United States

Abstract. Spatial patterning in the disturbance regime of a forest affects the vegetation dynamics. Therefore, the distribution of canopy gaps was examined in detail for a subalpine Abies-Picea forest in the northeastern United States. Gaps were not randomly distributed. The fraction of forest area in gaps and the abundance of gaps varied significantly with topographic position, elevation, and slope percent. On average, 15 % of the forest was influenced by gaps, but the gap fraction was greater near ridges (23 %) and near streams (27 %) than on the backslope (13 %). Also, gaps were larger and more abundant near streams and ridges. Gap fraction varied with elevation as well: more of the forest was disturbed at lower and higher elevations than at mid-elevations. Significantly more of the forest on steep slopes (≥ 30 %) was under gaps. As a result of this patterning, some parts of the Abies-Picea forest were predictably more disturbed than others. A remaining question is whether this patterning is sufficient to influence the regeneration environment and thus forest composition.     

The systematic significance of flavonoids in Persea of the Southeastern United States

The systematic significance of flavonoid distribution in the southeastern United States taxa of Persea is discussed. The compounds thus far elucidated are glycosides based on quercetin, kaempferol, apigenin or luteolin.     

History of vegetation and land use from 3200 B.P. to the present in the north-west Burren,a karstic region of western Ireland

A radiocarbon-dated pollen profile provides the first evidence for the vegetation history of the north-west Burren, a karstic region in Co. Clare, western Ireland. The profile, which spans the period ca. 3200-0 B.P., shows that Corylus was the main woody species and that, apart from the later eighteenth and nineteenth centuries, when woody species were almost totally cleared, it constituted a more important element in the landscape than at present. The data indicate that small populations of Quercus, Fraxinus, Betula, Alnus and possibly Ulmus were also present until at least the mid seventeenth century. Exceptionally high values of Plantago lanceolata (mostly in excess of 20% total terrestrial pollen) suggest intensive pastoral farming over most of the period. Between A.D. 200–580, a lull in activity is recorded during which woody species and, in particular, hazel scrub regenerated. The separation of pollen of cereal origin from large cereal-type pollen produced by non-cultivated grasses proved particularly problematic. Reliance has therefore been placed on the weed pollen flora as a guide to arable activity. This flora indicates that arable farming was of some importance in the Burren from A.D. 1000 onwards. The implications of the various findings for the longer term history of the rarer elements of the Burren flora and the relationship between the pollen record and the archaeological and historical evidence for settlement and land use are discussed.     

Nitrogen loss from deserts in the southwestern United States

A lower limit for nitrogen loss from desert ecosystems in the southwestern United States was estimated by comparing nitrogen inputs to the amount of nitrogen stored in desert soils and vegetation. Atmospheric input of nitrogen for the last 10 000 years was conservatively estimated to be 2.99 kg N/m². The amount of nitrogen stored in desert soils was calculated to be 0.604 kg N/m³ using extant data from 212 profiles located in Arizona, California, Nevada, and Utah. The average amount of nitrogen stored in desert vegetation is approximately 0.036 kg N/m².Desert conditions have existed in the southwestern United States throughout the last 10 000 years. Under such conditions, vertical leaching of nitrogen below a depth of 1 m is small (ca. 0.028 kg N/m² over 10 000 years) and streamflow losses of nitrogen from the desert landscape are negligible. Thus, the discrepancy found between nitrogen input and storage represents the amount of nitrogen lost to the atmosphere during the last 10 000 years. Loss of nitrogen to the atmosphere was calculated to be 2.32 kg N/m², which is 77% of the atmospheric inputs.Processes resulting in nitrogen loss to the atmosphere from desert ecosystems include wind erosion, ammonia volatilization, nitrification, and denitrification. Our analysis cannot assess the relative importance of these processes, but each is worthy of future research efforts.     

Height and death in the Antebellum United States: A view through the lens of geographically weighted regression

Factors related to geography such as climate, natural resources or waterways often affect human activities. However, traditional approaches such as ordinary least squares (OLS) have limitations in investigating such patterns. Unlike OLS regression, geographically weighted regression (GWR) allows the coefficients of explanatory variables to differ by locality by giving relatively more weight to geographically close observations. GWR depicts spatial patterns. This paper examines the pattern of height and crude death rate in the United States prior to the Civil War by this method. The GWR results show that access to water transportation increased mortality and decreased stature in the food exporting areas of the Midwest, and the opposite pattern appeared in the food importing areas of the Northeast.     

Relationships between the vegetation and soil conditions in beech and beech-oak forests of northern Germany

In this study we examine the relationships between the vegetation of beech and beech-oak forest communities (Hordelymo-Fagetum, Galio-Fagetum, Deschampsio-Fagetum, Betulo-Quercetum) and their soil conditions in the lowlands of northern Germany, based on 84 sample plots. In all plots the vegetation was recorded and soil parameters were analysed (thickness of the O- and the A-horizons, pH, S-value, base saturation, C/N, mean Ellenberg moisture indicator value). The vegetation classification according to the traditional Braun-Blanquet approach was compared with the result of a multivariate cluster analysis. Vegetation-site relationships were analysed by means of an indirect gradient analysis (DCA).Both traditional classification methods and the cluster analysis have produced comparable classification results. So far as the species composition is concerned, a similar grouping of sample plots was found in both approaches. Multivariate cluster analysis thus supports the classification found by the Braun-Blanquet method. The result of the DCA shows that the four forest communities mentioned above represent clearly definable ecological units. The main site factor influencing changes in the species composition is a base gradient, which is best expressed by the S-value. In addition, within the series Hordelymo-Fagetum - Galio-Fagetum - Deschampsio-Fagetum the C/N-ratios and the thickness of the organic layers (O-horizon) increase continuously. By contrast, the floristic differences between oligotrophic forest communities (i.e., Deschampsio-Fagetum and Betulo-Quercetum) cannot be explained by a base gradient and increasing C/N-ratios. It is suggested that a different forest management history in some cases (e.g., promotion of Quercus robur by silvicultural treatments) is responsible for differences in the species composition, but on the other hand the result of the DCA indicates that Fagus sylvatica is replaced by Quercus robur with increasing soil moisture (i.e., with the increasing influence of a high groundwater table). Summarizing these results, it can be concluded that the ecological importance of single site factors affecting the species composition changes within the entire site spectrum covered by the beech and beech-oak forests of northern Germany.     

International Law and Politics in U.S. Policymaking: The United States and the Svalbard Dispute

This article explains the position(s) of the United States in the maritime dispute adjacent to Svalbard. While the United States has regarded Norway's exclusive claim to the natural resources outside Svalbard as everything from “wishful thinking” to legally plausible, Washington maintains that it may have rights under the 1920 Svalbard Treaty to exploit the maritime zones adjacent to the archipelago. The U.S. reservation is the result of assessments and reassessments of legal considerations as well as national interests.     

美国生命科学两用性研究监管政策分析

目的:近年来高致病性禽流感H5N1病毒基因改造研究及美国国家生物安全科学咨询委员会的监管建议引起广泛争议,美国发布了生命科学两用性研究监管政策。本研究旨在通过对美国生命科学两用性研究监管政策进行分析,探讨两用性研究发展和监管趋势。方法:通过文献检索,获取生命科学两用性研究监管政策相关的情报源,对美国政府生命科学两用性研究监管的政策背景、主要内容、政策评论、政策动因进行系统梳理与分析。结果:美国政府出台的生命科学两用性研究监管政策,有助于美国对整个生命科学研究周期(从最初提交资助申请到研究结论和研究成果交流)潜在的误用风险进行管理。结论:美国生命科学两用性研究引发极大争议,两用性研究监管主要依靠生命科学团体的自我管理,政策具有指导性而非强制性。     

A survey of dermatophytes from human patients in the United States from 1985 to 1987

A survey of dermatophytes isolated from patients seeking medical advice was made from 1985 to 1987 in the United States. The survey included 54 locations with data from 45 cities and one state. Listing of the isolated dermatophytes and the frequency given by percentage of the total follows: Trichophyton rubrum 54.8%, T. tonsurans 31.3%, T. mentagrophytes 6.0%, Microsporum canis 4.0%, Epidermophyton floccosum 2.0%, M. gypseum 0.6%, and T. verrucosum 0.2%. Out of a total of 14,696 isolates M. audouinii was cultured 13 times, T. violaceum 12 times, M. nanum 6 times, T. terrestre 4 times, and T. soudanense twice. Single isolations were made of M.fulvum, M. ferrugineum and T. schoenleinii.Collection of dermatophyte data in Tucson, Arizona, began in 1966. In 1987, the first case of tinea capitis caused by T. tonsurans was observed. Other isolates of this organism as the cause of tinea capitis were made in this city during that year. These infections were in black children. With the recent growth of Tucson, the percentage of blacks in the population increased and this pathogen was introduced into the general population.     

Recommendations for the Implementation of a National Sediment Quality Policy in the United States

This article was prepared prior to the author's employment by the U.S. Environmental Protection Agency and therefore does not necessarily represent agency policy. Sediments provide needed habitats to valued aquatic and marine resources and are therefore an integral and valuable component of any coastal ecosystem. Studies have found that multiple anthropogenic activities have been the primary cause of sediment contamination. There is continued evidence that contaminated sediments pose adverse impacts to biological resources even when overlying water quality is acceptable. Efforts in environmental protection should therefore incorporate sediment quality management strategies. Despite a growing awareness and interest in management of contaminated sediment, no established systematic and standardized national management program has been established in the U.S. The establishment of a national sediment policy in the U.S. would reinforce the importance of integrating sediment issues with different management objectives and reiterate the importance of protecting sediments as a natural resource. A national sediment policy could also guide and direct states in developing state sediment policies that reflect national objectives while integrating local concerns. This perspective identifies and defines issues and concerns raised by 13 states’ agency staff in managing contaminated sediments in the absence of a national policy. Recommendations are developed that address sociopolitical and scientific issues and concerns. The recommendations are integrated with stakeholder comments, state manager needs and the existing regulatory and institutional framework.