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1.
桂南爬树蕨的组织结构研究   总被引:3,自引:1,他引:2  
黄玉源   《广西植物》1998,18(4):337-342
本文对桂南爬树蕨的主要器官进行了解剖结构的研究,结果表明:茎的中柱为仅有两个新月型的维管束,中柱内及其周围的机械组织形成“8”字形。与爬树蕨相同,为新的中柱类型,中柱周围的几层皮层组织特化为机械组织的起始位置为对正维管束长轴的中央。而髓部的机械组织分化的起点是在中轴的中间位置,呈线形,同时向两侧分化。木质部分化为沿圆周线方向两端式,为新的分化方式。叶为等面叶,与爬树蕨相比,叶的厚度小,叶肉细胞层数少且细胞排列紧密,机械组织明显少。叶缘近圆形。根为二原型,直径比后者略小些。孢子类型与爬树蕨相同,孢子囊群无囊群盖。  相似文献   

2.
贺兰山10种不同植物的旱生结构研究   总被引:3,自引:0,他引:3  
对贺兰山10种不同生活型植物的形态解剖研究表明,长期生长在干旱贫瘠环境中植物,均形成适应生境的一些特殊形态结构.叶器官主要表现在:(1)叶表面积与体积之比缩小,表面覆有厚的角质膜,并被有表皮毛,气孔下陷,具孔下室;(2)栅栏薄壁组织细胞明显增多,海绵薄壁组织细胞减少.有叶肉不分化叶、等而叶和异而叶3种结构形式;(3)叶内贮水组织和机械组织增强。轴器官主要表现在:(1)根普遍形成周皮,且木栓层细胞层数增多;(2)机械组织非常发达,在周皮、皮层和韧皮部中有许多厚壁细胞分布。维管组织中的木薄壁细胞的细胞壁也明显木质化加厚;(3)有些植物的根具有异常维管组织。另外,这些植物根、茎、叶中均有粘液细胞和含晶细胞分布。这些结构具有重要的适应干旱生境的意义。  相似文献   

3.
10种卷柏属植物茎的解剖学研究及其分类鉴定意义   总被引:4,自引:0,他引:4  
对湖北省及邻近地区产10种卷柏属(Selaginella)植物茎进行了比较解剖研究.结果表明它们在解剖学方面既存在共性特征,又各有其自身的特征.其共性特征表现在:(1)表皮细胞壁多少增厚.(2)皮层通常具机械组织(纤维、厚壁细胞,通常仅有厚壁细胞).(3)内皮层以内有大型气室,中柱由多数横桥悬于气室中.(4)中柱鞘由1至数层薄壁细胞环列而成.(5)维管束周韧型.但10种卷柏属植物茎的横切面外侧是否具脊状突起,皮层是否具纤维,皮层外侧是否具厚壁细胞以及厚壁细胞的层数,构成横桥的薄壁细胞的形态、数目,气室及中柱的形态、数目,木质部形态等方面具有种间差异,可作为分类鉴定的重要依据.同时,利用茎的解剖学特征,可辅助解决某些仅靠形态学性状难以解决的分类鉴定问题.根据其相似程度,还可进行部分种间亲缘关系的分析.  相似文献   

4.
运用形态解剖学和薄层色谱鉴定的方法,对植物药珠芽景天(Sedum bulbiferumMakino)进行了鉴定研究。结果显示该植物药鉴别特征明显,如茎横切面的皮层外侧及中柱薄壁组织散布有紫红色细胞,髓部细胞较小,壁不增厚;叶下表皮气孔密布,常可见2个(有时3个)相距较近;薄层色谱中具多个特征性荧光斑点。文中描述了该植物药材的性状、显微与薄层色谱特征,还与同属近缘植物药垂盆草、佛甲草、凹叶景天进行了薄层色谱比较,并结合四者茎的显微结构,探讨了相互之间的亲缘关系。研究结果为珠芽景天的应用提供了可靠的鉴定依据。  相似文献   

5.
初敬华  何显 《植物研究》1999,19(4):397-400
作者在对单瘤酸模(Rumex crispus var.unicallosus)幼苗维管组织做连续切片的解剖观察时看到,该植物维管组织从根到茎的变化过程中出现4个不同的中柱类型即(1)根-下胚轴的三元型星状中柱;(2)子叶节区下部的单体型原生中柱;(3)子叶节区上部的管状中柱和(4)上胚轴-苗端的真中柱。作者对此进行了讨论,认为子叶节区既然具有原始的单一管状类型的中柱,说明这一部位应为与古老 性顺官  相似文献   

6.
利用冰冻切片法在光学显微镜下观察暹罗苏铁茎的解剖特征。结果表明,暹罗苏铁茎由周皮、皮层、皮层维管束、中柱和髓组成,皮层和髓发达。皮层维管束可分为周韧型和外韧型两种,以周韧型为主。中柱的次生结构为同心环的次生维管组织构成,随着茎的不断成熟,环数逐渐增加;每个同心环均含有次生木质部、维管形成层、次生韧皮部的结构;环与环之间有次生的薄壁组织相间隔。皮层、中柱维管束均由木质部、形成层和韧皮部组成,木质部面积均大于韧皮部。苏铁植物茎在次生结构方面的主要特点是有皮层维管束和同心环结构的中柱维管束。  相似文献   

7.
滇黄芩的解剖学与组织化学研究及其与黄芩的比较   总被引:1,自引:0,他引:1  
应用植物解剖学方法和荧光显微技术研究了滇黄芩(Scutellariaalnoena)营养器官的解剖结构。结果表明:根的初生结构中木质部为二原型,凯氏带明显,并开始积累淀粉粒;在茎和叶维管束外有厚壁细胞群分布,较成熟的茎中厚壁细胞群连成一环。通过组织化学方法对黄酮类物质的定位研究表明,多年生根的中柱鞘及韧皮部、茎及叶的表皮和与皮层中的薄壁细胞是黄酮类物质的主要积累场所,其含量根〉茎〉叶。滇黄芩与同属植物黄芩(S.baicalensis)在结构和组化方面的差异主要表现为:黄芩茎中厚壁细胞群分布区域较小且未连成环,细胞壁加厚较不明显,叶中没有厚壁细胞;组化显色结果表明滇黄芩黄酮类物质含量高于黄芩。因此,深入对地方药用资源滇黄芩的研究与开发十分必要。  相似文献   

8.
福录考体细胞胚胎发生的观察   总被引:1,自引:0,他引:1  
福录考(PhloxdrummondiiHook)体细胞胚胎发生的方式有两种:一是由叶外植体具有一定功能的特化细胞脱化为胚性细胞,如叶表皮细胞,维管束鞘及韬皮薄壁细胞均可发生脱分化进行分裂,形成胚性细胞或分细胞团;二是叶外植体先脱分化形成愈伤组织。  相似文献   

9.
菰(Zizania latifolia)是一种多年生挺水植物,为了探讨该植物根、茎和叶的解剖结构、组织化学及其质外体屏障的通透性生理。该文利用光学显微镜和荧光显微镜,对菰的根、茎、叶进行了解剖学和组织化学研究。结果表明:(1)菰不定根解剖结构由外而内分别为表皮、外皮层、单层细胞的厚壁机械组织层、皮层、内皮层和维管柱;茎结构由外而内分别为角质层、表皮、周缘厚壁机械组织层、皮层、具维管束的厚壁组织层和髓腔。叶鞘具有表皮和具维管束皮层,叶片具有表皮,叶肉和维管束。(2)不定根具有位于内侧的内皮层及其邻近栓质化细胞和外侧的外皮层组成的屏障结构;茎具内侧厚壁机械组织层,外侧的角质层和周缘厚壁机械组织层组成的屏障结构,屏障结构的细胞壁具凯氏带、木栓质和木质素沉积的组织化学特点,叶表面具有角质层。(3)菰通气组织包括根中通气组织,茎、叶皮层的通气组织和髓腔。(4)菰的屏障结构和解剖结构是其适应湿地环境的重要特征,但其茎周缘厚壁层和厚壁组织层较薄。由此推测,菰适应湿地环境,但在旱生环境中分布有一定的局限性。  相似文献   

10.
红豆杉营养器官的解剖学研究   总被引:4,自引:0,他引:4  
通过对红豆杉营养器官的解剖学研究,阐明其根、茎、叶的结构特征,为红豆杉的开发利用研究提供了解剖学依据。研究结果表明红豆杉营养器官具有以下特征:(1)根、茎、叶中均无树脂道;(2)根、茎皮层中均有含单宁类物质的细胞分布;(3)根、茎中均有石细胞分布,(4)叶气孔器为双环型,略内陷;(5)根为二原型,(6)幼根的内皮层具凯氏带加厚,中柱鞘细胞富含单宁类物质;(7)根的次生维管组织中射线发达。  相似文献   

11.
A small permineralized osmundaceous stem has been collected from marine sediments of the Early Cretaceous (Aptian), Upper Chickabally Member of the Budden Canyon Formation near Ono, California. The specimen, 8.5 cm long and 5.4 cm wide, represents a stem surrounded by a mantle of stipular leaf bases and adventitious roots. A large number of sections were studied through the use of the cellulose acetate peel technique. The stem was erect, 11x13 mm in diameter, with a parenchymatous pith and two-layered cortex. The stele is an ectophloic siphonostele with 65-79 leaf traces in the stem per cross section. Leaf gaps are only produced in 13% of the departing traces. Most leaf traces have "delayed" gaps or completely lack leaf gaps. Leaf traces are C-shaped, endarch, with one protoxylem strand, and have sclerenchyma lining the adaxial concavity. Leaf bases have stipular wings with large patches of heterogeneous sclerenchyma and a few scattered strands outside of the heterogeneous sclerotic ring. Patches of sclerenchyma occur inside the ring and outside of the vascular tissues. Numerous diarch roots arise singly or doubly from the leaf traces as they depart the axis stele. Although the stem compares fairly closely to both Ashicaulis Tidwell and Millerocaulis Erasmus ex Tidwell emend. Tidwell, it is most similar to Millerocaulis. However, the combination of characters observed in our specimen differs from that of the seven known species of Millerocaulis. This stem is described as Millerocaulis embreei sp. nov. and is the youngest known species of the genus and the first to be found in the Northern Hemisphere.  相似文献   

12.
A new species of the Osmundaceae, Plenasium xiei sp. nov., is herein described from the Cretaceous of Northeast China. The specimens examined here represent the earliest unequivocal record of the extant genus Plenasium in Eurasia based on fossil rhizomes. The rhizome consists of a central stem with a mantle of petiole bases and adventitious roots. The stem contains an ectophloic‐dictyoxylic siphonostele and a two‐layered cortex. The C‐shaped leaf trace bears two protoxylem bundles at the point of separation from the stele. The pith is heterogeneous. The parenchymatous inner cortex is thinner than the sclerenchymatous outer cortex. Lobed sclerenchyma bands occur at the adaxial sides of the stem xylem strands, in the concavity of the leaf trace, and along the adaxial side of the vascular bundles of the petiole base. In distal petiole portions, the sclerenchyma band splits into several groups in the transverse view. Sclerenchyma rings are heterogeneous with an abaxial sclerenchymatous arc of thick‐walled fibers. Numerous sclerenchyma strands of thick‐walled fibers appear in the petiolar inner cortex and the stipular wing. These fossils provide unambiguous evidence for the existence of subgenus Plenasium of modern Plenasium by at least the Late Cretaceous, demonstrating the longevity of this extant subgenus. Altogether the leaf and rhizome fossil records of Plenasium indicate that this genus was widely distributed across North America and Eurasia from the Early Cretaceous to the Early Cenozoic, followed by a range restriction to Eurasia in the Late Cenozoic. Extant Plenasium species are only known from East and Southeast Asia.  相似文献   

13.
The genus Palaeosmunda was established by R. E. Gould in 1970 based upon some Late Permian Osmundaceous trunks with well-developed leaf gaps and rhomboidal sclerotic ring within petiolar base seen in cross section. As he thinks that the latter character is more important than the former, this genus could not be assigned to any subfamily of Osmundaceae. However, the leaf gap is one of the most important characters in the structure of the fern stem, so the author suggests that this genus should be assigned to subfamily Osmundoideae and its diagnosis must be emended as follows: The genus Palaeosmunda is represented by some rhizomes (or trunks), roots and leaf bases of ferns which structurally are preserved, resembling Osmundacaulis but which can’t be assigned to any group of this genus. Stem containing an ectophloic dictyoxylic siphonostele; if tracheids present in the pith, they being multiseriate scalariform pitted; pith or cortex sometimes contain ing groups of secretory cells or sclerenchyma; number of leaf traces seen in a tran sverse section of cortex more than 30; leaf traces adaxially curvature, rarely oblong shaped; petiolar bases with or without stipular expansion, containing a C-shaped vascular strand; root diarch. Type species——Palaeosmunda williamsii. According to this diagnosis some primitive osmundaceous species with the leaf gaps, which have already found in Upper Permian and Lower Triassic, could be assigned to this genus. Two of them are P. williamsii Gould and P. playfordii Gould, and Osmundacaulis beardmorensis, which was from Lower Triassic of Antarctica in 1978, should be assigned to the genus Palaeosmunda. In this paper two osmundaceous new species: P. primitiva and P. plenasioides were found in the coal balls of Upper Permian age from Wangjiazhai of Shuicheng of Guizhou Province, China. P. primitiva is represented by two trunks; stem about 4 cm in diameter; stele actophloic dictyoxylic siphonostele; pith cavity about 3—4 mm in diameter, contianing parenchyma and tracheids; xylem cylinder thin, less than 10 tracheids in radial thickness, dissected by leaf gaps. Inner cortex about 1.5 cm thick, mainly parenchymatous, but sometimes containing a few sclerenchymatous; number of leaf traces seen in a transverse section about 50—60; leaf traces departing at 35—45º,open C-shaped at point of departure, gradually becoming shallow C-shaped or V-shaped in different parts; protoxylem in base of leaf traces single, endarch; when leaf traces pass through inner cortex, protoxylem biturcating. Petiole bases without stipular expansion, probablyloosely embracing the stem; xylem strand of potiole trace shallow C-shaped, surrounded by selerenchyma; sclerotic ring round, connected with single sclerenchyma mass in the concavity of the petiole trace. Root arising singly from leaf trace, diarch, with inner and outer cortex. P. plenasioides is represented by a rhizome; stem more than 4 cm in diameter; stele actophloic dictyoxylic siphonostele; xylem cylider with about 20 tracheids in radial thickness, dissected by leaf gaps; xylem bundle U-, O-, or crosier- (i.e. query-) shaped; pith and inner cortex parenchymatous, with many groups of secretory cells; leaf trace C-shaped, its base containing two endarch protoxylem groups; root diareh,with inner and outer cortex, arising singly from leaf trace or its base.  相似文献   

14.
木姜子油细胞的发育解剖学研究   总被引:2,自引:1,他引:1  
利用薄切片法对木姜子茎叶油细胞的发育以及油细胞分布的研究结果表明:油细胞最早发生于第一叶原基以及茎端皮层和髓的基本分生组织中,在未出现油细胞以痛,上述器官的基本分生组织和原分生组织,难以区分油细胞的原始细胞与周围细胞。当油细胞原始细胞呈现出体积较大,液泡化程度较低,细胞核大而明显的特征才明显可辩,以后经过液泡融合,油细胞成熟和油细胞细胞质解体阶段而成为一贮油的囊,油细胞中未出现杯形构造。叶和茎中,  相似文献   

15.
A developmental study of the normal shoot apex of Matteucciastruthiopteris suggested that patterned stelar differentiationis initiated immediately beneath the single layer of promeristemand occurs prior to the initiation of the youngest leaf primordium.A developmental study in which all leaf primordia were suppressed,with or without lateral isolation of the terminal meristem byvertical incisions, has confirmed this interpretation of stelardifferentiation. Experimentally-induced changes in the tissueimmediately below the promeristem were reflected in the resultingmature structure of the stele. Failure of leaf gap initialsto differentiate, if all leaf primordia were suppressed at theincipient stage, resulted in a mature stele without leaf gaps.Similarly the disappearance of pith mother cells after severalweeks of leaf removal was associated with the formation of astele without pith. Leaf influence was further assessed by allowingone primordium to develop while all others were suppressed.The developing leaf had a small promoting effect on caulinevascular tissue differentiation but its major impact on theexpansion of the parenchymatous tissues of the stele. Characteristicprotoxylem and protophloem failed to differentiate when allleaves were suppressed and, when leaf was allowed to develop,formed only in relation to the leaf.Copyright 1995, 1999 AcademicPress Leaf influence, vascular pattern formation, experimental surgery, shoot apex development, protoxylem, protophloem, Matteuccia struthiopteris  相似文献   

16.
Basipetal to the shoot apex, a procambial ring with parenchymatous gaps is present. The protoxylem poles are endarrh in both the ectophloic siphonostele and the collateral vascular bundle which comprises the leaf trace. Each leaf trace has an anastomosing system of protoxylem poles that decreases in number basipetally from five to three to two. Differentiation of the leaf trace procambium and protoxylem is bidirectional, that is the differentiation first occurs near the base of the leaf and acropetally in the leaf and basipetally in the stem. Then a fascicular cambium differentiates betweem the primary xylem and phloem in the leaf. This vascular cambium which is also present in the stem is unidirectional and only produces secondary xylem centripetally. Limited secondary growth also occurs in roots. Medullary tracheids when present are longitudinally continuous with the vascular system. The stele of the stem is interpretated as a sympodium of leaf traces and the pith is considered to be fundamental tissue enclosed by the anastomosing of leaf traces.  相似文献   

17.
This study was aimed to evaluate anatomical responses to waterlogging of mangrove seedlings (Avicennia marina (Forsk.) Vierh.) grown in experimentally simulated semidiurnal tides. The following treatments were used: 0, 2, 4, 6, 8, 10 and 12 h submergence period with two daily tidal cycles. With increasing waterlogging duration, the leaf thickness, mesophyll thickness, palisade parenchyma thickness, palisade–spongy ratio and hypodermis thickness decreased, but the mesophyll to leaf thickness ratio, stem and pith diameter, and cortex thickness increased. The tangential vessel diameter, vessel wall thickness in stem and leaf and fiber wall thickness in stem showed a similar tendency in response to waterlogging, remaining constant between 0 and 4 h waterlogging duration, but decreasing with more prolonged waterlogging. When the waterlogging duration exceeded 4 h, no sclerenchyma cells in leaves or gelatinous fibers in stems were observed. The response of these leaf and stem features indicated that water transport and mechanical support could remain relatively stable in the 0–4 h waterlogging duration, but they would be negatively influenced by longer flooding. Tissues for gas exchange were stimulated by waterlogging, while the functions of water storage, photosynthesis, mesophyll conductance were weakened with increasing waterlogging.  相似文献   

18.
MIDDLETON, D. J., A systematic survey of leaf and stem anatomical characters in the genus Gaultheria and related genera (Ericaceae). Leaf and stem anatomical characters of 123 species within the Gaultheria group of genera of the tribe Andromedeae are surveyed. A number of characters show considerable variation within a species which limits their taxonomic use. Other characters such as marginal sclerenchyma and, to a lesser extent, the hypodermis, the presence of adaxial stomata, the presence of free fibres and the pith type were of more taxonomic interest. These characters are discussed in relation to the generic and infrageneric classification of the group.  相似文献   

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