Reef slope geometries and facies distribution: controlling factors (Messinian,SE Spain)

Sea-level fluctuations and changes in sediment grain size are widely thought to be the main factors controlling carbonate platform slope geometries. Two successive clinoform bodies from the Upper Miocene Cariatiz carbonate platform (SE Spain) were selected to analyze geometry and facies distribution in relation to sea-level oscillations. Facies occurring in these clinoform bodies are from top to bottom reef-framework, reef-framework debris, Halimeda breccia, Halimeda rudstone, and bioclastic packstone, as well as siltstone and marl. Slope geometry and facies, composition, and distribution, are significantly different in each clinoform body. These differences are the result of the interaction of several factors such as coral growth, in situ slope carbonate production, rockfalls and sediment gravity flows, hemipelagic rain, reworking of reef-slope facies and siliciclastic input. Changes in accommodation were related to sea-level fluctuations and controlled the relative impact of these factors. A sea-level fall took place in the time between deposition of the selected clinoform bodies and changed the hydrographical conditions of the basin. These changes influenced the presence of Halimeda and the grain-size distribution, and consequently the slope geometries. Reef-slope geometry is not exclusively controlled by changes in grain size. The stabilization by organic binding is proposed to be a significant factor controlling the slope deposition.     

Composite Trace Fossil Assemblage in a Distal Carbonate Setting from the Tethys (Middle Jurassic,Betic Cordillera,Southern Spain)

Ichnological analysis of a Middle Jurassic carbonate surface from the Betic Cordillera (southern Spain) reveals a complex trace fossil assemblage, including softground Ophiomorpha, firmground Arenicolites, Thalassinoides and Gastrochaenolites, and hardground Trypanites as well as possible Gastrochaenolites. The ecological replacement in the macrobenthic community is interpreted according to successive suites that are controlled mainly by substrate consistency. Variations in composition and abundance of trace fossils between suites can be ecologically and/or taphonomically determined.     

Stratigraphic framework, discontinuity surfaces, and regional significance of Campanian slope to ramp carbonates from central Dalmatia, Croatia

The sedimentology, microfacies, and stratigraphic age (from planktonic and benthic foraminifera and strontium-isotope stratigraphy) of a 300-m-thick Upper Cretaceous carbonate succession from the Island of ?iovo (central Dalmatia, Croatia) were analyzed in order to determine the lithostratigraphic, depositional, and chronostratigraphic framework. The Cretaceous strata were deposited in the southern part of the long-lasting (Late Triassic to Paleogene) Adriatic-Dinaridic Carbonate Platform (ADCP), one of a few late Mesozoic, intra-Tethyan, peri-Adriatic (sub)tropical archipelagos. The succession is separated by a firmground formational boundary into two lithostratigraphic units: the underlying Middle to Upper Campanian Dol Formation consisting of slope pelagic limestone with intercalated turbidites and debrites, and the overlying Upper Campanian ?iovo Formation composed of outer-ramp bioclastic-lithoclastic and echinoderm-dominated packstone. Age, lithology, and depositional settings of the ?iovo Formation are different from other penecontemporaneous, regionally important inner-platform carbonate successions within the ADCP domain. Therefore, the ?iovo Formation is proposed here as a new lithostratigraphic unit. Regionally important condensed intervals in the form of at least two firmground surfaces, characterized by Thalassinoides burrows (with phosphatic mineralization) that belong to the Glossifungites ichnofacies, occur in the lowermost part of the ?iovo Formation. Abrupt shallowing of depositional environments at the boundary between the Dol and the ?iovo Formation, and the generation of the formational boundary firmground, likely correlate with the regionally recorded Upper Campanian Event that represents a global eustatic sea-level fall. A regionally important subaerial exposure surface with nodular calcrete, rhizoliths, and Microcodium aggregates in the upper part of the ?iovo Formation represents a regional subaerial unconformity that was recorded across the ADCP domain and was interpreted as a consequence of diachronous and differential uplift of various parts of the platform in response to the formation of a forebulge in front of the approaching Dinaridic orogen.     

Contribution of calcareous nannoplankton to carbonate deposition: a new approach applied to the Lower Jurassic of central Italy

The Upper Pliensbachian–Lower Toarcian Somma section (Central Italy), which displays alternation of limestone–marl lithotypes, was investigated for its calcareous nannofossil and CaCO₃ contents. In order to evaluate the contribution of calcareous phytoplankton to the carbonate fraction of rocks a new method was applied combining a recently developed technique for nannofossil absolute quantification and measurements of nannofossil size. Our study suggests that nannofossils did not produce the bulk of carbonate mud in the studied section, although they contribute to different extents to the pelagic carbonate fraction. The contribution of both coccoliths and (mainly) the Incertae sedis Schizosphaerella to the carbonate fraction was important only in periods of reduced accumulation rates and/or during clay deposition. The highest estimated carbonate production by nannofossils in the studied section is around 83% of the total carbonate content, but in a sample containing only 12.8 wt% CaCO₃. The observed limestone–marl alternations therefore result from cyclical export of carbonate mud from the shallow Latium–Abruzzi carbonate platform to the Umbria–Marche Basin and, only subordinately, from changes in nannoplankton productivity.     

Crayfish burrows from Late Jurassic–Late Cretaceous continental deposits of Patagonia: Argentina. Their palaeoecological,palaeoclimatic and palaeobiogeographical significance

The trace fossil, Loloichnus baqueroensis igen. and isp. nov., from Late Jurassic–Late Cretaceous continental deposits of Patagonia, Argentina, includes thickly lined, mostly passively-filled, and Y-branched burrows. Other important features of this ichnofossil are the inner surface texture of lining showing transversal, elongated, and adjacent grooves, and less commonly, the pelletal filling of burrows. L. baqueroensis is recorded from the Bajo Grande, Bajo Tigre, Punta del Barco, and Laguna Palacios Formations, which were deposited in different volcaniclastic environments of the Deseado Massif and San Jorge Basin geological provinces, respectively. The described burrows are found in many levels of palaeosols developed in reworked piroclastic deposits, where they are the main component of the ichnofabrics, in association with meniscate and thinly lined burrows and a diffuse and complex boxwork of small diameter burrows. Root traces are also present, and in many cases, occur inside L. baqueroensis. Considering general morphology, surface texture, filling types, palaeoenvironments in which they occur, and comparisons with extant and fossil decapod burrows, the likely trace makers of L. baqueroensis were crayfishes (Decapoda: Astacidea), probably Parastacidae. The producers of L. baqueroensis inhabited soils, where their burrows probably reached the water table, and contained roots that were used for feeding. Considering climatic preferences of extant parastacids, it is proposed a temperate climate for central Patagonia during the deposition of the studied units. The widespread presence of crayfishes during Late Jurassic–Late Cretaceous times in central Patagonia, supports the monophyletic origin of this group during the Triassic, and suggests that the present restricted geographic distribution in southern South America is a relict of a broader one.     

Ethological and environmental significance of Bergaueria hemispherica from the Late Cretaceous of the Bagh Group,Western India

The probable trace maker of B. hemispherica is the fact that one encounters different age groups of sea anemone burrows extended at variable depths in soft, unconsolidated, non-fluidized mud to get stability. It occurs as densely packed, vertical to subvertical, usually unornamented, occasionally showing faint, thin, ring-like structures, allochemic sandstone-filled cylinders with hemispherical base extending to variable depths in the shale. The diameter-to-height ratio calculated for 44 burrow specimens shows that the diameters of most of the specimen are smaller than their height. The probable trace maker of B. hemispherica is the fact that one encounters different age groups of sea anemones, which extended their column at variable depth in soft, unconsolidated, non-fluidized mud to get stability. The inclined nature of paired burrows towards each other suggests social aggression while unpaired inclined burrows suggest swaying in search of food. The monodominant occurrence of B. hemispherica as pre-storm colonization of r-selected organisms (sea anemones) suggest sstressed environment and simultaneous vacation of the burrower reflects rapid sedimentation due to high-energy storm events. Further, storm and inter-storm events deposited sandy allochemic limestone and shale series, respectively, but did not form an identical condition for the colonization of the sea anemone. The species B. hemispherica of the Bagh Group revealed physicochemical parameters (energy conditions, turbidity, sedimentation rate, bathymetry, suspended organic matter, substrate consistency, oxygen, and salinity) of the Late Cretaceous transgressive sea.     

An integrated analysis (microfacies and ichnology) of a shallow carbonate-platform succession: upper Aptian,Lower Cretaceous,Betic Cordillera

Four lithofacies and 12 microfacies types recognized in an upper Aptian section in the Sierra de Bedmar-Jódar (Prebetic of Jaén) represent shallow lagoonal environments (marl and marly limestone) and sand bars that delimited the lagoon. The lagoonal facies reflect subtidal restricted water circulation with low energy. The sand bar facies (intertidal environment) have upper surfaces that show the effects of supratidal and subaerial conditions. The presence of early fractures in particular lithofacies shows the importance of local synsedimentary tectonics during sedimentation. Thalassinoides, ?Arenicolites, Diplocraterion, Circolites, Gastrochaenolites and Trypanites are recorded in different beds of this section, reflecting various states of substrate consistency, in the form of firmground, hardground, and rockground. Whereas firmground conditions were dominant in the lower part of the section, hardgrounds and rockgrounds are mainly present in the upper part of the section. Four types of shallowing-upward elementary sequence are recognized. All the sequences show at the base mudstone or wackestone microfacies representing a lagoonal environment, overlain by sand-bar grain-pack-stone facies corresponding to a bar bounding the lagoon. The factors that controlled their development were carbonate production and tectonic movements.     

Silurian trace fossils in carbonate turbidites from the Alexander arc of southeastern Alaska

Early to Late Silurian (Llandovery‐Ludlow) body and trace fossils from the Heceta Formation of southeastern Alaska are preserved in the oldest widespread carbonates in the Alexander terrane. These fossils represent the earliest benthos to inhabit diverse shallow and deep subtidal environments in the region and are important indicators of early stages in benthic community development within the evolving Alexander arc. The ichnofossils are particularly significant because they add to a small but growing body of knowledge about trace fossils in deep‐water carbonates of Paleozoic age.

Carbonate turbidites that originated along a deep marine slope within the arc yield a low‐diversity suite of trace fossils consisting of five distinct biogenic forms. Simple burrows (Planolites, two forms), ramifying tunnels (Chondrites), and tiny cylindrical burrows (?Chondrites) represent the feeding activities (fodinichnia) of pre‐turbidite animals that burrowed in the lime mud before the influx of coarser sediment deposited by turbidity currents. These trace fossils are associated locally with cross‐cutting burrows created as domichnia (Palaeophycus). Rarer hypichnial burrows and endichnial traces were created by post‐turbidite animals that fed soon after the deposition of coarse detritus from turbidity flows.

Trace fossils in these deposits reflect much lower diversity levels than in Paleozoic siliciclastic turbidites. This difference may represent unfavorable environmental conditions for infaunas, differential preservation, or significant paleogeographic isolation of the Alexander terrane during the Silurian. Greater utilization of trace fossils in terrane analysis may help to resolve this issue and provide new data for reconstructing the paleogeography of circum‐Pacific terranes.     

Genetic relationships of tui chub populations in the northwestern Great Basin and conservation implications for the Cow Head Tui Chub

The Cow Head Tui Chub, Siphateles thalassinus vaccaceps, is restricted to the small Cow Head basin in far northeastern California and northwestern Nevada. Using microsatellite DNA loci, we found extensive genetic variation among tui chub populations in the Cow Head and surrounding basins of the northwestern Great Basin. Our data support the systematic conclusions of earlier authors, while showing distinct intraspecific differences between populations in most subbasins within the pluvial basins examined. The Cow Head Tui Chub forms a distinctive population within S. thalassinus with greatest affinity to the tui chub in the Warner Valley into which the Cow Head Lake drains. Tui chubs from the Goose Lake and Pit River basins are genetically very similar. Warner, Abert, Summer Lake, Catlow and Guano basin tui chubs appear distinct, both from each other and from tui chubs in the other basins. However, genetic proximity and some overlap between populations in presumably isolated hydrologic basins (i.e. Summer–Goose–Warner) suggest the possibility of unrecognized hydrologic connections or anthropogenic transport across barriers. Current genetic diversity in the Cow Head Tui Chub population is similar to that of stream-associated populations. We suggest that the Cow Head Tui Chub should remain a species of conservation concern due to its highly restricted distribution in an arid environment. Successful conservation of this unique fish will depend on stewardship of its limited habitat and maintenance of connectivity between subpopulations in a manner that ensures its long-term viability.     

Early Cambrian Teichichnus-dominated ichnofabrics and palaeoenvironmental analysis of the Caerfai Group, Southwest Wales, UK

The early Cambrian Caerfai Group of Pembrokeshire in Southwest Wales yields an intensely bioturbated Teichichnus-dominated ichnofabric, and provides evidence for the early evolution of infaunal organisms. Three units of the Caerfai Group comprise the focus of this study in sections exposed at Caerfai Bay. In the St. Non's Sandstone few primary sedimentary structures are observed, bedding being predominantly massive with subordinate parallel and current ripple cross-lamination. The topmost St. Non's Sandstone contains common soft-sediment deformation phenomena, and collectively deposition is believed to have taken place in the proximal subaqueous reaches of a steep-fronted delta system. Teichichnus is the dominant trace, with subordinate Planolites, Palaeophycus and possible Rhizocorallium.The overlying Caerfai Bay Shales were deposited from mass flows with evidence of downslope creep suggesting deposition on a steep delta front. Sandstone debrites are common, as are tuffaceous beds with evidence of reworking by bottom currents. Sandstone and tuff event beds contain common Skolithos and Arenicolites with less frequent Planolites. Interbedded mudstones contain infrequent Teichichnus.The Caerfai Bay Shales coarsen and thicken-upwards into the Caerbwdy Sandstone. Debrite sandstone beds dominate, and bed thickening upsection indicates progradation of the delta system. The dominant trace fossil within the unit is Teichichnus which is constrained to the top few centimetres of individual sandstone beds. Less common are Planolites and Palaeophycus.The size and abundance of Teichichnus in the Caerfai Group are similar to, or exceed those in contemporaneous sections in Avalonia and Baltica. Local environmental factors were probably the primary control on trace fossil distribution. Substrate and salinity are proposed as the main influences on ichnodiversity and size in the Caerfai Group. Teichichnus is most abundant in the St. Non's Sandstone, where the proximal reaches of the delta experienced reduced salinity and possibly higher oxygen levels. Burrow diameter progressively decreases upsection, and bioturbation becomes less pervasive in subsequent units of the Caerfai Group. This indicates that by the Cambrian Stage 2, significant advances into reduced salinity environments had occurred, along with the dispersal of the Teichichnus producer throughout the Avalonia-Baltica seaway. Correlation within Avalonia indicates that a transgression in the Cambrian Stage 2 initiated the deposition of the Caerfai Group marking the onset of the Welsh Basin.     

First occurrence of the Ichnogenus Selenichnites from the Middle Jurassic Strata of the Skoura Syncline (Middle Atlas,Morocco); Palaeoecological and palaeoenvironmental context

Mesozoic strata of North Africa yield the first occurrence of the ichnogenus Selenichnites. The trace fossils occur on the top surface of a sandy carbonate deposit in the axis of a Middle Atlas syncline (Skoura Syncline, NE Morocco). The ichnofossil-bearing horizon belongs to the Late Bajocian–Early Bathonian Ich Timellaline/Bou Akrabene Formation. The trace fossils are crescent-shaped and the best preserved exhibits a posterior central axial impression (possible telson tail impression). They are interpreted as feeding burrows (fodinichnia) or hiding depressions of Xiphosurids or Limulids (horseshoe crabs) on a sandy carbonate substrate beneath a veneer of muddy deposits. The sedimentological character suggests a relatively protected shallow water subtidal palaeoenvironment preceding the Bathonian regression of the Atlas domain. This discovery provides the first evidence of xiphosurans or xiphosuran-like organisms inhabiting the southern shores of the Tethys in the Middle Jurassic.     

Bio-deposition of a calcium carbonate layer on degraded limestone by Bacillus species

To obtain a restoring and protective calcite layer on degraded limestone, five different strains of the Bacillus sphaericus group and one strain of Bacillus lentus were tested for their ureolytic driven calcium carbonate precipitation. Although all the Bacillus strains were capable of depositing calcium carbonate, differences occurred in the amount of precipitated calcium carbonate on agar plate colonies. Seven parameters involved in the process were examined: calcite deposition on limestone cubes, pH increase, urea degrading capacity, extracellular polymeric substances (EPS)-production, biofilm formation, ζ-potential and deposition of dense crystal layers. The strain selection for optimal deposition of a dense CaCO₃ layer on limestone, was based on decrease in water absorption rate by treated limestone. Not all of the bacterial strains were effective in the restoration of deteriorated Euville limestone. The best calcite precipitating strains were characterised by high ureolytic efficiency, homogeneous calcite deposition on limestone cubes and a very negative ζ-potential.     

Stratigraphy and microfossils (radiolarians and planktonic foraminifers) of the Upper Cretaceous (upper Santonian–lower Campanian) Struganik limestone (Western Serbia)

Radiolarians and planktonic foraminifers were studied from several sections of the Upper Cretaceous lithographic limestones near Struganik Village (Western Serbia). These deposits were formed in a local basin, in a low-energy sedimentary environment surrounded by a shallow-water carbonate platform. The Struganik limestone is represented by alternating micritic limestone with calcarenite, rudite and tuffaceous rocks. The sections of the Struganik limestone have a narrow stratigraphic interval that covers the upper Santonian and probably includes the lower Campanian, with a total thickness reaching 120–150 m. The radiolarian zones Crucella robusta and Afens perapediensis were traced within the unit. The foraminiferal zone Dicarinella asymetrica was tentatively identified.     

Balanoglossites Mägdefrau, 1932 from the middle triassic of Germany: part of a complex trace fossil probably produced by burrowing and boring polychaetes

The ichnogenusBalanoglossites Mägdefrau, 1932 is re-described from its type area and is interpreted as part of a complex trace fossil comprising both burrow and boring components. The type ichnospecies,B. triadicus, consists of predominantly deep U- or Y-shaped tunnel elements, whereasB. ramosus n. isp. encompasses irregularly ramified galleries. Both ichnospecies occur frequently in firmground and hardground substrates with sporadic epikarst features. In the underlying softground, they may integrate intoLabyrintichnus terrerensis Uchman &; Álvaro, 2000, a network of irregular galleries and communicating shafts. On the omission surface, the tunnels are linked to elongated or winding grooves in hard or firm substrates. Further elements of the complex trace fossil include lobate burrows similar toZoophycos andRhizocorallium, mainly occurring in firmground and softground, shallow to deep U-shaped pouches and winding tunnels (Maeandropolydora sulcans Voigt, 1965) in hard and firm substrates. Finally, some tunnel portions and well preserved top surfaces of the ichnofabrics are scattered with the faecal pelletsCoprulus oblongus Mayer, 1952.C. oblongus has a rounded shape and a homogeneous structure and belongs to the Coprulidae n. ichnofam. Based on the numerous fingerprints preserved in the complex trace fossil, polychaete worms are identified as possible tracemakers. This is in accordance with the close combination of borings and burrows, a tunnel system highly variable in shape and size, blind ending side-branches and tunnel terminations, association with scolecodonts and exceptionally preserved polychaete remains. In addition, different commensals may have contributed to the modification of the complex trace fossil.     

Upper ordovician turbidites in western Asturias: A facies analysis with particular reference to vertical and lateral variations

Ab Upper Ordovician sandstone—shale sequence in Asturias, Northwest Spain, is analysed with particular regard to lithology, sedimentary structures and trace fossils. It is deduced that deposition of the sandstones was from axially flowing turbidity currents in a deep-water trough. Trace fossils are rare but there is a predominance of deep-water turbidite-facies types including Helminthopsis, Spirophycus, Cosmorhaphe, and Protopaleodictyon. Arenicolites, normally regarded as typical of shallow water, is, however, also recorded. The low abundance and diversity of trace fossils in this succession is in marked contrast to the situation in many younger turbidite sequences.The initiation of turbidite deposition followed a long period of quiet-water mud sedimentation. Recognition of this abrupt facies change in two sections separated by a down-current distance of 40 km, provides a reference level from which vertical and lateral facies changes are quantitatively documented. In each section, deposition starts with a fining and thinning-upwards cycle and continues with several similar cycles. The onset of turbidite deposition, abrupt facies transition, and repetition of the cycles, are probably largely the result of fault block uplift at source, while initial development of the deep-water trough may have resulted from associated downthrow. The lateral facies variations, which include downcurrent reduction in sandstone/shales ratio and the Walker ABC index, probably result from a combination of factors, including reduced flow velocity and decreased sediment load.     

Temperature effects on calcification rate and skeletal deposition in the temperate coral, Plesiastrea versipora (Lamarck)

The geographic range of the coral, Plesiastrea versipora (Lamarck, 1816), extends into temperate waters outside the southern limit for hermatypic corals. In the present study, calcification in Plesiastrea collected from Port Phillip Bay, Victoria was examined over the coral's normal annual temperature range (10-21 °C), which is well below the normal optimum for coral calcification in tropical corals (25-28 °C). Calcification rate in Plesiastrea was considerably lower than in reef corals, but showed a similar pattern in temperature responses, with a trend towards higher rates at ∼18 °C. The light/dark calcification ratio was markedly lower than that in tropical corals. Autoradiography showed that calcification occurred primarily by deposition of calcium carbonate at the upper surfaces of the septo-costae. Scanning electron microscopy (SEM) showed that skeletal deposition in Plesiastrea had a temperature-dependent diel pattern. In the light, calcium carbonate was deposited as small spheroidal crystals and, at higher temperatures, small needle-shaped crystals. In the dark, calcium carbonate deposition appeared to be in the form of an amorphous sheet-like cementation. Compared with other scleractinian corals, calcification rate in Plesiastrea was relatively slow and showed different patterns of skeletal deposition.     

Mayfly Burrows in Firmground of Recent Rivers from the Czech Republic and Poland,with Some Comments on Ephemeropteran Burrows in General

U-shaped, pouch-like burrows with parallel limbs, covered with short scratches arranged in sets, occur in the thalweg of the Oh?e river in NW Czech Republic. Similar, but smaller burrows with rare scratches, not arranged in sets, occur in the thalweg of the Drw?ca river in N Poland. Probably, they are produced by larvae and/or nymphs of Palingenia and Polymitarcis (Ephoron), respectively. In both localities, they burrowed in firmground surfaces at shallow depths. The burrowed surfaces were emerged during low water levels. A review of recent mayfly burrows shows that they are 1) U-shaped pouches with parallel limbs and septum, which may be covered with short scratches and are oriented perpendicular to the bottom, irrespective of its inclination, or 2) wide U-shape burrows with divergent limbs, which may be branched. In the fossil record, the ichnogenera Fuersichnus, Asthenopodichnium, and Rhizocorallium are partly ascribed to mayfly burrows, but their comparison to the recent burrows shows that such interpretations are somewhat problematic. The mayfly burrows are potentially good indicators of aquatic, non-marine, well oxygenated, clean water environments.     

A facies model for an Early Aptian carbonate platform (Zamaia, Spain)

The Cretaceous (Early Aptian, uppermost Bedoulian, Dufrenoyia furcata Zone) Zamaia Formation is a carbonate unit, up to 224 m thick and 1.5 km wide, which formed on a regional coastal sea bordering the continental Iberian craton. A high-resolution, facies-based, stratigraphic framework is presented using facies mapping and vertical-log characterization. The depositional succession consists of a shallow estuarine facies of the Ereza Fm overlain by shallow-water rudist limestones (Zamaia Fm) building relief over positive tectonic blocks and separated by an intraplatform depression. The margins of these shallow-water rudist buildups record low-angle transitional slopes toward the adjacent surrounding basins. Syn-depositional faulting is responsible for differential subsidence and creation of highs and lows, and local emplacement of limestone olistoliths and slope breccias. Two main carbonate phases are separated by an intervening siliciclastic-carbonate estuarine episode. The platform carbonates are composed of repetitive swallowing-upward cycles, commonly ending with a paleokarstic surface. Depositional systems tracts within sequences are recognized on the basis of facies patterns and are interpreted in terms of variations of relative sea level. Both Zamaia carbonate platform phases were terminated by a relative sea-level fall and karstification, immediately followed by a relative sea-level rise. This study refines our understanding of the paleogeography and sea-level history in the Early Cretaceous Aptian of the Basque-Cantabrian Basin. The detailed information on biostratigraphy and lithostratigraphy provides a foundation for regional to global correlations.     

Unusual nonmarine burrows from the Middle Jurassic of Scotland

The Kilmaluag Formation of the Great Estuarine Group (Middle Jurassic) of Scotland represents deposition of mixed carbonate and clastic sediments in a low‐salinity coastal lagoon to floodplain lake setting. Large, unusual trace fossils occur at two horizons within the formation. One type consists of platelike structures about 50 cm in diameter, which are found on wave‐rippled sandstone. These structures, strikingly similar to burrows produced by modern mudskippers, are assigned to fish that shallowly burrowed into the lagoon‐shore sediment.

The second type of burrows, found in brecciated, dolomitic limestones, are pipelike, about 4 to 7 cm in diameter and as much as 50 cm deep. One example has a chamber at the base of the pipe. Although most features of these structures appear similar to modern lungfish burrows, the chamber is most similar to structures produced by modern crayfish. The animal probably burrowed into the moist, mudflat sediment to escape desiccation during seasonal aridity.     

Trace fossils from lower Cambrian Hongjingshao Formation,Yunnan, China: Taxonomy,palaeoecology, palaeoenvironment

The Hongjingshao Formation (Cambrian Series 2, Stage 3) yields abundant, complex trace fossils that have not been systematically investigated up till now. Our detailed ichnological study of the traces from this formation in Malong area, Yunnan, China, recognizes four groups: (1) simple horizontal or sub-horizontal burrows; (2) complex branched burrow systems; (3) arthropod trails and; (4) simple vertical burrows belonging to the Cruziana ichnofacies. A trophic web is reconstructed for the community in Hongjingshao Formation based on both trace and body fossils. The ichnological and sedimentological features of the formation indicate an intertidal setting. The trace fossil assemblage demonstrates that Early Cambrian organisms were able to colonize very shallow marine environments, which further supports the landward expansion of the Early Cambrian ecosystems.