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1.
Summary During the Late Albian, Early and Middle Cenomanian in the NW part of the Adriatic Carbonate Platform (presentday Istria) specific depositional systems characterised by frequent lateral and vertical facies variations were established within a formerly homogeneous area, ranging from peritidal and barrier bars to the offshore-transition zone. In southern Istria this period is represented by the following succession: thin-bedded peritidal peloidal and stromatolitic limestones (Upper Albian); well-bedded foreshore to shoreface packstones/grainstones with synsedimentary dliding and slumping (Vraconian-lowermost Cenomanian); shoreface to off-shore storm-generated limestones (Lower Cenomanian); massive off-shore to shoreface carbonate sand bodies (Lower Cenomanian); prograding rudist bioclastic subaqueous dunes (Lower to Middle Cenomanian); rudist biostromes (Lower to Middle Cenomanian), and high-energy rudist and ostreid coquina beds within skeletal wackestones/packstones (Middle Cenomanian). Rapid changes of depositional systems near the Albian/Cenomanian transition in Istria are mainly the result of synsedimentary tectonics and the establishment of extensive rudist colonies producing enormous quantities of bioclastic material rather than the influence of eustatic changes. Tectonism is evidenced by the occurrence of sliding scars, slumps, small-scale synsedimentary faults and conspicuous bathymetric changes in formerly corresponding environments. Consequently, during the Early Cenomanian in the region of southern Istria, a deepening of the sedimentary environments occurred towards the SE, resulting in the establishment of a carbonate ramp system. Deeper parts of the ramp were below fair-weather wave base (FWWB), while the shallower parts were characterised by high-energy environments with extensive rudist colonies, and high organic production leading to the progradation of bioclastic subaqueous dunes. This resulted in numerous shallowing- and coarsening-upwards clinostratified sequences completely infilling formerly deeper environments, and the final re-establishment of the shallow-water environments over the entire area during the Middle Cenomanian.  相似文献   

2.
Summary Paleozoic carbonate ramp sedimentation has generally been described in terms of downlapping clinoforms composed of allochthonous sediment derived from shallower environments. However, during transgressive episodes when carbonate sediment production is low and down slope sediment transport by gravity becomes inactive, autochthonous carbonate sediment accumulates in vertical stacks of essentially in situ sediment. Autochthonous outer ramp deposition is probably a part of many Paleozoic ramp strata, but has heretofore not been recognized because of the general absence of adequate exposures. Evidence of autochthonous, in situ deposition and preservation of sediments in a starved setting is well displayed in the Alamogordo Member of the Lake Valley Formation in south central New Mexico. This evidence includes: 1) beds and bed sets that are individually continuous and traceable along ramp slope for 32 km, 2) down-ramp sequential distributions of depth-sensitive organisms and assemblages but patchy distribution of rock types, 3) lack of sedimentary structures indicative of transport, 4) well preserved, unabraded fossils, 5) the common occurrence of fossils in life position, 6) beds traceable into and through mounds 7) bed thickness trends ascribed to biotic productivity, and 8) geopetal structures in original position. Integrated paleontologic, sedimentologic, and stratigraphic data provide information about depositional processes and setting. The depositional slope was approximately 0.5o based on the distribution of fossil algae; this is comparable to dips reported for other Mississippian homoclinal ramps. An oxygen minimum zone may have impinged on the ramp during a major flooding event. Shifts in biotic gradients from bed to bed reveal transgressive-regressive patterns that would not be resolvable without detailed paleontological evidence. The Alamogordo Member formed as a result of transgressive and early highstand starved carbonate sedimentation along a narrow, homoclinal outer ramp. The surface of maximum flooding and the boundary between the TST and HST are within the Alamogordo Member.  相似文献   

3.
Worldwide Late Cambrian—Silurian lithofacies patterns indicate that the platforms of that time were sites of accumulation of two essentially different rocks suites: the platform carbonate rocks and the platform terrigenous rocks. Most of the platform rocks accumulated as sediments in shallow marine environments similar to those of the present but far more widely spread.Present-day marine benthic faunas are distributed in depth zones which are primarily controlled by temperature. Faunas tend to occur in substrate-related discrete clusters (communities) within each life zone; similar substrates in different depth zones commonly have different faunal associations. Individual phyletic stocks may encounter environmental optimum or near-optimum conditions in certain areas, that commonly are revealed by an abundance of species and individuals within species in each stock. Environmental optimum conditions depend upon availability of food that may be utilized, modes of feeding of the animals present, water motion, and substrate, among other factors. Organisms in past seas were distributed in patterns similar to those of the present.Carbonate platforms were particularly widespread during the latest Cambrian—Early Ordovician. Intertidal environments spread widely across those platforms during that time and characteristic faunal associations developed in them. Saukiid and related tribolites dominated latest Cambrian carbonate platform intertidal faunas. The Early Ordovician carbonate platform intertidal was dominated by archeogastropod-nautiloid cephalopod faunas. These animals were joined by tabulate corals and certain brachiopods during the latter part of the Ordovician and Silurian as prominent faunal elements in the carbonate platform intertidal—shallow subtidal. Cruziana and related trace fossils, bivalves, and certain tribolites (notably homalonotids and dalmanitids) dominated most terrigenous platform intertidal—shallow subtidal faunas of the Ordovician and Silurian.Articulate brachiopods (primarily orthoids, strophomenoids, and rhynchonelloids) appear to have been relatively prominent during the Early Ordovician in shallow subtidal environments on both carbonate and terrigenous platforms and to have spread down the bathymetric gradient into increasingly deeper subtidal areas of both platforms during the latter part of the Ordovician. Tribolites dominated faunas in relatively moderate to deep subtidal environments on both platforms during the early part of the Ordovician. They were gradually replaced by brachiopods in first the shallower, and later the deeper subtidal as dominant members of the faunas. Brachiopods (primarily pentameroids and spiriferoids) dominated nearly all Silurian warm-water subtidal environments from the shallow subtidal to the edges of the platforms.Platform uplifts in the Middle Ordovician and glacio-eustatic sea-level fluctuations in the Late Ordovician caused environmental changes across the platforms that were accompanied by marked replacements among marine benthic faunas in all environments. The distribution of Ordovician carbonate platforms and glacial deposits suggests that an Ordovician polar region may have been close to present-day equatorial Africa and that Ordovician warm temperate-tropical regions lay close to the present-day North Pole.  相似文献   

4.
In the clastic Genesee Group of the Catskill delta, lateral changes of the fauna are believed to reflect onshore-offshore physicochemical gradients. A shoreward increase of infauna is interpreted as adaptation to increased environmental stress. Free immobile taxa were concentrated offshore, while vagile forms, presumably able to cope with shifting substrata, are dominant nearshore. A shorewards replacement of brachiopods by bivalves reflects the eurytopy and infaunal habits of the bivalves.
In Gencsee time, progradation was first rapid, then slow. The sequence is reversed in the superjacent Sonyea Group and the accompanying reversal of faunal patterns is strong evidence of faunal control by the rate of progradation. This indicates the hazardous nature of attempts to trace 'community evolutioneditor' using only a few studies from each period.  相似文献   

5.
6.
Hardground surfaces from the Late Cambrian Snowy Range Formation in Montana/Wyoming are the oldest known non-reefal hard substrates exhibiting encrusting fossils. These surfaces range in age from Early Franconian to early Trempealeauan. Hardgrounds were developed on slightly hummocky to planar, truncated surfaces of glauconite-rich, carbonate, flat pebble conglomerates, which were deposited during episodes of storm scouring in shallow subtidal environments of the Montana/Wyoming shelf. Snowy Range hardgrounds are encrusted by a low diversity assemblage of fossils dominated by simple discoidal holdfasts of pelmatozoans, probably crinoids, and including small conical spongiomorph algae? and probable stromatolites. Macroborings (e.g. Trypanites) are notably absent from all hardground surfaces, although sharp-walled, vertical, cylindrical holes (borings?) occur in micrite clasts imbedded in certain flat pebble conglomerates. No evidence of faunal succession or microecologic partitioning of irregular surfaces was observed on these Cambrian hardgrounds.  相似文献   

7.
The Paleocene–Eocene Taleh Zang Formation of the Zagros Basin is a sequence of shallow-water carbonates. We have studied carbonate platform, sedimentary environments and its changes based on the facies analysis with particular emphasis on the biogenic assemblages of the Late Paleocene Sarkan and Early Eocene Maleh kuh sections. In the Late Paleocene, nine microfacies types were distinguished, dominated by algal taxa and corals at the lower part and larger foraminifera at the upper part. The Lower Eocene section is characterised by 10 microfacies types, which are dominated by diverse larger foraminifera such as alveolinids, orbitolitids and nummulitids. The Taleh Zang Formation at the Sarkan and Maleh kuh sections represents sedimentation on a carbonate ramp.

The deepening trends show a gradual increase in perforate foraminifera, the deepest environment is marked by the maximum occurrence of perforate foraminifers (Nummulites), while the shallowing trends are composed mainly of imperforate foraminifera and also characterised by lack of fossils in tidal flat facies.

Based on the facies changes and platform evolution, three stages are assumed in platform development: I; algal and coralgal colonies (coralgal platform), II; coralgal reefs giving way to larger foraminifera, III; dominance of diverse and newly developing larger foraminifera lineages in oligotrophic conditions.  相似文献   

8.
A litho-biostratigraphic analysis has been carried out in the Gador-Turon unit of the Sierra de Gador (Alpujarride complex, Betic Cordillera, SE Spain). The Triassic succession of this unit is composed of a lower meta-detrital formation overlain by an upper meta-carbonate formation divided in six members. In the latter, a Ladinian–Carnian-rich fossil association has been found (foraminifers, algae, bivalves, microproblematica, trace fossils). Facies analysis has enabled the recognition of 22 facies of platform origin. This succession accumulated as a subsiding margin-type carbonate platform with homoclinal ramp geometry (Anisian?–Ladinian) evolving into a fault-block-type platform with a steeper-margined geometry (Ladinian–Carnian). Slope deposits of this latter platform show a prism-like geometry with progradational patterns and include syn-sedimentary structures associated with normal faults capped by younger beds. The results of the present research indicate that the architecture of the platform studied has been controlled mainly by climate and oceanic factors during the development of the ramp, and by syn-sedimentary extensional tectonics during the development of the steeper-margined platform. The Ladinian–Carnian tectonic activity was probably also responsible for the siliciclastic input and the shift to a mixed terrigenous-carbonate platform.  相似文献   

9.
We have obtained new insights into the behavior of a class of excitable systems when a stimulus, or parameter, is slowly tuned through a threshold value. Such systems do not accommodate no matter how slowly a stimulus ramp is applied, and the stimulus value at onset of repetitive activity shows a curious, nonmonotonic dependence on ramp speed. (Jakobsson, E. and R. Guttman. Biophys. J. 1980. 31:293-298.) demonstrated this for squid axon and for the Hodgkin-Huxley (HH) model. Furthermore, they showed theoretically that for moderately slow ramps the threshold increases as the ramp speed decreases, but for much slower ramp speeds threshold decreases as the ramp speed decreases. This latter feature was found surprising and it was suggested that the HH model, and squid axon in low calcium, exhibits reverse accommodation. We have found that reverse accommodation reflects the influence of persistent random fluctuations, and is a feature of all such excitable systems. We have derived an analytic condition which yields an approximation for threshold in the case of a slow ramp when the effect of fluctuations are negligible. This condition predicts, and numerical calculations confirm, that the onset of oscillations occurs beyond the critical stimulus value which is predicted by treating the stimulus intensity as a static parameter, i.e., the dynamic aspect of a ramp leads to a delay in the onset. The condition further demonstrates a memory effect, i.e., firing threshold is dependent on the initial state of the system. For very slow ramps then, fluctuations diminish both the delay and memory effects.(ABSTRACT TRUNCATED AT 250 WORDS)  相似文献   

10.
The Late Cretaceous Laguna Palacios Formation in Central Patagonia (San Jorge Basin), southern South America, is composed of tuffaceous deposits supplied by periodical volcanic ash falls partly reworked by rivers, on broad plains. Variations in ash-fall rates allowed the formation of stacked, mature paleosols, which are one of the most characteristic features of this formation. The mature paleosols show well-developed horizons, ped structure and bear an intricate network of trace fossils mostly produced by small roots and invertebrates. Two different insect trace fossils could be recognized in this formation: sweat bee nests and coleopteran pupal chambers. Fossil bee nests are composed of inclined tunnels with cells attached to them by means of short necks, a typical construction of bees of the subfamily Halictinae. Similar halictine constructions were reported from the Cretaceous of the USA. Coleopteran pupal chambers are discrete, ovoid structures, having an internal cavity with a smooth surface, and an outer wall of lumpy appearance composed of different layers of soil material. They are commonly constructed by the larvae of different families of Coleoptera. Similar trace fossils were previously reported from the Asencio Formation (Late Cretaceous–Early Tertiary) of Uruguay and from the Djadokhta Formation of Mongolia (Late Cretaceous). These trace fossils constitute some of the only paleontological data from the Laguna Palacios Formation, allowing inferences about its paleoecology, paleoclimatology and paleogeography. Ecological preferences of Halictinae, as well as some features of the nests, suggest a temperate, seasonal climate and an environment dominated by low vegetation for the Laguna Palacios Formation, which is also compatible with sedimentologic and pedogenic evidence. The morphology of the nests, typical of North American halictinae, adds more evidence to the hypothesis of the existence of faunal interchange between North and South America by the Late Cretaceous. The fossil nests constitute some of the oldest evidence of bees in the fossil record, the third known record of bees of Cretaceous age and the first for the Southern Hemisphere. The two traces described are, together with those of Dakota and the Gobi Desert, the only trace fossils from paleosols of Cretaceous age that can be certainly attributed to insects.  相似文献   

11.
Four benthic marine communities occur in the clastic facies of the prograding Upper Frasnian-Lower Famennian (Upper Devonian) Foreknobs Formation in the Central Appalachians along the Allegheny Front in Maryland, West Virginia, and Virginia. Deep-water, rapidly prograding environments were inhabited by the Ambocoelia-Chonetes Community, dominated by an epifauna of unattached brachiopods. Offshore bar environments were inhabited by the Cyrtospirifer-Camarotoechia Community, exhibiting adaptations to shallow-water, high-energy conditions and probably lowered salinities. Shallow-water, sublittoral environments were inhabited by the Atrypa-Cypricardella Community, a community in which existed a variety of life habitats and a diverse epifaunal and infaunal association of brachiopods and bivalve molluscs. The Leptodesma-Tylothyris Community flourished in nearshore bar-protected environments in the southern region of the study area, whereas in the north the Cyrtospirifer-Camarotoechia Community inhabited nearshore environments in conjunction with the onshore development of a large fluviodeltaic system.  相似文献   

12.
Summary The Cabo Cope Unit, which outcrops east of Aguilas (Murcia), belongs to the Maláguide tectonic Complex (Betic Internal Zone) and displays stratigraphic characteristics of particular interest, including Triassic bioclastic carbonate beds which are not common in the Maláguide units. Biostratigrafic fossils have been found in these beds and may correlate with Triassic alpine biofacies. Alpine fauna fossils only appeared in those palaeogeographic units of the Internal Zone of the Cordillera referred to as Alpujárride units, while the influence of the Sephardic faunal province is evident in almost all the cordillera. For these reasons it is noteworthy that new alpine fauna fossils have been found in an Internal Zone unit in which relevant fossils rarely appear. The Triassic succession of the unit studied in this paper can be subdivided into two members: a lower one, which is clastic and contains thick gypsum beds, and an upper one, consisting of carbonate rocks. The lower member has been interpreted as a fluvial-coastal deposit. The upper member is interpreted as a sequence of carbonate ramp deposits. This ramp evolved into a shallow platform with tidal flats typical of a coastal zone. The bivalve fossilsDaonella cf.lommeli (Wissmann) and “Posidonia” sp. have been found in the carbonate member, along with the conodontSephardiella mungoensis (Diebel). These fossils are of the Late Ladinian age and have been found only in this outcroup of the Betic Cordillera. The presence of this fossil assemblage, which belongs to the alpine faunal province, indicates a connection during the Late Ladinian between the Tethys sea and this area of the Maláguide palaeogeographic domain. The palaeogeographic location of the Cabo Cope Unit during the Middle Triassic was at the south-easternmost part of the Betic Basin, implying that the connection between the Tethys and the Betic Basin was established in the easternmost domains of the basin.  相似文献   

13.
The late Miocene is a crucial interval for global climate evolution as well as for the regional geodynamic evolution of the Central Mediterranean area. It spans the transition from the warm Mid Miocene Climatic Optimum, associated with the major Monterey Carbon Isotope Excursion, to the cooler Pliocene, characterized by a bipolar glaciation. Within this climatic transition, during the early Tortonian, a positive carbon isotope excursion related to a global carbon cycle perturbation is recorded, named Carbon Maximum 7 (CM7). In this study, two upper Serravallian–lower Tortonian carbonate ramps of the Central Mediterranean have been analysed: the Latium‐Abruzzi and the Apula ramps (Central Apennines, Central Italy). Carbon isotope ratios have been coupled with facies and microfacies analyses with the aims: (1) to identify the CM7 in the Central Mediterranean carbonate ramp successions; (2) to evaluate potential carbonate factory changes or demises related to the CM7; and (3) to discriminate the role of global vs regional factors in affecting the Mediterranean carbonate ramps. The two studied ramps show different evolutions due to regional geodynamics causes. The Latium‐Abruzzi ramp drowns in the Tortonian, while the Apula shows a regressive trend, with upper Tortonian middle ramp facies overlying the lower Tortonian outer ramp. Despite the complex geodynamic setting, a positive carbon isotope shift has been identified in both ramps and correlated with the CM7. This positive carbon isotope shift may be linked to the strengthening of the global ocean circulation during the late Miocene cooling. This strengthening of the circulation enhanced the coastal upwelling, bringing nutrient‐rich waters to the surface and triggering an enhanced primary productivity consistent with the CM7.  相似文献   

14.
S.G.Pemberton   《古生物学报》2001,40(1):134-142
介绍地质记录中在不稳定的沉环境下,由于生活条件的恶化(盐度的变,严酷的温度,低含氧量及移动的沉积基底等)以及沉积事件(浊流沉积和风暴沉积)所产生的机会(r-选择)遗迹化石和均衡(K-选择)遗迹化石及其特点以及机会行为方式的古生物证据。对现代底栖生物的生态学研究,揭示现代机会生物均衡生物的生态特征(居群策略)描述由于破坏事件导致动物群消除后重新移居的现代实例,有助于了解地质记录中机会(r-选择)遗迹化石和均衡(K-选择)遗迹化石的成因及其古生态特征。  相似文献   

15.
The oldest Mesozoic nearshore Zoophycos: evidence from the German Triassic   总被引:1,自引:0,他引:1  
The trace fossil Zoophycos has been described from the Middle Triassic carbonates of the German Basin for the first time. It occurs in a calcilutite bed at the top of a shallowing-upward cycle (parasequence) in the transgressive systems tract of the Middle to Upper Muschelkalk sequence of Thuringia (Germany). Based on sedimentological and palaeontological features, the studied interval is interpreted as deposited in a marine nearshore environment with proximal storm deposits (tempestites). Zoophycos occurs in a very simple planar form with lobate spreiten, which were most likely produced by a worm-like animal by strip mining. The upper tier of the ichnofabric consists of Zoophycos, whereas the lower tier is occupied by cylindrical trace fossils of unknown taxonomic affiliation and with decreasing size towards the bottom. Associated trace fossils such as Rhizocorallium, Balanoglossites and Trypanites indicate a partly firm to hard substrate. No mixed layer is developed at the top of the trace fossil bearing succession. The ichnofabric together with the sedimentological features (disseminated pyrite, blue-grey colour) and palaeontological circumstances (poor benthic fauna, meiofauna with a small body size) support an interpretation of a dysaerobic environment. In the view of evolutionary change, Palaeozoic Zoophycos occurs in both deep and shallow marine deposits, whereas Mesozoic and Cenozoic Zoophycos is only common in shelfal and deeper-marine deposits. The new finding from the shallow-marine Middle Triassic represents the first reliable occurrence of Zoophycos after the end-Permian mass extinction and shows close similarities to its Palaeozoic precursors. It demonstrates that the producer survived the end-Permian mass extinction, became re-established in the nearshore realm and progressively colonized deeper-marine environments during the Mesozoic and Cenozoic.  相似文献   

16.
The Jahrum Formation was deposited in the foreland basin in southwest Iran (Zagros Basin). The Zagros mountain belt of Iran, a part of the Alpine–Himalayan system, extends from the NW Iranian border through to SW Iran, up to the strait of Hormuz. The various facies of the Jahrum Formation were deposited in four main genetically related depositional environments, including: tidal flat, lagoon, shoal and open marine. These are represented by 14 microfacies. The Jahrum Formation represents sedimentation on a carbonate ramp. Tidal flat facies are represented by fenestral fabric, stromatolitic boundstone and thin-bedded planes. Carbonate deposition in a shallow marine lagoon was characterised by wacke–packstone, dominated by various taxa of imperforate foraminifer. The shoals are made up of medium- to coarse-grained skeletal and peloidal grainstone. This facies was deposited predominantly in an active high energy wave and current regime, and grades basinward into middle ramps facies are represented by wackestones–packstones with a diverse assemblage of echinoderm and large benthic foraminifers with perforate wall. Outer ramp facies consist of alternating marl and limestones rich in pelagic foraminifera. There is no evidence for resedimentation processes in this facies belt. The sequence stratigraphy study has led to recognition of three third-order depositional sequences.  相似文献   

17.
The timing of recovery after the end‐Permian mass extinction has been a matter of debate, with some authors favouring a more rapid faunal recovery during the Early Triassic and others considering a more protracted biotic reestablishment spanning until the Middle Triassic. In this work, we investigated the lowermost Middle Triassic (Ladinian) carbonate deposits in the Catalan Basin to evaluate the potential environmental mechanisms and evolutionary constrains involved in the kilometre‐scale predominance of microbialites and the low‐diversity and high‐density Planolites association in a low‐latitude epicontinental setting. The studied sedimentary succession records the development from a low‐gradient, homoclinal microbial‐dominated carbonate ramp evolving towards a slightly inclined swell‐dominated type. Sedimentological analysis suggests that facies heterogeneity was controlled by pulses of syn‐rift tectonic activity, which compromised Peri‐Tethyan basin connectivity, reducing palaeobathymetry gradients. Although the monospecific nature of the studied trace‐fossil association may reflect the delayed recovery after the end‐Permian mass extinction, this is inconsistent with widespread, relatively high‐diversity ichnofaunas in carbonates elsewhere in the region. Since other Ladinian basins were characterized by the recurrence of microbial carbonates, low‐diversity ichnoassemblages and limited skeletal production, we hypothesize that shallow and restricted carbonate ramp settings harboured limited ecological complexity and widespread opportunistic colonization of the sediment when compared to coeval open marine locations.  相似文献   

18.
For lacking of absolute dating, and because of the strong mammalian faunal endemism, the Chinese late Neogene has long been subdivided into mammalian ages mostly based on isolated local faunas by evolutionary stages of fossil taxa. The correlation with European stages and or mammalian units remains unsatisfied. We report the recent advances of the land mammal biochronology of late Neogene based on fossils found from continuous sections located at Lantian, Shaanxi Province and Lingtai, Gansu Province. There recognized three biozones for the late Miocene by fossils from the Bahe and Lantian formations, and three biozones for Pliocene on the data from Lingtai. The detailed biochronology work in combination with paleomagnetic data may improve the precision of correlation in the continental scale.  相似文献   

19.
The Maastrichtian shallow-water carbonate platform (Tarbur Formation) is described from outcrop in southwest Iran. It is characterised by eight microfacies types, which are dominated by larger foraminifera, rudist debris and dasycladacean algae. They are grouped into four distinct depositional settings: tidal flat, lagoon, barrier and open marine. The depositional settings include stromatolitic boundstone of tidal flat, peloidal dasycladacean miliolids wackestone and peloid bioclastic imperforate foraminifera wackestone of restricted lagoon, Omphalocyclus miliolids bioclast packstone–grainstone and miliolids intraclast bioclast packstone–grainstone of open lagoon, rudist bioclast grainstone of inner-platform shoals and rudist bioclast floatstone–rudstone and bioclastic wackestone of open-marine environments.

The facies and faunal characters are typical of a ramp-like open shelf. The lack of reef-constructing organisms resulted in a gently dipping ramp morphology for the margin and slope. On the basis of facies analysis, three depositional sequences (third order) are defined.  相似文献   

20.
The Purbeckian facies of the northern margin of the Aquitaine Basin is exposed in the Cherves-de-Cognac section (SW France), considered as basal Cretaceous in age. Two lithological units have been identified. The basal unit (U1) is composed of an alternation of gypsum and finely bedded black dolomitic marlstone, occasionally stromatolitic in nature. It represents mostly hypersaline, dolomitic tidal flat, lagoonal, and sabkha-type environments. The upper unit (U2) consists of variably fossiliferous limestone-marl alternations that contain a diverse fauna and flora. In the upper part of this unit, a 4-m-thick level has yielded a rich vertebrate fauna. The preservational quality of the vertebrate fossils varies within the section. On the basis of faunal, floral, sedimentological and mineralogical information the vertebrate remains accumulated in brackish water environments in which the influence of freshwater has progressively increased towards the top of the section. The close association of environments representing varying degrees of marine and continental influence suggest an estuarine setting.  相似文献   

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