A morphological revision of the genus Herbertus S.Gray (Herbertaceae,Marchantiophyta) in Africa,including the East African islands

Abstract

The genus Herbertus S.Gray in Africa and the East African islands is revised. The division of the genus into two broad species complexes centred on H. dicranus (Tayl.ex Gottsche et al.) Trev. and H. juniperoideus (Sw.) Grolle is confirmed. H. capensis (Steph.) Sim, H. capillaris (Steph.) H. doggeltianus (Steph.) Demaret, H. lobatus (Steph.) Demaret, H. mascarenicus (Steph.) S.Arn., H. mollis (Steph.) Dusén, H. stuhlmannii (Steph.) Demaret and Schisma kilimandjarense Steph. are synonymous with H. dicranus. H. grossevittatus (Steph.) S.Arn. ex Grolle is synonymous with H. juniperoideus. Three new species are described: H. spicatus N.G.Hodgetts sp. nov., which is related to H. juniperoideus and appears close to the neotropical H. pensilis, with long, narrow leaf lobe apices and a sheathing leaf base; H. mauritianus N.G.Hodgetts sp. nov., also related to H. juniperoideus, has fewer basal slime papillae and the vitta bifurcating lower down; and H. pocsii N.G.Hodgetts sp. nov., which is related to H. dicranus but has setaceous leaf lobe apices composed of long, narrow cells. The relationships of the taxa and patterns of distribution, taking account of recent molecular work, are discussed.     

Miscellanea hepaticologica 141-150

Abstract

141. Anomacaulis fiaccidus (Steph. 1917) Grolle c. n. has to replace A. hamatilobus (Grolle 1965) Schust. 142. Colura leratii (Steph. 1908) Steph. Has to replace C. apiculata Schiffn. ex Steph. 1916. 143. Herbertus sendtneri (Nees) Lindb. 1874 has to replace H. sendtneri (Nees) Evans 1917. 144. The redetected type of Herbertus stramineus (Dum.) Trev. in BR wholly justifies the recent usage of that name instead of H. aduncus auct. 145. The large styli of Jovetastella Tixier were misinterpreted as amphigastria. Jovetastella is reduced to subgeneric rank within Cololejeunea (Spruce) Schiffn., necessitating C. subg. Jovetastella (Tixier) Grolle n. st. et c. and C. paniensis (Tixier) Grolle c. n. 146. Lichenastrum Dill. 1811 is lectotypified with Jungermannia Ianceolata L. emend. Grolle, thus becoming a synonym of Jungermannia L. s. str. 147.Lophozia elongata Steph. has to replace L. elongata (Lindb. ex Kaal.) Steph. 148. Mannia triandra (Scop. 1772) Grolle c. n. has to replace M. rupestris (Nees 1817)Frye et Clark. A redetected type specimen of M archantia ludwigii Schwaegr. 1814 turned out to be Mannia triandra. Therefore Asterella gracilis (F. Web.) Underw. has to replace A. ludwigii auct. 149. Grimaldia chilensis Lindenb. ex Mont. 1839 was newly placed in synonymy of Sauteria berteroana Mont. 1839. 150. Telaranea nematodes (Gottsche ex Aust. 1879) Howe has to replace T. sejuncta auct., whereas the true T. sejuncta (Angstr. 1877) S. Arn. is a synonym of Arachniopsis diacantha (Mont. 1856) Howe. T. sejuncta yare breviseta (Herz.) Fulf. is still another species, thus the single record of T. nematodes or T. sejuncta from Juan Fernandez is rejected.     

Miscellanea hepaticologica 201-210

Abstract

201. Calycularia crispula Mitt. new to Mexico and the whole America. It is the first record outside Asia. 202. A type study of Fimbriaria nana Lindenb. revealed that it is synonymous with Athalamia hyalina (Sommerf.) Hatt. 203. Fossombroniaceae Hazsl. 1885 has to replace Codoniaceae Klinggr.1858 nom. illeg., because Codonia Dum. 1822 is illegitimate. 204. Herzogobryumaterrimum (Steph.) c. n. (≡ Cephalozia aterrima Steph.) has to replace H. filarium Grolle 1975. 205. Korrnickia Steph. 1917 is regarded as a synonym of Symphyogyna Nees et Mont. 1836 and K. cylindrica Steph. 1917 is placed in synonymy of S. mexicana Steph. 1909.206. The detection of the perianth of Lophocolea spectabilis Steph. (≡ Heteroscyphus arnellii Fulf.) necessitates a transfer as Leptoscyphus spectabilis (Steph.) c. n. 207. All records of Preissia Corda from Latin America are rejected. P. barbata Herz. 1921 is referred to as syn. nov. of Marchantia chenopoda L. 1753. 208. Riccia crustata Trab. was validly published in 1916 already, but has been generally overlooked hitherto. 209. Symphyogyna brasiliensis. Nees et Mont. is widespread in the Neotropics and Africa. Several African names are treated as synonyms. 210. The monotypic genus Wiesnerella Schiffn. is newly recorded from New Guinea and the Mascarenes. The Mascarene record is the first one outside Asia.     

Jamesoniella carringtonii—eine Plagiochila in Nepal mit Perianth

Abstract

Jamesoniella carringtonii (Balfour) Schiffn. is transferred to Plagiochila (P. carringtonii (Balfour) Grolle, comb.nov.). P. carringtonii ssp. carringtonii is regarded as an endemic subspecies of the British Isles and the Faeroes, whereasssp. lobuchensis Grolle, ssp.nov. with wholly entire leavesis restricted to Nepal and bears perianths. Jamesoniella carringtonii var. recurvata Nicholson is an independent not closely related species of Plagiochila (P. recurvata (Nicholson) Grolle, stat. et comb.nov.) from N.W.-Yünnan. P. recurvata and p. poeltii Inoue et GroUe, sp.nov. (from Nepal) probably belong to section Renitentes Carl. New differences between Jamesoniella autumnalis (DC.) Steph. and J. undulifolia (Nees) K. MOO., whichlatter is type of Biantheridion Grolle, sect.nov. Notes on the distribution of hepatics in Europe restricted to the cool euatlantic regions.     

On Plagiochila section Cobanae Carl in Asia and melanesia

Section Cobanae Carl of the genus Plagiochila is characterized. Several Asian and Melanesian species are included in the section: Plagiochila bicornuta Steph., P. detecta So & Grolle, P. singularis Schiffn., P. tagawae Inoue, P. tixierí Inoue, P. trabeculata Steph. and P. zhuensis Grolle & So. Plagiochila laxissima Schiffn. is placed under the synonymy of P. bicornuta Steph.     

Mrican hepatics. XXXIV. Some little-known or new Lejeuneaceae

Abstract

Lejeunea aethiopica E. W. Jones, sp. nov., L. amaniensis E. W. Jones sp. nov., L. hepaticola Steph., and Cheilolejeunea cordistipula (Steph.) Grolle ex Jones, 397 comb. nov. (Strepsilejeunea cordistipula Steph., Pycnolejeunea angustiflora Steph., syn. nov.) are described.     

Studies on Plagiochila (Hepaticae) in Australasia and the Pacific II

Fourteen species names of Plagiochila are newly placed in synonymy : P. aneitiana Steph. (= P. trapezoidea Lindenb.), P. colonialis Steph. (= P. arbuscula (Brid. ex Lehm. & Lindenb.) Lindenb.), P. comptonii Pearson (= P. arbuscula), P. innovans Steph. (= P. trapezoidea), P. laciniata Pearson (= P. arbuscula), P. leptocaula Gottsche ex Dugas (= P. bicornuta Steph.), P. longa Dugas (= P. arbuscula), P. microphyces Gottsche ex Dugas (= P. bicornuta Steph.), P. palmicola Steph. (= P. arbuscula), P. santoensis Steph. (= P. bicornuta), P. subangulata Steph. (= P. bialata Mitt.), P. taitica Gottsche ex Dugas (= P. arbuscula), P. tenuitexta Steph. (= P. propinqua Sande Lac.), and P. vulgarifolia Steph. (= P. trapezoidea Lindenb.). Female plants of P. bialata, P. bicornuta. and P. inflata Steph. are illustrated.     

Distribution Maps of Bryophytes in Britain

Abstract

The following species of Cheilolejeunea subgen. Strepsilejeunea are recognised from Africa and described: C. krakakammae (Lindenb.) Schust., C. pluriplicata (Pears.) Schust., C. usambarana (Steph.) Grolle, C. convexa S. Arn., C. pocsii E. Jones, sp. nov., C. camerunensis S. Arn. Strepsilejeunea georgensis S. Arn. is a synonym of C. krakakammae.     

New records of bryophytes from the Canary Islands

Abstract

Two members each of the Adelanthaceae and Schistochilaceae and three of the Porellaceae are recorded from Mulanje Mountain, of which Adelanthus lindenbergianus (Lehm.) Mitt., Porella abyssinica (Nees) Trev. var. hoehnelii (Steph.) Pócs, Porella capensis (Gott.) Mitt. and P. subdentata (Mitt.) Steph. are first records for Malawi.     

Studies on Plagiochila in Asia,supplements to sections Abietinae,Annotinae, Ciliatae,Contiguae, Cucullatae,Poeltiae, Subtropicae and zonatae

Twenty-seven names are newly placed in synonymy, Plagiochila blepharophora (Nees) Lindenb. var. δ major Schiffn. (= P. sandei Dozy), P.boninensis Inoue (= P. fordiana Steph.), P. cadens Inoue (= P. sciophila Nees), P. cameronensis Inoue (= P. hampeana Gottsche), P. ceramica Inoue (= P. pulvinata Steph.), P. decidua Inoue & Grolle (= P. sciophila Nees), P. didyma Inoue (= P. parvifolia Lindenb.), P. euryphyllon Herzog (= P. sciophila Nees), P. flavovirens Steph. (= P. sciophila Nees), P. fraseri Steph. (= P. teysmannii Sande Lac), P. gedeana Schiffn. (= P. hampeana Gottsche), P. kaernbachii Steph. (= P. sandei Dozy), P. lagunensis Inoue (= P. junghuhniana Sande Lac), P. massalongoana Sande Lac. (= P. junghuhniana Sande Lac), P. minor Horik. (= P. fordiana Steph.), P. nilgherriensis Steph. (= P. semidecurrens (Lehm. & Lindenb.) Lindenb.), P. nubila Steph. (= P. sandei Dozy), P. nymanii Steph. (= P. sandei Dozy), P. orientalis Taylor (= P. sciophila Nees), P. seemannii Mitt. (= P. sandei Dozy), P. sepikensis Inoue (= P. inflata Steph.), P. spinosissima Steph. (= P. semidecurrens (Lehm. & Lindenb.) Lindenb.), P. subplanata Inoue (= P. sciophila Nees), P. trabeculata var. bifida S. Hatt. (= P. fordiana Steph.), P. trochantha Steph. (= P. sciophila Nees), P. tsutomui Inoue (= P. ungarangana Sande Lac), and P. wichurae Steph. (= P. renitens (Nees) Lindenb). Lectotypes are designated for P. junghuhniana, P. semidecurrens, and P. ungarangana. Plagiochila inflata Steph. is reinstated as a species of sect. Annotinae Carl.     

Morphological and phytochemical studies of Plagiochila papillifolia Steph., a Neotropical liverwort new to Europe

Abstract

The Neotropical liverwort Plagiochila papillifolia Steph. is conspecific with P. caespiticia Spruce ex Herzog (nom. inval.), P . deciduifolia Steph., P . solmsii Steph. and P . verruculosa R.M.Schust.; it is reported from the Azores, new to Europe. Morphologically it shows affinity with sect. Bidentes Carl; phytochemically, several of its lipophilic secondary metabolites are the same as those of P . retrorsa Gottsche, another Neotropical species reported only recently from Macaronesia.     

Distribution and synonymy of Plagiochila punctata (Taylor) Taylor, with hypotheses on the evolutionary history of Plagiochila sect. Arrectae (Plagiochilaceae, Hepaticae)

For a long time, Plagiochila (sect. Arrectae) punctata (Taylor) Taylor has been treated as an endemic of Atlantic Europe. Studies of larger specimen sets demonstrated that the species is widespread in mountainous regions of the Neotropics where it is known under several names including P. stolonifera Lindenb. & Gottsche, P. choachina Gottsche and P. patzschkei Steph. In tropical Africa P. punctata is established as Plagiochila subalpina Steph., nom. illeg. The sporophyte of P. punctata is described for the first time, based on material from Costa Rica. A maximum likelihood analysis of a nrITS sequence alignment with sequences of P. sects. Fuscoluteae (outgroup), Arrectae, and Rutilantes as well as of P. rubescens (Lehm. & Lindenb.) Lindenb. results in a tree with well bootstrap supported Arrectae and Rutilantes clades. ITS sequences of Plagiochila punctata from the Comoros and Zaire are placed in an unsupported monophyletic lineage together with P. punctata sequences from the British Isles and Ecuador. The species is nested in a robust clade with P. spinulosa (Dicks.) Dumort. and P. stricta Lindenb. The weak genetic separation of these three species indicates a recent diversification; the disjunct ranges may be the result of long range dispersal events. Plagiochila rubescens from southern South America is placed sister to P. sect. Arrectae. The latter section possibly originated from southern South American ancestors, diversified in tropical America after the uplift of the Andes and reached the Holarctics and tropical Africa by long range dispersal of diaspores.We are grateful to the curators and owners of the herbaria cited in the text for the loan of specimens, especially to Tamás Pócs, Volker Buchbender and Eberhard Fischer for their continuous support with recent collections from tropical Africa. David Rycroft kindly sent a duplicate of P. rubescens. Financial support of the Deutsche Forschungsgemeinschaft (grant HE 3584/1) is gratefully acknowledged.     

Recent Bryological Literature, 68

Abstract

Five species of Colura are reported from Mulanje Mountain, of which four, C. ealyptrifolia (Hook.) Dum., C. digitalis (Mitten) Steph., C. hattoriana Pócs and C. tenuicornis (Evans) Steph. are new for Malawi. C. hattoriana is also reported new for mainland Africa. A key to sub-Saharan taxa is provided.     

Bryophytes from the southeastern Yukon Territory,Canada

One hundred and forty-five taxa of bryophytes are reported from the extreme southeastern Yukon. Eight species of Hepaticae and 13 of mosses represent new records for the Yukon, which are Calypogeia suecica (S. Arnell & Perss.) K. Müll., Chiloscyphus pallescens (Ehrh.) Dum., Conocephalum conicum (L.) Dum., Jamesoniella autumnalis (DC.) Steph., Pellia neesiana (Gott.) Limpr., Ptilidium pulcherrimum (Web.) Hampe, Riccardia palmata (Hedw.) Carruth., Tritomaria exsecta (Schmid.) Schiffn., Amblystegium varium (Hedw.) Lindb., Brachythecium rivulare B.S.G., B. rutabulum (Hedw.) B.S.G., Bryum blindii B.S.G., Didymodon rigidulus Hedw., D. tophaceus (Brid.) Lisa, Drepanocladus trichophyllus (Warnst.) Podp., Hygroamblystegium noterophilum (Sull. & Lesq. ex Sull.) Warnst., Hygrohypnum molle (Hedw.) Loeske, Plagiomnium ciliare (C. Müll.) Kop., P. cuspidatum (Hedw.) Kop., Platydictya minutissimum (Sull. & Lesq. ex Sull.) Crum, and Pylaisiella selwynii (Kindb.) Crum, Steere & Anderson. Many of the other collections represent wide extensions of range within the Yukon Territory.     

African Hepatics. XXXI. Some new or little-known Lejeuneaceae and extensions of range

Abstract

Lejeunea brenanii sp. nov. and Microlejeunea ankasica sp. nov. are described. Lejeunea autoica Schuster and Pycnolejeunea contigua (Nees) Grolle are described and recorded as new to Africa. Lejeunea longirostris (Steph.) E.W. Jones, nom. inval., is a synonym of L. grossecristata (Steph.) E.W. Jones. Critical notes and/or new localities are provided for Colura digitalis (Mitt.) Steph., C. dusenii Steph., C. humbertii S. Jovet-Ast, C. sp. ad inornata Jovet-Ast, Marchesinia moelleriana Pearson.     

Chromosomes of African Hepatics—Jubulae

Abstract

The chromosome numbers of five species belonging to the Jubulae have been described, and are as follows:

Lejeuneaceae

Cololejeunea cf. dissita, n = 9.

Arehilejeunea autoiea Vanden Berghen, n=9.

Caudalejeunea hanningtonii (Mitt.) Schiffn., n =9.

Mastigolejeunea florea (Mitt.) Steph., n=9.

Frullaniaceae

Frullania spongiosa Steph., female, n = 9.

F. spongiosa Steph., male, n=8.

The author wishes to thank Dr E. W. Jones for assistance in identifications, especially with Cololejeunea cf. dissita.     

中国大陆一苔类新记录种——膨体异萼苔

报道了产自中国大陆贵州省的异萼苔属(Heteroscyphus)一新记录种--膨体异萼苔[H. turgidus (Schiffn.) Schiffn.]。本种与圆叶异萼苔[H. tener (Steph.) Schiffn.]主要的区别特征在于腹叶边缘具粗齿且植株明显较小。依据采自贵州梵净山的标本,提供了详细的形态描述和线条图。     

Crittogame brasiliane,a review of Giuseppe Raddi bryophyte collections in the state of Rio de Janeiro

Abstract

The bryophytes collected by Giuseppe Raddi during his travels in Brazil (1817–1818) housed at Pisa (PI) and Firenze (FI) herbaria were critically studied. The present study provides information about types, nomenclature, taxonomy and geographical distribution for 51 species. One comb. nov. (Taxilejeunea serpyllifolioides (Raddi) Costa (basionym: Jungermannia serpillifolioides Raddi 1822) and two new synonyms (Jungermannia serpillifolioides Raddi 1822 = Taxilejeunea isocalycina (Nees) Steph. 1914; Anthoceros brasiliensis Raddi 1818 = Dendroceros crispus (Sw.) Nees 1846) are made.     

Studien über dieJungermannioideae (Hepaticae) 8.Jungermannia subg.Plectocolea und subg.Solenostoma in Australien,Neuseeland und Ozeanien

Die achte Studie über die SubfamilieJungermannioideae (Jungermanniaceae, Hepaticae) ist dén Arten der GattungJungermannia L. emend.Dum., die in Australien, Neuseeland und Ozeanien vorkommen, gewidmet. Die meist endemischen Arten, die auf Hawaii vorkommen, werden als eine selbständige (neunte) Studie bearbeitet. Im genannten Gebiet wurden 13 Arten der GattungJungermannia L. emend.Dum. festgestellt.J. (P.) hasskarliana (Nees) Steph.,J. (P.) obliquifolia (Schiffn.) Váňa,J. (P.) tetragona Lindenb.,J. (P.) hirticalyx Steph.,J. (P.) minutiverrucosa Amak.,J. (P.) wattsiana Steph.,J. (S.) ariadne Tayl.,J. (S.) totipapillosa Hodgs.,J. (S.) inundata Hook. fil. etTayl. emend.Mitt. undJ. (S.) orbiculata (Col.) Grolle sind eingehend taxonomisch bearbeitet,J. (P.) micrantha (Mitt.) Steph. wird in die nächste Studie eingereiht.J. (P.) boninensis (Horik.) Inoue wurde vonInoue (Inoue etIwatsuki 1969) eingehend beschrieben und abgebildet,Haplozia comptonii Pears. wurde schon im 2. Teil diskutiert. Die anderen aus dem Gebiet bekannten Arten sind mit den obenerwähnten Arten synonymisiert oder zu anderen Gattungen gestellt.