Characterization of the Light Environment in Canopies Using 3D Digitising and Image Processing

A method to measure light interception by vegetation canopiesis presented which uses a 3D digitiser and image processingsoftware. The 3D digitiser allows for simultaneous acquisitionof the spatial co-ordinates of leaf locations and orientations.Software for image synthesis is used to make virtual photographsof the real canopy. Information on light interception is derivedfrom the virtual images by using simple features of image analysissoftware. The method is applied to cotton, grapevine and youngmango plants. Calculations are made of light interception atthe canopy level, light partitioning between plant organs, verticalprofiles of light interception, fisheye photographs and leafirradiance distribution.Copyright 1998 Annals of Botany Company 3D digitising, image analysis, light interception,Gossypium hirsutumL.Vitis viniferaL.,Mangifera indicaL., cotton, grapevine, mango, canopy.     

行距配置对‘兰考矮早八’小麦后期群体冠层结构及其微环境的影响

 在大田试验条件下,研究了在同一密度（375×104株&;#8226;hm^－2）下,行距配置对大穗型冬小麦(Triticum aestivum)品种‘兰考矮早八’后期群体 冠层结构、冠层微环境及产量构成的影响。结果表明,冠层叶面积指数随着行距的增大而减小;而冠层开度随着行距的增大而增加。行距配置 还可改变小麦冠层微环境,冠层不同层次的光截获及消光系数均随着行距的增大而减小;随着行距增加,冠层不同层次的温度升高,而湿度下 降; 随着行距缩小,冠层不同层次的CO₂分布更趋均匀,有利于群体光合作用。通过缩小行距,能够使植株分布均匀,竞争减弱,使产量构成 因素实现最佳配置,从而获得较高的产量。从该研究看,15 cm行距的产量最高,可作为该类品种的较佳行距配置。     

Plant light interception can be explained via computed tomography scanning: demonstration with pyramidal cedar (Thuja occidentalis, Fastigiata)

BACKGROUND AND AIMS: Light interception by the leaf canopy is a key aspect of plant photosynthesis, which helps mitigate the greenhouse effect via atmospheric CO(2) recycling. The relationship between plant light interception and leaf area was traditionally modelled with the Beer-Lambert law, until the spatial distribution of leaves was incorporated through the fractal dimension of leafless plant structure photographed from the side allowing maximum appearance of branches and petioles. However, photographs of leafless plants are two-dimensional projections of three-dimensional structures, and sampled plants were cut at the stem base before leaf blades were detached manually, so canopy development could not be followed for individual plants. Therefore, a new measurement and modelling approach were developed to explain plant light interception more completely and precisely, based on appropriate processing of computed tomography (CT) scanning data collected for developing canopies. METHODS: Three-dimensional images of canopies were constructed from CT scanning data. Leaf volumes (LV) were evaluated from complete canopy images, and fractal dimensions (FD) were estimated from skeletonized leafless images. The experimental plant species is pyramidal cedar (Thuja occidentalis, Fastigiata). KEY RESULTS: The three-dimensional version of the Beer-Lambert law based on FD alone provided a much better explanation of plant light interception (R(2) = 0.858) than those using the product LV*FD (0.589) or LV alone (0.548). While values of all three regressors were found to increase over time, FD in the Beer-Lambert law followed the increase in light interception the most closely. The delayed increase of LV reflected the appearance of new leaves only after branches had lengthened and ramified. CONCLUSIONS: The very strong correlation obtained with FD demonstrates that CT scanning data contain fundamental information about the canopy architecture geometry. The model can be used to identify crops and plantation trees with improved light interception and productivity.     

间伐改形对陇东旱塬密闭苹果园树体冠层结构和发育后期叶片质量的影响

为研究间伐改形对红富士成龄乔化密植果园树体冠层特征、生育后期叶片生理特性、养分积累分配规律、光合生产力和土壤水分时空分布动态的影响,以18年生‘红富士’苹果密闭园为试材,对苹果树单株结构参数、枝量、枝类组成、覆盖率、叶片光合速率等参数进行测定.结果表明: 密闭果园叶片光合作用受非气孔因素限制,导致PSII最大光化学效率、PSII光合潜能、光合性能指数下降了1.2%、11.5%、13.9%.间伐改形后,叶面积指数、树冠覆盖率有所降低,使得冠层直射光透过系数增加了79.0%,树形结构有所改善.苹果园总枝数降低到约1100400条·hm^-2,单株枝量增加了5.0%,短枝比例提高至73.0%.由于冠层光照条件的改善,叶面积、比叶质量、百叶重、叶绿素含量有不同程度的提高.叶片光合速率的提高促进了光合产物的积累,淀粉、蛋白质含量为密闭果园的143.5%、107.8%.叶片的发育质量与其所处的光照辐射环境有着密切联系,密闭果园经间伐、改形后,果园群体结构和冠层光照得以改善,促进了叶片生长发育,提高了叶片光合效能,降低了果园土壤水分的无效消耗,是陇东旱塬苹果产区密闭果园适宜的调整、优化方案.     

Simulated insect defoliation on soybean: influence of row width

An ongoing change in soybean production gaining popularity in the United States is a reduction in row spacing. Plant canopy closure is quicker and leaf area index is greater, thus yield is usually higher. Because yield response to insect defoliation is primarily a function of how defoliation causes changes in light interception, the possibility exists that the insect-injury-yield-loss relationship might differ among row widths. Soybean was grown in four states using similar methodologies. Insect defoliation was simulated by picking leaflets based on an insect defoliation model. Plant growth measurements were taken immediately following the end of defoliation. Numerous independent variables were measured or calculated, including percentage light interception, leaf area index, percentage defoliation, and leaf area per plot. Analyses of covariance were conducted on the resultant data to determine whether insect-injury-yield-loss relationships interact with row width. A significant interaction would indicate that the impact of the variables on yield was dependent on the row width, whereas a nonsignificant interaction would suggest that the relationship between the variables and yield is similar at all row widths. Few significant interactions were obtained, indicating that the impact of the variables on yield is similar across row widths. Because of the lack of significant interactions, the insect-injury-yield-loss relationships previously developed should be usable across varying row widths. Thus, treatment decisions based on light interception and leaf area indices, both considered more appropriated measures of insect injury, should be applicable for all row spacings.     

Harnessing Genetic Variation in Leaf Angle to Increase Productivity of Sorghum bicolor

The efficiency with which a plant intercepts solar radiation is determined primarily by its architecture. Understanding the genetic regulation of plant architecture and how changes in architecture affect performance can be used to improve plant productivity. Leaf inclination angle, the angle at which a leaf emerges with respect to the stem, is a feature of plant architecture that influences how a plant canopy intercepts solar radiation. Here we identify extensive genetic variation for leaf inclination angle in the crop plant Sorghum bicolor, a C4 grass species used for the production of grain, forage, and bioenergy. Multiple genetic loci that regulate leaf inclination angle were identified in recombinant inbred line populations of grain and bioenergy sorghum. Alleles of sorghum dwarf-3, a gene encoding a P-glycoprotein involved in polar auxin transport, are shown to change leaf inclination angle by up to 34° (0.59 rad). The impact of heritable variation in leaf inclination angle on light interception in sorghum canopies was assessed using functional-structural plant models and field experiments. Smaller leaf inclination angles caused solar radiation to penetrate deeper into the canopy, and the resulting redistribution of light is predicted to increase the biomass yield potential of bioenergy sorghum by at least 3%. These results show that sorghum leaf angle is a heritable trait regulated by multiple loci and that genetic variation in leaf angle can be used to modify plant architecture to improve sorghum crop performance.     

叶幕PAR光能截留和分配对葡萄群体光合同化物源库关系的调控

 田间自然条件下在葡萄园群体水平上多年的研究证明：叶幕PAR光能截留率与葡萄群体净光合速率、叶幕和单叶PAR光能截留率与果实干物质总量占地上部生物量干重的百分比、与果实总糖产量占果实本身干物质总量的百分比、以及与果皮色素产量占果实本身干物质总量的千分比之间呈现显著或极显著的线性正相关关系。说明利用叶幕结构变异调节叶幕PAR光能截留和分配,可以对光合同化物源库关系和果实中物质代谢方向进行有效的调控。PAR光能截留率较高和分配合理的叶幕,不但通过较高的群体光合速率为产量和品质形成提供了丰富的同化物“源”,而且通过调节器官间“库”关系使同化物以较高的比例流向果实,同时使果实中物质代谢过程有利于合成构成品质的要素。     

苹果园光能截获率的数学模型

应用气象学原理推导了不同纬度、不同栽植行向、不同树形的苹果园光能截获率的数学模型,给出在保证树冠基部外围日照时间大于25％总日照时间的条件下,生长季中充分截获光能的最佳树形、行距、冠高、冠径的优化组合方案。实例分析表明,计算结果与专家经验基本一致。本研究为果园合理栽植,充分利用光能,合理整形修剪提供理论依据与参考方案。     

秦巴山地植被冠层降雨截留时空分异特征及驱动因素

冠层截留研究对于了解区域水资源分配和评估生态水文功能至关重要,山地复杂多样的环境使其存在较大的不确定性,遥感的发展为揭示山地系统冠层截留的特征提供了机遇。以秦巴山地为研究区,基于降雨数据和叶面积指数遥感数据,耦合植被冠层降雨截留模型,定量模拟和分析秦巴山地2003-2020年植被冠层降雨截留能力及其时空变化特征,并验证其精确性;采用地理探测器、相关分析和约束线法探究冠层截留的驱动因素。结果表明:(1) 与PML_V2数据集和实测数据相比,3.5以下的均方根误差和0.75以上的有效系数证实了A.P.J.DE ROO模型模拟的可靠性。(2) 近18年截留量和截留率整体呈上升趋势,截留率在2015年发生逆转,由增(0.08%/a)向减(-0.15%/a)转变。(3) 秦巴山地冠层截留总体上呈西部高山区和东北部边缘低,秦岭和大巴山区高的空间格局,其随海拔上升呈现"上升-稳定-下降"的分布特征;空间变化以上升趋势为主,显著下降的区域主要分布在汉江河谷的中心;低海拔区域变化差异较大,中海拔区域以显著增加为主,高海拔区域无显著变化。(4) 叶面积指数和降雨量是影响冠层截留的主要因子,约束关系分别为正线型和正凸型;阔叶林截留率与小降雨事件的相关性高,针叶林、灌丛截留率与强降雨事件相关性较强,气候因子对冠层截留的影响在类别和解释程度上存在空间差异。研究可为区域尺度冠层截留的估测提供思路,且有助于评估气候变化背景下生态系统对水循环的影响。     

Changes in the vertical distribution of leaf area enhanced light interception efficiency in maize over generations of selection

Breeders select for yield, thereby indirectly selecting for traits that contribute to it. We tested if breeding has affected a range of traits involved in plant architecture and light interception, via the analysis of a panel of 60 maize hybrids released from 1950 to 2015. This was based on novel traits calculated from reconstructions derived from a phenotyping platform. The contribution of these traits to light interception was assessed in virtual field canopies composed of 3D plant reconstructions, with a model tested in a real field. Two categories of traits had different contributions to genetic progress. (a) The vertical distribution of leaf area had a high heritability and showed a marked trend over generations of selection. Leaf area tended to be located at lower positions in the canopy, thereby improving light penetration and distribution in the canopy. This potentially increased the carbon availability to ears, via the amount of light absorbed by the intermediate canopy layer. (b) Neither the horizontal distribution of leaves in the relation to plant rows nor the response of light interception to plant density showed appreciable trends with generations. Hence, among many architectural traits, the vertical distribution of leaf area was the main indirect target of selection.     

Growth of Lettuce, Onion, and Red Beet. 1. Growth Analysis, Light Interception, and Radiation Use Efficiency

A field experiment was carried out to analyse the growth oflettuce, onion and red beet in terms of: (a) canopy architecture,radiation interception and absorption; (b) efficiency of conversionof absorbed radiation into biomass; and (c) dry matter partitioning.Growth analysis, total solar radiation interception, PAR interceptionand absorption by the crop canopy, ground cover, maintenancerespiration of onion bulbs and red beet storage roots were measured.Models for different leaf angle distribution and ground coverwere used to simulate light transmission by the crop canopy. The three crops are shown to have contrasting growth patternsfrom both a morphological and a physiological point of view.Lettuce showed very high light interception and growth afterthe early growth stages but, throughout the growth cycle, thisleafy crop showed the lowest radiation use efficiency due tothe respirational cost of the high leaf area. Onion showed alower early relative growth rate than lettuce and red beet.This was due partly to the low light interception per unit leafarea in the later stages of growth and partly to the low initialradiation use efficiency compared with the other two crops.On the other hand, thanks to more uniform distribution of theradiation inside the canopy, to the earlier termination of leafdevelopment and to the very low level of bulb respiration, onionshowed high radiation use efficiency and was able to producea large amount of dry matter. Red beet leaf posture and canopystructure resulted in high light interception and absorption.Its radiation use efficiency was lower than that of onion, partlyperhaps because of the more adverse distribution of the interceptedradiation fluxes within the canopy and partly because of thehigh respiration cost of a continuous dry-matter allocationto the leaves. However, this crop can accumulate a very largeamount of dry matter as leaf blade development and storage rootgrowth can both continue almost indefinitely, providing continuouslyavailable sinks. Ground cover gave a good estimate of the PAR interception onlyat low values of light interception but, in general, it underestimatedPAR interception in all three crops. Ratios between attenuationcoefficients established by considering PAR or total solar radiationand LAI or ground cover were calculated. Lettuce,Lactuca sativa L. var.crispa ; onion,Allium cepa L.; red beet; Beta vulgaris L. var.conditiva ; growth analysis; light interception and absorption; canopy architecture; ground cover; radiation use efficiency; maintenance respiration rate; dry matter distribution     

Effects of spacing interval of wide bed planting on canopy characteristics and yield in winter wheat

2010-2012连续两年在大田试验条件下以多穗型品种‘百农矮抗58’为供试材料, 研究了宽幅播种(播幅8 cm)种植方式下不同带间距7 cm (KF7)、12 cm (KF12)和17 cm (KF17)对冬小麦(Triticum aestivum)冠层特征及产量的影响。结果表明: 与常规条播比较, 宽幅播种群体叶面积指数、平均叶倾角、光截获量、相对湿度、成穗数、生物量和产量较高, 而冠层开度和温度较低; 2010-2011年和2011-2012年宽幅播种成穗数和产量显著高于常规条播, 成穗数分别增加4.8%-16.4%和8.9%-21.0%, 产量分别提高2.96%-15.94%和4.09%-14.23%。宽幅播种下随带间距增大, 叶面积指数、平均叶倾角、光截获量、湿度及成穗数降低, 而冠层开度和温度升高; 穗粒数、千粒重、产量、生物量和收获指数以KF12最高, KF7最低。综合分析, 宽幅播种下12 cm带间距处理的小麦冠层结构合理, 微环境适宜, 产量最高, 可作为该种植方式的适宜带间距。     

Exploring the spatial distribution of light interception and photosynthesis of canopies by means of a functional-structural plant model

Background and Aims

At present most process-based models and the majority of three-dimensional models include simplifications of plant architecture that can compromise the accuracy of light interception simulations and, accordingly, canopy photosynthesis. The aim of this paper is to analyse canopy heterogeneity of an explicitly described tomato canopy in relation to temporal dynamics of horizontal and vertical light distribution and photosynthesis under direct- and diffuse-light conditions.

Methods

Detailed measurements of canopy architecture, light interception and leaf photosynthesis were carried out on a tomato crop. These data were used for the development and calibration of a functional–structural tomato model. The model consisted of an architectural static virtual plant coupled with a nested radiosity model for light calculations and a leaf photosynthesis module. Different scenarios of horizontal and vertical distribution of light interception, incident light and photosynthesis were investigated under diffuse and direct light conditions.

Key Results

Simulated light interception showed a good correspondence to the measured values. Explicitly described leaf angles resulted in higher light interception in the middle of the plant canopy compared with fixed and ellipsoidal leaf-angle distribution models, although the total light interception remained the same. The fraction of light intercepted at a north–south orientation of rows differed from east–west orientation by 10 % on winter and 23 % on summer days. The horizontal distribution of photosynthesis differed significantly between the top, middle and lower canopy layer. Taking into account the vertical variation of leaf photosynthetic parameters in the canopy, led to approx. 8 % increase on simulated canopy photosynthesis.

Conclusions

Leaf angles of heterogeneous canopies should be explicitly described as they have a big impact both on light distribution and photosynthesis. Especially, the vertical variation of photosynthesis in canopy is such that the experimental approach of photosynthesis measurements for model parameterization should be revised.     

草坪型结缕草冠层截留性能试验研究

草坪植物冠层对雨水或灌溉水的截留量大小对于草坪水分利用效率具有重要影响.针对草坪型结缕草,用吸水法分别试验研究了单株和群落草坪草的冠层截留特性.结果表明:结缕草的叶片数量、叶面积对单株结缕草截留量影响显著.随着叶片数量的增多,单株结缕草截留量增大,5叶枝条的结缕草截留量高达0.16g,占其植株浸水前自身重的81.22%,而3叶枝条的结缕草截留量仅为5叶枝条结缕草的1/2.单株结缕草截留量与叶面积呈线性正相关关系.对于结缕草草坪,修剪留茬高度越高,截留量就越多.建植3a的草坪在修剪高度14cm时冠层截留可达1.05mm,修剪6cm时截留量为0.48mm.在修剪留茬高度相同的情况下,不同生长年限间草坪草截留量、截留率差异极显著(P＜0.01),建植3a的草坪比建植1a的草坪能拦截更多的水分,在修剪高度6cm时,3a生草坪草截留量为0.48mm,而1年生草坪草截留量为0.38mm,并且随着修剪高度的增加,不同年限间截留量的差异越明显,差值最大可达0.44mm.     

An Analysis of Spatial Variation in the Nitrogen Content of Leaves from Different Horizons Within a Canopy

An analysis of spatial variation in the specific nitrogen contentof leaves from different positions within a canopy is developed.It is used to examine data obtained for contrasting crop species;the legume crop guar (Cyamopsis tetragonoloba), and crops ofa forage sorghum. The analysis distinguishes two componentsof the leaf nitrogen content: one associated with the metabolicapparatus within the leaf tissues and the other with structuralelements of the leaf. The analysis allows the spatial variationin specific leaf nitrogen content to be quantitatively described. Leaf nitrogen content, crop growth, light interception     

黄土高原农牧交错带人工乔灌木林冠截留

在黄土高原农牧交错带的六道沟流域,通过2年观测,研究了主要人工造林树种油松、刺槐、柠条和沙柳的冠层截留特征。结果表明：4个树种林冠截留量和林内降雨量均与大气降水量呈正相关,其中林冠截留量与大气降水量呈幂函数关系;油松林平均截留率为16.7％,刺槐林为11.6％,沙柳林为12.8％,柠条林为11.2％。模拟结果表明,修正的王彦辉模型较原模型能够更好地反映黄土高原半干旱地区典型乔灌木树种冠层截留的大小。     

川西亚高山典型森林生态系统截留水文效应

截留是水文循环的一个重要过程,水文功能是森林生态系统功能的重要方面,林冠和枯落物截留实现对大气降水的二次分配过程.为深入认识生态系统截留的水文效应,采用野外观测和人工降雨模拟试验相结合的方法,研究了2008年和2009年5-10月贡嘎山亚高山峨眉冷杉中龄林、峨眉冷杉成熟林和针阔混交林的冠层枯落物截留能力.结果表明,峨眉冷杉中龄林2008年林冠截留率为20.9％,针阔混交林2008年和2009年林冠截留率分别为23.0％和23.6％,林冠截留率的年际间变化不大,林冠截留主要受到降雨特征影响.3种林型枯落物饱和持水能力分别为5.1、5.1和5.7 mm,显著高于林冠的饱和持水能力,但由于冠层的截留蒸发速率较高,林冠截留蒸发仍是生态系统截留蒸发的主要组成部分.     

苏南丘陵区毛竹林冠截留降雨分布格局

降雨穿过林冠层时,由于林冠的拦截作用,改变了降雨分布格局。林冠截留是一个复杂的水文过程,受降水特征及林分特性的影响较大。本文以北亚热带苏南丘陵地区人工毛竹林(Phyllostachys edulis)为研究对象,利用2007年度各场次降雨观测数据,分析了降雨量和降雨强度与林冠截留降雨的关系,研究了林冠截留过程的特点。结果表明：（1） 研究期间共观测到102次降水事件,降水总量为1110.8mm,单次最大降雨量为110.0㎜,最小为0.55㎜,事件平均降水量为10.89mm,且绝大部分降雨为低雨强、中雨量级的降雨事件。（2） 研究期间林冠截留总量为171.72mm,占同期降雨总量的15.46%。单次林冠截留量变幅为0.21—4.55mm,截留率变幅为1.3%—100%,且随林外降雨量的递增,林冠截留率呈现递减的变化趋势,二者的关系用幂函数（I0=117.34P-0.9106）拟合效果较好。（3）在林外次降雨量小于5㎜的条件下,事件降雨量占年降雨总量的5.0%,相应的降雨事件频率为9.8%,此时林冠截留量随降雨量的增大而增加,其变幅为0.55—1.9㎜,截留量与降雨量的关系用对数函数（I=0.4931Ln(P) 0.9493）进行拟合效果较好;在单场降雨量大于5㎜时,林冠截留量随各场次降雨量的增加,其变动幅度和频率均大大增强,变化范围在0.21—4.55㎜之间,二者相关性较差。（4）降雨在7—10mm雨量级范围内时,林外降雨量和林冠截留量分别为245.14mm和47.9㎜,占其全年总量的比例均为最大,分别为22.07%和27.9%;各雨量级林冠平均截留量与平均降雨量的关系表现为对数函数关系（R2=0.7287）,而截留率与平均降雨量的关系表现为极显著的幂函数关系（R2=0.9817）,且林冠对降雨的截留作用在雨量级较小时,表现十分显著。（5）全年单场降雨强度小于0.06mm/min时,降雨事件频数为73.53%,降雨量占其总量的57.93%,林冠截留量占其总量的89.34%,降雨事件平均截留率（23.84%）远高于雨强大于0.06mm/min时的降雨事件平均截留率（3.92%）。本研究结果为长江中下游丘陵山区水土保持林和水源涵养林体系建设提供了理论依据。     

The effects of tree characteristics on rainfall interception in urban areas

Trees in urban areas have significant effects on the urban ecosystem. They can be used to improve the water cycle in urban areas by increasing evaporation and reducing runoff through rainfall interception. Street trees placed in planters on impervious areas reduce runoff by intercepting rainfall and by temporarily storing raindrops on leaves. Therefore, understanding tree canopy geometry and the effect of rainfall interception is important in urban hydrology. In this study, we assessed the effect of tree canopy morphology on rainfall interception using four major street tree species, Sophora japonica L., Ginkgo biloba L., Zelkova serrata (Thunb.) Makino, and Aesculus turbinata Blume, in Seoul, South Korea. We measured throughfall for each tree and also derived three-dimensional data of tree canopy morphology with a terrestrial laser scanner. Tree height, canopy crown width, leaf area index (LAI), leaf area density, mean leaf area, and mean leaf angle were used to determine canopy morphology. The interception rate was mostly affected by the LAI; a higher LAI tended to result in a higher interception rate. Leaf area affected the rainfall interception rate when trees had similar LAIs. These findings on individual tree canopy rainfall interception can help us to understand the importance of rainfall interception in hydrology and for ecological restoration when planning urban green spaces.

     

模拟降雨条件下玉米植株对降雨再分配过程的影响

为系统测定玉米(Zea mays)不同生长阶段的穿透雨、茎秆流和冠层截留,采用室内模拟降雨法测定了不同降雨强度、不同叶面积指数玉米冠下穿透雨和茎秆流,采用喷雾法测定了玉米不同生长阶段的冠层截留。对其进行了量化分析,并探讨了三者与玉米叶面积指数和降雨强度的关系,阐明了玉米冠下穿透雨的空间分布特征。结果表明:玉米冠下穿透雨量占冠上总降雨量比例为30.97%—94.02%,平均为63.92%;茎秆流量占降雨量比例的变化范围为5.68%—75.70%,平均为35.28%;冠层截留量在其全生育期内变化范围为0.02—0.43 mm,平均为0.16 mm,所占总降雨量比例最大仅为1%。随玉米生长,穿透雨量逐渐降低,茎秆流量和冠层截留量逐渐增加。穿透雨与茎秆流呈现此消彼长的关系,其中穿透雨率平均由93.55%降至36.23%;茎秆流率平均由5.98%增加至70.42%。降雨强度与穿透雨量和茎秆流量呈正相关关系,但是二者占总降雨量的比例与降雨强度关系不显著(P0.05)。随着玉米生长,穿透雨冠下空间分布由均匀逐渐趋向于不均匀,使降雨经过冠层后趋于向行中汇集,但在玉米生长后期,集中于行中的穿透雨量也因叶片衰败而随之降低。揭示了玉米对降雨的再分配作用特征,可为农田水分有效利用、农田生态水文过程机理和坡耕地土壤侵蚀防治提供理论依据。