导语

<正>森林生态系统保护与恢复是新时期国家战略——生态文明建设的主体。提升森林生态系统功能、做强林业产业、精准提升森林质量不仅是我国《林业发展"十三五"规划》的重大战略任务,更是维持国家生态安全、促进生态文明建设的核心目标。开展森林生态与经营研究,是完成上述重大任务和核心目标的基础与保障。森林生态与经营(Forest Ecology and Management)是研究森林生物之间及其与环境之间相互关系的规律,并应用于营林实践以获得林木或其他林产品以及森林生态效     

泾河流域县域尺度生态系统服务相互关系及影响因子

以泾河流域31个县粮食供给、肉类供给、薪柴供给、水源涵养和土壤保持5项生态系统服务为研究对象,分析了县域尺度5项生态系统服务的相互关系、相互关系模式和总生态系统服务指数的空间差异,以及影响空间差异的自然环境和人为因子.结果表明:泾河流域县域尺度5项生态系统服务之间相互关系的差异较大,粮食供给与肉类供给呈极显著正相关,与土壤保持呈极显著负相关,水源涵养与薪柴供给和土壤保持呈显著正相关.生态系统服务相互关系模式中,调节服务主导模式、供给与调节平衡模式和粮食供给主导模式的县分别为24、3和4个;不同模式的总生态系统服务指数差距较大,最大值(泾源县)与最小值(盐池县)差距5.1倍.自然环境因子中,总生态系统服务指数与降水和土壤全氮呈极显著正相关,与日照时数呈显著负相关;排除自然环境因子的差异后,耕地增加对总生态系统服务指数有负面影响,乔木林地影响甚微,灌木林地和草地有较大的正面影响.     

“一带一路”地区生态系统服务关系的时空分异与区域响应

"一带一路"沿线多数发展中国家地处生态脆弱区,土地资源空间配置亟待优化。研究生态系统服务间的相互关系可以为土地利用决策提供定量化支持,但该关系在不同地域的表现形式有较大的差异性。基于InVEST模型,评估"一带一路"区域2000、2005、2010和2015年土壤保持、产水量、碳固定和生境质量4种生态系统服务时空变化,通过相关性分析和局部空间自相关分析等方法识别生态系统服务相互关系,以及其对不同地域特征变量的响应。就区域整体而言,土壤保持、产水量、碳固定和生境质量呈现中亚和西亚为一致低值区,而高值区分布不一致的空间格局;生态系统服务之间均呈显著协同关系,但冷热点格局的区域异质性显著。在生态区单元中,泰加林、温带草原和灌木的多种生态系统服务间为权衡关系;在国家单元中,生态系统服务间协同与权衡关系并存。产水量与碳固定间的协同关系对人口分布有良好的响应,而土壤保持与碳固定对人口分布的响应不明显。定量化评估生态系统服务及权衡关系,为区域土地资源优化配置提供依据,研究结果也为"一带一路"地区沿线国家间生态保护合作提供空间指引。     

我国农田生态系统重金属的循环与调控

本文系统分析了我国农田土壤重金属的含量状况,探讨了农田生态系统中重金属的输入、输出及其平衡.农田生态系统中重金属存在一定的累积效应,但整体上超标风险较小,而部分作物、特别是蔬菜可能存在一定的超标风险.在采矿区和冶炼厂周边地区,以及废弃物利用强度较大的农区,土壤和作物中重金属超标的风险较大,是普通农区的十倍甚至数十倍以上.针对我国农田重金属污染现状,提出了调控农田生态系统重金属循环的有效措施,并对相关领域今后的研究进行了展望.     

基于PSR的黄河河口区生态系统健康评价

根据压力-状态-响应(PSR)框架模型,从广义上定义河口区生态系统,将河口及毗邻的陆域、海域生态系统作为一个整体,从压力指标、状态指标、响应指标3个方面构建了黄河河口区生态系统健康评价的指标体系,以研究区1991年数据和相关国家标准为基准,2013年代表现况,利用综合指数法(CEI)评价了黄河河口区的生态系统健康状况。结果显示:黄河河口区生态系统健康评价的响应指数最高(0.9055),压力指数居中(0.8288),状态指数最低(0.6458),综合指数为0.7427。总体来看,与1991年相比,目前黄河河口区生态系统仍处于"健康"状态,但健康状况明显下降,其中状态指数下降最为严重。从区域轻度开发到人类活动强烈干扰阶段,黄河河口区存在过度捕捞、湿地不合理开发、浅海养殖过度及污染物排放等一系列影响生态系统健康的问题,应进行区域的生态恢复和科学管理。     

污染生态学

污染生态学是研究生物与污染环境之间相互关系和基本规律的科学,是环境科学的一个重要分支。目前正在进行生态系统本底值的普查和污染情况的监测,并从宏观和微观两方面深入研究污染物在生态系统各个层次上的迁移、转化、积累规律,探索生态系统的净化能力,确定各项有关参数,为评价环境质量和保护环境提供科学依据。     

黄河三角洲湿地生态系统的形成及其保护

通过对黄河三角洲新生湿地的野外调查,利用TM陆地卫星影像图的判读,结合黄河尾闾变动的历史资料分析,结果表明,黄河三角洲新生湿地是多泥沙的黄河淤积和海岸蚀退所形成.受区域气候、地貌、地质沉积、土壤、植被的相互作用和互相影响,形成4.5×10⁵hm²的湿地,其中有6.84×10⁴hm²的人工湿地.湿地生态类型随着黄河尾闾摆动和海退陆进程度而变化,自海洋向陆地依次为潮下带水生生态系统潮间带湿地生态系统潮上带盐生生态系统芦苇獐茅湿地生态系统草甸湿地生态系统陆上农田生态系统.该系统资源丰富,共有野生动物1524种,鸟类300种,鱼类1040种.石油开发和近年来黄河断流对湿地产生一定影响,应在开发利用中加强管理和保护.     

乌兰布和沙漠生态系统服务价值时空动态

以乌兰布和沙漠为研究区,基于1990—2018年4期土地利用类型数据,以"当量因子法"为基础,运用网格分析、热点分析、权衡协同分析等方法,探究乌兰布和沙漠近30年生态系统服务价值的时空动态特征。结果表明:(1)生态系统服务价值总量以2000年为拐点先下降后逐期上升,由1990年17.03亿元上升至2018年25.94亿元,其中调节服务价值贡献量最大,4期平均占71.34%,供给服务增幅最大,近30年间增加85.04%;(2)生态系统服务价值呈东部和东北部高、中部和西南部低的空间分布特征,1990—2010年空间格局基本不变,但2018年高值区有所扩展;(3)1990—2018年间生态系统服务冷点区面积占比由2.85%增至4.23%,热点区面积占比由0.22%增至10.45%,热点区分布在黄河沿岸及部分生态恢复区,但东南部冷点区逐渐增大,生态治理时应着重关注;(4)1990—2018年生态系统服务间以协同关系为主,权衡关系存在于初期供给服务与其他类型服务之间。     

竺山湾湖泊缓冲带湿地生态系统EWE模型构建与分析

湖泊缓冲带在湖泊流域空间布局中具有特殊地位,缓冲带内的湿地对于保障流域生态健康和湖泊水环境质量具有十分重要的意义.本研究以太湖竺山湾湖泊缓冲带内的竺山湖湿地生态系统为研究对象,将生物组分划分为16个功能组,构建了生态通道(EWE)模型,并分析了生态系统的特征、状态以及功能组之间的相互关系.结果表明: 竺山湖湿地生态系统的有效营养级范围在1~3.72,营养流动主要发生在前4个营养级,开始于沉水植物和有机碎屑的食物链较多.湿地生态系统的总的能量转换效率为5.1%,并未达到“1/10定律”,说明当前的能量转换效率较低.物质流量在生态系统中的平均传输效率为4.3%.系统的总生产量为2496.66 t·km^-2·a^-1,总流量为10145.2 t·km^-2·a^-1.生态系统的多种特征参数表明当前生态系统处于幼态化阶段.     

浅谈微生物与人类生态系统的相互影响

概括介绍了微生物、人类生态系统的基本知识以及二者之间的相互影响。提出：人类首先应该处理好微生物和人类生态系统之间的相互关系,使其协调发展,进而使生物多样性得以维持和发展。在这个基础上再尽量减少或消除微生物对人类生态系统的消极影响,充分发挥微生物的积极作用,达到利用微生物为人类服务的最终目的。     

华北平原种养一体规模化农场氮素流动特征及利用效率——以河北津龙循环农业园区为例

种养一体规模化、集约化是华北平原农业发展的必然趋势,而氮素是连接种植养殖的主要养分资源,以河北津龙循环农业园区为例,采用文献资料、实地调查方法分析农场水平氮素流动特征及利用率,并通过情景分析方法提出农场氮素管理措施,为实现农场水平氮养分资源高效利用、提高农场生产系统生产力和改善华北平原循环农业模式提供技术支撑和科学依据.结果表明: 在农场水平下,化肥和有机肥输入氮量674.6 kg·hm^-2·a^-1,占总输入氮量的88.3%,氮利用率为41.5%,种植系统氮盈余量190.7 kg·hm^-2·a^-1,施氮量过多是造成种植系统氮利用率低和氮素盈余量高的主要原因.养殖系统中外购饲料提供氮量占饲料总输入氮量的83.2%,粪尿排氮量为776.6 t·a^-1,而还田比例仅为36.3%,氮利用率19.7%.农场水平氮总利用率为40.7%.情景分析表明,农田减少化肥施氮量50%(情景1)、增加来自农场内部玉米籽粒产量(情景2)措施,可分别使种植系统氮利用率提高34.6%和15.6%,同时农场水平氮总利用率分别提高18.7%和9.8%;另外,优化养殖系统饲料结构(情景3),可使氮总利用率提高19.1%.因此,减少化肥氮施用量、调整作物种植结构、优化饲料结构等,是提高农场氮生产力和实现环境友好双赢效果的措施和途径.     

不同农业景观背景下传粉昆虫群落的分布差异

农业景观中的自然、半自然生境等非农生境可为传粉昆虫提供丰富的食物来源、栖息地、繁殖地、避难所等,对维持生物多样性的稳定起着不可替代的作用。以巩义典型的山地-丘陵-河川混杂的复杂景观和民权的平原农业简单景观为研究区,分析不同景观背景下传粉昆虫群落的物种组成及其在不同生境中的分布特征。结果表明:(1)巩义研究区内累计捕获传粉昆虫18582头,民权研究区内累计捕获传粉昆虫18518头,优势传粉昆虫功能群为双翅目、膜翅目、鞘翅目和鳞翅目等;(2)景观复杂度更高的巩义研究区的传粉昆虫多样性、丰富度以及均匀度均显著高于民权研究区;存在大面积农田斑块的平原景观中则有更多的优势传粉昆虫个体;(3)农田斑块中具有更高的物种丰富度,但林地物种的多样性和均匀度则相对稍高些。农田斑块在作物花期能有效的提高传粉者种群密度,但林地等自然、半自然生境对于维持传粉者多样性和食物缺乏期种群的稳定具有重要作用。因此在未来为确保农业景观中传粉者的多样性与传粉服务的稳定,一方面要关注自然、半自然生境的作用,同时也要考虑不同景观背景下异质性特征对不同生物类群的影响差异。     

基于不同预测变量的天然椴树可加性地上生物量模型构建

森林生物量是森林生态系统的最基本数量特征,生物量数据是研究许多林业问题和生态问题的基础,因此,准确测定生物量对于计算碳储量以及研究气候变化、森林健康、森林生产力、养分循环等十分重要.目前,测算森林生物量常用的方法为生物量模型估算法.本研究基于小兴安岭地区和张广才岭地区97株实测生物量数据,建立了3个天然椴树立木可加性生物量模型系统(基于胸径的一元可加性生物量模型系统、基于胸径和树高的二元可加性生物量模型系统、基于最优变量的最优可加性生物量模型系统),采用非线性似乎不相关回归法进行参数估计,用加权方法解决模型的异方差问题,并采用“刀切法”进行模型检验.结果表明: 3种可加性生物量模型系统均能较好地对椴树各部分生物量进行拟合和预测(调整后确定系数R_a²>0.84,平均预测误差百分比MPE<8.5%,平均绝对误差MAE<16.3 kg,平均百分标准误差MPSE<28.5%),其中,树干和地上生物量的拟合效果优于树叶、树枝和树冠;在引入树高和树冠因子后,提高了模型的拟合效果和预测能力(R_a²提高0.01～0.04,MAE降低0.01～4.55 kg),缩小了预测值置信区间的范围,树干、树叶和地上生物量提高较多,树枝和树冠提高较少.总体来看,最优生物量模型系统效果最好,其次为二元生物量模型系统,再次是一元生物量模型系统,添加树高和树冠因子进行生物量模型的构建十分必要.     

Livestock reduces juvenile tree growth of alien invasive species with a minimal effect on natives: a field experiment using exclosures

Many alien invasive tree species were originally introduced to their non-native ranges for use in forestry and as urban trees. These alien species were selected for their fast growth and not necessarily for possessing mechanisms which deter browsing. Instead, many tree species native to semiarid areas of the world evolved mechanisms which deter browsing, presumably at the cost of slower growth. In a semiarid rangeland we observed that livestock exclusion greatly promoted the growth of juveniles of several alien species but not of native species, and we hypothesized that this increase in growth of aliens was due to livestock preference for alien and not native trees. With the objective of quantifying our observations and understanding the mechanism underlying the increased growth rates of alien juvenile trees under livestock exclusion, we assessed growth and browsing levels in juveniles of two alien invasive and four abundant native tree species within three parcels where livestock was excluded and three parcels with livestock at 0.20 cattle equivalents.ha^?1. Alien species grew around four-fold faster under livestock exclusion than with livestock and, as predicted, received five times more browsing than natives. Instead, native species did not significantly increase their growth rate with livestock exclusion. The results support our hypothesis and the implications for management would be that stocking paddocks with livestock to browse existing alien juveniles and re-growth of felled adults should be effective in delaying invasions of trees used for forestry without significantly affecting the growth of the most abundant native trees.     

城市化背景下广佛都市圈农林生态系统服务价值

从广佛都市圈1990、1995、2000、2005和2010年5期土地利用矢量数据中提取出耕地、林地、园地和水域4类农林业用地.基于Costanza等生态系统服务价值计算方法,在对谢高地等提出的“中国陆地生态系统单位面积生态服务价值”进行区域修正及功能性和经济性调整的基础上,估算了1990—2010年间广佛都市圈农林生态系统的生态服务价值.结果表明: 1990—2010年间,随着经济的不断发展以及人们生活水平的不断提高,居民对农林生态系统服务的支付意愿和支付能力逐年增强,使得广佛都市圈农林生态系统服务价值总体上呈波动上升趋势,1990、1995、2000、2005和2010年的服务价值分别为413.74×10⁸、612.83×10⁸、582.88×10⁸、773.44×10⁸和698.67×10⁸元,20年间提高了284.93×10⁸元,平均年增幅为2.7%.研究期间,总服务价值增加最多的为水域,达 166.74×10⁸元,其次为林地和园地,分别增加了103.01×10⁸和47.74×10⁸元.耕地的服务价值则降低了32.56×10⁸元,降幅达23.1%.在空间分布上,广佛都市圈各年的农林生态系统服务价值均表现为从东北和西南部向中部减少的趋势.各类型农林生态系统服务价值所占比例在年际间变化不大,总体表现为:涵养水源>废物处理>生物多样性保护>气候调节>土壤形成与保护>气体调节>娱乐休闲与文化>原材料提供>食物生产.     

Phytomass utilization and deposition of feces by ungulates on steppe pastures of eastern Mongolia

The amount of phytomass removed by a complex of livestock (horses, cattle, sheep, and goats) and wild ungulates (Mongolian gazelle Procapra gutturosa Pall.) grazing in plain and mountain pastures of eastern Mongolia has been estimated by taking account of feces deposited by these animals. The results show that at an animal density of up to 30 head/km², the total annual amount of feces reaches 140 kg/ha (dry weight), with the greater part (up to 90 kg/ha) being deposited by horses. The contribution of Mongolian gazelles in some pastures reaches 20–40 kg/ha per year. Decomposition of feces proceeds very slowly, with the annual loss of their weight averaging only 9–12%. This is evidence for gradual accumulation of nondecomposed matter in the soil. The removal of phytomass by the complex of ungulates, calculated from the amount of feces with regard to their annual loss and forage digestibility, varies in different grazing areas from 240 to 400 kg/ha (25–60% of the total aboveground phytomass). The greatest amount of phytomass is utilized by horses, reaching 200 kg/ha (13%), and Mongolian gazelles utilize up to 86 kg/ha per year. In the growing season, ungulates remove no more than 11–16% of the total aboveground phytomass. It is concluded that the impact of total ungulate stock does not impair the productivity of vegetation in the pastures studied.     

Impact of balsam fir flowering on pollen and foliage biochemistry in relation to spruce budworm growth, development and food utilization

The impact of balsam fir (Abies balsamea (L.) Miller) flowering on nutritional and allelochemical quality of pollen, current-year and one-year-old foliage is studied in relation to spruce budworm (Choristoneura fumiferana Clem.) (Lepidoptera: Tortricidae) growth, development and utilization of food and nitrogen. In the laboratory, using fresh food from the field, we simulated conditions of low larval population density, in which there is no current-year foliage depletion during the spruce budworm feeding period. Similarly, we simulated conditions of high larval population density when current-year foliage depletion occurs.Because of the high nutritive value of pollen (high amounts of amino acids and minerals, especially nitrogen; low monoterpene content), insects from flowering trees reached the fifth instar five days earlier than those from non-flowering trees, and had heavier dry- and nitrogen-weights at the beginning of the fifth instar. At budbreak, switching from pollen to current-year foliage negatively affected conversion efficiencies and digestibilities of food and nitrogen (AD; ADN; ECDN; ECI; ECIN). The switch from pollen to new foliage had a detrimental impact on fifth-instar survival and on newly-moulted sixth-instar dry- and nitrogen-weights. Moreover, during the fifth instar, balsam fir flowering reduced the nutritive value of current-year foliage, which in turn, might have contributed to the reduced larval growth. Nevertheless, during the sixth instar, balsam fir flowering affected the biochemistry of current-year foliage in ways that enabled larvae to compensate for their low fifth-instar biological performance; larvae also managed to reach pupal dry weight similar to larvae reared on non-flowering trees. Current-year foliage from flowering trees contained less nitrogen, total soluble sugars and total monoterpenes. Those foliar characteristics enabled larvae to increase food and nitrogen consumption rates (RCR; RNCR), because of lower repellency and/or post-ingestional feedback from monoterpenes.As for current-year foliage, balsam fir flowering reduced nitrogen, total soluble sugar and total monoterpene contents in one-year-old foliage during the sixth-instar feeding period. These characteristics enabled sixth-instar larvae, fed on old foliage from flowering trees, to have high relative food and nitrogen consumption rates (RCR; RNCR). Larvae were also able to reach higher relative growth rates (RGR) and relative nitrogen accumulation rates (RNAR) compared to larvae reared on one-year-old foliage from non-flowering trees. Finally, larvae on flowering trees had pupal dry weight similar to those from non-flowering trees, but reached the adult stage nine days earlier.Regardless the foliage type consumed by spruce budworm larvae during the sixth instar, pollen consumption during early larval stages reduced total development time, and thus exposure time to natural enemies. This phenomenon might increase larval survival. Balsam fir flowering changed the biochemistry of one-year-old and current-year foliages, but did not affect pupal dry weights of larvae reared on flowering trees compared to those reared on non-flowering trees. Thus, at low population density, spruce budworm populations in balsam fir flowering stands might be favoured over those in balsam fir non-flowering stands. In addition, when larvae consumed one-year-old foliage during the entire sixth instar, those on flowering trees are probably favoured over those on non-flowering trees. However, because flowering trees produce less new foliage than non-flowering trees, current-year foliage depletion may occur earlier on flowering trees than on non-flowering trees. Thus, at similar larval population density, larvae on flowering trees might have to feed on one-year-old foliage earlier than those on non-flowering trees. In that case, spruce budworm populations on non-flowering stands would be favoured over those on flowering stands.     

Communities of insect herbivores foraging on saplings versus mature trees of Pourouma bicolor (Cecropiaceae) in Panama

The arthropod fauna of 25 saplings and of three conspecific mature trees of Pourouma bicolor (Cecropiaceae) was surveyed for 12 months in a tropical wet forest in Panama, with particular reference to insect herbivores. A construction crane erected at the study site provided access to tree foliage in the upper canopy. A similar area of foliage (ca. 370 m²) was surveyed from both saplings and trees, but samples obtained from the latter included 3 times as much young foliage as from the former. Arthropods, including herbivores and leaf-chewing insects with a proven ability to feed on the foliage of P. bicolor were 1.6, 2.5 and 2.9 times as abundant on the foliage of trees as on that of saplings. The species richness of herbivores and proven chewers were 1.5 (n=145 species) and 3.5 (n=21) times higher on trees than on saplings, respectively. Many herbivore species preferred or were restricted to one or other of the host stages. Host stage and young foliage area in the samples explained 52% of the explained variance in the spatial distribution of herbivore species. Pseudo-replication in the two sampling universes, the saplings and trees studied, most likely decreased the magnitude of differences apparent between host stages in this forest. The higher availability of food resources, such as young foliage, in the canopy than in the understorey, perhaps combined with other factors such as resource quality and enemy-free space, may generate complex gradients of abundance and species richness of insect herbivores in wet closed tropical forests.     

Light interception in apple trees influenced by canopy architecture manipulation

Improvement of light penetration within tree canopies has been a constant objective of fruit tree architecture manipulation through the setting up of training systems. Recently, centrifugal training, i.e. the removal of fruiting shoots in the tree centre and on the underside of branches, has been proposed to improve fruit size and colour as well as return-bloom as compared to conventional solaxe-trained trees with equivalent crop loads. The present study was conducted to quantify the benefits of centrifugal training on light interception by the fruiting shoots via computer-assisted three-dimensional representations of foliage geometry. Data were collected on six 5-year-old apple trees cv.Galaxy, trained either with solaxe or centrifugal training systems, using an electromagnetic 3D digitiser. The 3D distribution of the foliage in the tree canopy was recreated by combining both the spatial locations of shoots (as measured from 3D digitising) and foliage reconstruction. Light interception efficiency properties of the trees were characterised by silhouette to total area ratio (STAR) values computed from images of the 3D mock-ups. Compared to the solaxe system, centrifugal training significantly improved the STAR of the whole tree by 20%. It also increased both leaf area and STAR of the fruiting shoots by approximately 15%, regardless of their position in the canopy. In this paper, we discuss the role of this enhanced light interception by the canopy in increasing the autonomy of the fruiting shoot, i.e. improved fruit size and colour, and return-bloom.     

The biomass of New England peppermint (Eucalyptus nova-anglica) in relation to insect damage associated with rural dieback

Two adjacent mature trees of New England peppermint (Eucalyptus nova-anglica) were harvested with the aid of a cherry picker to determine their biomass distribution and insect damage. One suffered from obvious symptoms of rural dieback and the other was healthy. Weights of foliage and wood were measured, and insect damage for all leaves and branches was quantified. For each tree 25% of the roots were extracted from the soil using a bulldozer and manual methods; they were then weighed and damage by insects estimated. The healthy tree lost more leaf surface area to insects (11% or 1.1 kg vs 9.2% or 0.3 kg); but the dieback tree had four times more wood affected by boring insects (19% cf. 5%); and only 20% root biomass remaining (92 kg cf. 488 kg). The accuracy of sampling techniques needed to measure defoliation and the consequences of insect damage to dieback of rural eucalypts are discussed.