Taphonomy and palaeoecology of Late Triassic (Carnian) ammonoid concentrations from the Taurus Mountains,Turkey

The deposits of the Carnian Kas?mlar Formation within the Taurus Platform Units of south‐western Turkey represent an important archive of a Late Triassic ecosystem. New palaeontological information was obtained by analysing the Kasimlarceltites mass occurrence, located within the Kas?mlar Formation and named after the Lower Carnian (Julian) ammonoid genus Kasimlarceltites. This is the dominant taxon (> 94%) within the mass occurrence: nearly 775 million ammonoids and 50 million gastropods were extrapolated for the whole extension (at least 5 km²) of the Kasimlarceltites beds. This calculation is one of the main findings within this study, as it is the first time that such a fossil mass occurrence was quantified. Additionally, orientation measurements of the planispiral ammonoids and the helical gastropods enabled reconstructing the history of the mass occurrence and interpreting the underlying transport mechanisms. Further taphonomic aspects (e.g. biofabric, preservation, bioerosion or genetic classification) as well as comparisons with samples of the same acme zone from different localities near A?a?iyaylabel (AS IV, KA I‐II) point to a two‐phased genetic history. Accordingly, local mass mortality within the Kasimlarceltites fauna due to oxygen fluctuations or methane degassing may have initially led to a primary accumulation. These deposits were then reworked and redeposited basinward by gravity flows to create the present‐day secondary allochthonous concentrations.     

New ammonoid taxa from the Lower Cretaceous Giumal Formation of the Tethyan Himalaya (Northern India)

Knowledge of the Early Cretaceous ammonoids of the NW‐Himalayas was poor until recent discoveries. Intense sampling from the Giumal Formation exposed near the village of Chikkim (Spiti Valley, Himachal Pradesh, India) led to the recognition of a new Early Cretaceous ammonoid fauna. The succession consists of arenitic sandstone interbedded with shale that was deposited by turbidity currents on an unstable shelf in the Early Cretaceous. Ammonoids have been obtained only from sandstone beds in the lower one‐third and close to the top of the c. 350‐m‐thick section. Eight new ammonoid taxa (1 genus and 7 species) are described: Sinzovia franki sp. nov. (rare), Giumaliceras giumaliense gen. et sp. nov. (abundant), Giumaliceras bhargavai gen. et sp. nov. (rare), Neocomites (Eristavites) platycostatiformis sp. nov. (rare), Cleoniceras oberhauseri sp. nov. (abundant), Australiceras himalayense sp. nov. (rare) and Deshayesites fuchsi sp. nov. (rare). Sinzovia and Deshayesites are reported for the first time from the Tethyan Himalaya. According to the biostratigraphic relevance of some ammonoid taxa described here, the age of the Giumal Formation can be constrained from Berriasian (Giumaliceras assemblage) to Aptian (Cleoniceras assemblage). The discovery of the new fauna substantiates the significance of the Giumal Formation around Chikkim and facilitates comparison with faunal assemblages from other regions in the Tethys Ocean and beyond.     

Late Jurassic decapod crustaceans from northeast Japan

Abstract: Astacidean and thalassinidean macrurans (Glyphea sp., ?Eryma sp. and Protaxius sp.) and a new longodromitid crab, Planoprosopon kashimaensis, are recorded from the Upper Jurassic (upper Kimmeridgian to lower Tithonian) of Fukushima Prefecture, northeast Japan. Material was collected from the Tatenosawa Sandstone Member of the Nakanosawa Formation, Somanakamura Group, from which abundant Tethyan‐type marine invertebrates are known. Planoprosopon kashimaensis sp. nov. closely resembles P. heydeni (von Meyer), a common form in the Upper Jurassic of the Tethyan realm in Europe, and represents the oldest record of a brachyuran from the circum‐Pacific region. Similarities to contemporaneous decapod assemblages in southern Germany indicate that closely comparable, parallel decapod faunas in the Tethyan realm, inclusive of brachyurans, had already been established in the western circum‐Pacific region by the Late Jurassic.     

SHARKS OF THE ORDER CARCHARHINIFORMES FROM THE BRITISH CONIACIAN, SANTONIAN AND CAMPANIAN (UPPER CRETACEOUS)

Abstract: Bulk sampling of phosphate‐rich horizons within the British Coniacian to Campanian (Upper Cretaceous) yielded very large samples of shark and ray teeth. All of these samples yielded teeth of diverse members of the Carcharhiniformes, which commonly dominate the fauna. The following species are recorded and described: Pseudoscyliorhinus reussi ( Herman, 1977 ) comb. nov., Crassescyliorhinus germanicus ( Herman, 1982 ) gen. nov., Scyliorhinus elongatus ( Davis, 1887 ), Scyliorhinus brumarivulensis sp. nov., ?Palaeoscyllium sp., Prohaploblepharus riegrafi ( Müller, 1989 ) gen. nov., ?Cretascyliorhinus sp., Scyliorhinidae incertae sedis 1, Scyliorhinidae incertae sedis 2, Pteroscyllium hermani sp. nov., Protoscyliorhinus sp., Leptocharias cretaceus sp. nov., Palaeogaleus havreensis Herman, 1977 , Paratriakis subserratus sp. nov., Paratriakis tenuis sp. nov., Paratriakis sp. indet. and ?Loxodon sp. Taxa belonging to the families ?Proscylliidae, Leptochariidae and Carcharhinidae are described from the Cretaceous for the first time. The evolutionary and palaeoecological implications of these newly recognised faunas are discussed.     

<Emphasis Type="Italic">Noriphyllia</Emphasis>, a new Tethyan Late Triassic coral genus (Scleractinia)

Noriphyllia gen. n. is a distinctive coral representing the Coryphylliidae, a group of Late Triassic scleractinian corals. Coral faunas of this age are poorly known. The new coral is distinguished from related corals belonging to the reimaniphylliids by key features of septal microstructure as discerned in thin sections. This microstructure consists of a straight/wavy midseptal zone and lateral septal stereome organized into thin fascicles of fibres, producing a fine and sharp micromorphology of the septal sides. The solitary genus Noriphyllia gen. n. contains two Early Norian species: N. anatoliensis sp. n. chosen as the type species and N. dachsteinae sp. n., and a Carnian species referred to as N. monotutoensis sp. n. The new genus is widely distributed in the Late Triassic, Early Norian reef facies of the Tethys region (Northern Calcareous Alps, Austria; Taurus Mountains, Turkey) and it also occurs in the Carnian of Timor. Noriphyllia gen. n. is unique and details of its microstructural features add new understanding to the composition of both Late Carnian and Early Norian corals.     

Early Silurian (Telychian) rugose coral fauna of Daguan area, northeast Yunnan province, China

Study on rugose coral fauna of the Sifengya Formation (early Telychian) and Daluzhai Formation (mid-late Telychian) in Daguan area, northeast Yunnan Province, China was carried out. Rugose coral fauna of the Sifengya Formation included 18 genera and 34 species, while Daluzhai Formation with nine genera, ten species. We described rugose coral fauna (12 genera, 19 species) including one new genus and five new species, i.e. Protoketophyllum daguanense gen. et sp. nov., Crassilasma huanggexiense sp. nov., Pseudophaulactis heae sp. nov., P. convolutus sp. nov., and Shensiphyllum minor sp. nov. The characteristics and geological significance of rugose coral fauna of Sifengya Formation and Daluzhai Formation were analyzed. Particularly, rugose coral fauna of the Sifengya Formation represent early Telychian rugosan fauna in the Upper Yangtze region and improve the sequences of early Silurian (Llandovery) rugose coral assemblages in Yangtze region. It is therefore very meaningful to further analyze radiation period of rugose coral fauna in such epoch. __________ Translated from Acta Palaeontologica Sinica, 2005, 44(2): 229–246 [译自: 古生物学报, 2005, 44(2): 229–246]     

Taxonomy of the fossil grylloblattid nymphs (Insecta: Grylloblattida)

Nymphs of fossil grylloblattid insects are revised. Newly described taxa are Lemmatonympha gracilissima gen. et sp. nov. and Kaltanympha vorcutensis sp. nov. from the Verkhne-Syr’yaginsk locality (Ufimian, Lek-Vorkuta Formation in the Vorkuta Coal Basin), genera Sylvalitoralis gen. nov. and Tshebardanympha gen. nov. from the Tshekarda locality (Kungurian, Koshelevka Formation, Middle Urals), Tataronymphakamensis gen. et sp. nov. from the Tikhie Gory locality (Lower Kazanian, linguloid beds of the Baitugan Formation, Tatarstan), and Kaltanympha ornata sp. nov. from the Kerbo locality (Upper Tatarian, lower part of the Degali Formation, Evenki Autonomous Region). Liomopterites (?) gracilis Sharov, 1961 from Lower Kazanian deposits of the Kuznetsk Formation in the Kuznetsk Basin is transferred to the genus Kaltanympha Sharov, 1961; Permonympha arcuata Sharov, 1957 from the same locality is synonymized under Permonympha gracile Sharov, 1957; the nymph described from the Karatau locality (Upper Jurassic, Karabastau Formation in southern Kazakhstan) as Blattogryllus karatavicus Rasnitsyn, 1976 is excluded from grylloblattids. Keys to extinct grylloblattid nymphs are provided.     

Revision of the family Idelinellidae,with a review of the Permian Eoblattida (Insecta)

Permian Eoblattida, which include the families Tillyardembiidae, Protembiidae (=Sylvardembiidae), Soyanopteridae, and Idelinellidae, are reviewed. The latter family is revised; it includes Idelinella macroptera Storozhenko, 1997 from the Middle Permian (Lower Kazanian Substage) of northern European Russia, Permostriga augustalis Novokshonov, 1999, Sylvastriga miranda Aristov, 2004, Strigulla cuculiophora (Aristov, 2002) comb. nov., Cucullistriga cucullata Aristov et Rasnitsyn, gen. et sp. nov., Scutistriga scutata Aristov et Rasnitsyn, gen. et sp. nov. from the Lower Permian (Kungurian Stage) of the Urals, Permeoblatta borealis Rasnitsyn et Aristov, 2010 from the Upper Permian (Upper Severodvinian Substage) of northern European Russia, and probably also Rasstriga americana Aristov et Rasnitsyn, gen. et sp. nov. from the Upper Carboniferous (Desmoinesian Stage) of Illinois.     

Oldest known Dicoelosia and Epitomyonia,deep water brachiopods from the Beiguoshan Formation (Middle Katian,Upper Ordovician), Shaanxi,north China

Abstract: A diverse brachiopod fauna from a relatively deep water carbonate facies of the Upper Ordovician Beiguoshan Formation (uppermost Caradoc – lower Ashgill, middle Katian) is characterized by small shells and contains the oldest known Dicoelosia and Epitomyonia, two diagnostic taxa of deep water brachiopod palaeocommunities during the Late Ordovician and Silurian. Three new species are recognized: Dicoelosia cordiformis sp. nov., Dicoelosia perbrevis sp. nov. and Epitomyonia fui sp. nov. These pioneer forms of the family Dicoelosiidae show a relatively high degree of morphological plasticity. The shells of Dicoelosia from the Beiguoshan Formation range from the typical slender‐lobed form with a concavoconvex profile to the strongly equibiconvex, fat‐lobed morphotype that was not known previously until the late Silurian. The Beiguoshan dicoelosiids point to an important attribute of the deep water brachiopods: small generalists with high morphological plasticity, which make them ideal candidates as progenitors for the evolution of shallow water brachiopod faunas in shelf and platform depositional environments.     

中国陕西中三叠世中蝎蛉科三新属新种(昆虫纲,长翅目)

记述中蝎蛉科Mesopanorpodidae 4属4种,其中包括3新属4新种：Longifurcula hejiafangensi gen.et sp.nov,Ladinochorista lata gen.et sp.nov.,Allochorista erdosensis gen.et sp.nov.,Itaphlebia tongchuanensis sp.nov..化石标本采自陕西铜川中三叠统铜川组下段上部的灰绿色泥岩和页岩.这些新属、种系陕西昆虫群铜川昆虫组合的新成员（属陕西动物群、陕西生物群的一个类别）.根据前人的植物、孢子、双壳、叶肢介、介形虫、昆虫化石等的研究结果,铜川组的时代划归中三叠世,相当于欧洲中三叠世拉丁尼期（Ladinian stage）.     

<Emphasis Type="Italic">Orientosirenites</Emphasis>, a New Ammonoid Genus (Sirenitidae; Ammonoidea) from the Upper Carnian of the Boreal Realm

The ammonoid family Sirenitidae from the Upper Carnian yakutensis Zone of northern Middle Siberia and Northeastern Asia is revised. A new genus Orientosirenites gen. nov. with the type species O. bytschkovi sp. nov. is established based on the study of the main morphological characters of the shell. The new genus and the species O. bytschkovi are described and the species O. yakutensis (Kiparisova) is redescribed. The new data allow the taxonomic composition and the distribution of the Carnian ammonoids of the Boreal Realm to be updated.     

滇东南泸西县上三叠统卡尼阶小凹组菊石*

本文对云南省东南部的泸西县白水镇小兴安村上三叠统卡尼阶小凹组菊石类进行了详细研究,共发现菊石9属13种,其中Anasirenites属为华南首次发现。依据各物种组合特征及其地层分布情况,自下而上识别出菊石Trachyceras multituberculatum带和Austrotrachyceras triadicum带,时代对应于晚三叠世早卡尼期的Julian 1和Julian 2。该菊石组合不仅记录了卡尼期Julian期Trachyceratidae的辐射事件,同时该菊石组合与卡尼期特提斯域和泛大洋域的菊石组合均有良好的对比性,并且为华南古地理研究提供了新的补充材料。另外,通过对剖面的岩性特征与菊石生物地层特征分析,认为在滇东南地区由于卡尼期湿润幕事件(CPE)所导致的碳酸盐岩台地消亡起始时间应为Julian 1末期略早于陆源碎屑输入的Julian 2亚期。     

New hymenopteran insects (Insecta: Vespida) from the lower or middle Jurassic of India

Hymenopterans Xyelula alexandri Rasnitsyn, sp. nov. (Sepulcidae) and Kotaphialtites frankmortoni Rasnitsyn, gen. et sp. nov. (Ephialtitidae) are described from the Lower or Middle Jurassic Kota Formation, Andhra Pradesh, India. These finds prove a considerable similarity between the Gondwanan and Laurasian hymenopteran faunas during the Jurassic, as well as during the Triassic and Cretaceous.     

Elasmobranchs from the Lower Triassic Sulphur Mountain Formation near Wapiti Lake (BC, Canada)

Hybodontoid and nonhybodontoid sharks are described from the Lower Triassic Vega‐Phroso Siltstone Member of Sulphur Mountain Formation on the basis of newly discovered material. The age of the classic fossil site ‘Wapiti Lake’ in the Canadian Rocky Mountains is discussed on the basis of new field data and one conodont found in association. Preliminary results suggest that these elasmobranch remains are between early Smithian and Spathian in age. Apart from the enigmatic genus Listracanthus and previously reported edestoids, the shark fauna consists of at least one hybodont, at least two questionable hybodontoid genera and an elasmobranch of enigmatic affinities, represented by peculiar denticles only and described as ‘genus A’incertae sedis. The presence of the only previously reported hybodont genus, cf. Palaeobates, is erroneous. The largest specimen represents the most complete Early Mesozoic shark known. The heterodonty of its dentition, fin spine morphology and the short, robust body shape imply it represents a member of a new family of shark, Wapitiodidae fam. nov. , and is described here as Wapitiodus aplopagus gen. et sp. nov. The unique dental morphology shows affinities to Polyacrodus but clearly differs in the complete lack of side cusps. Wapitiodus gen. nov. possesses a primitive fin spine structure. The tooth crowns are entirely blunt in the distal (posterior) tooth files, and are acuminate‐unicuspid in several anterior files. Tooth morphology, the shape of the basal cartilages, the proximal insertion of the fin spines and the pectoral fin structure are interpreted as diagnostic characters for this new genus, and possibly for the Wapitiodidae fam. nov. The majority of observed characters appear to be primitive and are reminiscent of Palaeozoic sharks, however, and these features include dorsal fin spine morphology and gross skull anatomy. A second species, provisionally placed in the same genus, is described as Wapitiodus homalorhizo sp. nov. Wapitiodus homalorhizo sp. nov. can be distinguished from W. aplopagus gen. et sp. nov. by the proportions of the fin spines, tooth morphology and possibly the body shape. Several isolated teeth and other fragmentary material are referred to either Wapitiodus gen. nov. sp. indet. or to ?Polyacrodus sp. (Polyacrodontidae gen. et sp. indet.). A third genus of elasmobranch (incertae sedis) is described as ‘Genus A’ and is recognized by its peculiar scales. © 2007 The Linnean Society of London, Zoological Journal of the Linnean Society, 2007, 149 , 309–337.     

吉林李家窑范家屯组中的二叠纪北温带牙形刺动物群

在吉林省九台市三台乡李家窑附近的二叠系范家屯组中采到一特殊的牙形刺动物群。多数牙形刺都较小,锯齿发育,齿脊细齿不愈合。由一些未见报道的新分子组成。本文描述了４个新种（Ｍｅｓｏｇｏｎｄｏｌｅｌｌａ ｃｈａｎｇｃｕｎｅｎｓｉｓ ｓｐ．ｎｏｖ．,Ｍ．ｊｉｌｉｎｅｎｓｉｓ ｓｐ．ｎｏｖ．,Ｍ．ｍｕｌｔｉｓｅｒｒａｔａ ｓｐ．ｎｏｖ．,Ｍ．ｐｓｅｕｄｏａｌｔｕｄａｅｎｓｉｓ ｓｐ．ｎｏｖ．）和３个未命名新种     

New radiolarian species from the Upper Cretaceous-Lower Paleogene of the eastern slope of the northern Ural Mountains

Five new radiolarian species, Dictyophimus? orbiculiporosus sp. nov., Spongomelissa imperceptusa sp. nov., Theocoronium puncticulatum sp. nov., Pterocyrtidium porrectum sp. nov., and Tripodiscinus corona sp. nov., from the Campanian-Danian diatomites of the Leplinskaya Formation of the northern Ural Mountains are described. They are useful for stratification and correlation of the Upper Cretaceous and Lower Paleogene beds of the Ural Region of Western Siberia.     

Lacewings of the family Osmylidae (Insecta: Neuroptera) from the Upper Jurassic of Asia

Three new genera and five new species of osmylid lacewings are described from the locality Karatau (Kazakhstan, Upper Jurassic): Jurosmylus parvulus sp. nov. (Protosmylinae), Ensiosmylus acutus gen. et sp. nov. (Spilosmylinae), Kolbasinella elongata gen. et sp. nov. (Gumillinae), Arbusella bella gen. et sp. nov. and Jurakempynus arcanus sp. nov. (Kempyninae). Another new osmylid species (Jurakempynus sublimis sp. nov.) is described from the Upper Jurassic of Mongolia (Shar-Teg locality). The quantitative compositions of the osmylid faunas of Karatau and Shar-Teg are discussed.     

Toarcian and Aalenian (Jurassic) ammonites from the Shemshak Formation of the Jajarm area (eastern Alborz, Iran)

In the Jajarm area (eastern Alborz Mountains, NE Iran) the ?Upper Triassic — Lower Bajocian siliciclastic Shemshak Formation is up to 2000 m thick. Whereas the lower third of the formation is nearly exclusively non-marine, the upper two-thirds are fully marine. The middle part is characterized by several levels containing a relatively diverse and well preserved assemblage of ammonites of the Toarcian to Aalenian stages. Two sections of the ammonite-bearing strata, spaced 20 km, are presented and correlated by means of ammonite biostratigraphy. The ammonite fauna consists of 27 taxa, some of which are recorded for the first time from the Alborz Mountains. The ammonites are briefly described and their palaeobiogeographic context is reviewed. The ammonite fauna of the Shemshak Formation at Jajarm, as elsewhere in North and Central Iran, is exclusively Northwestern Tethyan in character and is closely related to the faunas of Northwestern and Central Europe.     

NEOSELACHIANS FROM THE UPPER CAMPANIAN AND LOWER MAASTRICHTIAN (UPPER CRETACEOUS) OF THE SOUTHERN PYRENEES, NORTHERN SPAIN

Abstract: Bulk sampling of upper Campanian to lower–middle Maastrichtian coastal and lagoonal deposits in five sections of the Tremp Formation in the south‐central Pyrenees yielded numerous neoselachian teeth. The fauna comprises nine taxa of which three species and one genus are new: Hemiscyllium sp., Lamniformes indet., Paratrygonorrhina amblysoda gen. et sp. nov., Coupatezia trempina sp. nov., Coupatezia sp., Coupatezia? sp., Rhombodus ibericus sp. nov. and Igdabatis indicus. The neoselachian fauna is dominated by small nectobenthic rays. This composition resembles assemblages known from the marine Upper Cretaceous, but differs from nearby localities of the Basque‐Cantabrian region and continental selachian associations of the French Pyrenees. The results indicate that Rhombodus might not be a reliable biostratigraphic marker for the Maastrichtian. The faunal composition suggests a shallow trans‐Tethyan connection between Eurasia and India at the end of the Cretaceous Period.