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1.
Aim The influence of winter temperatures and other climate variables are explored to determine which variables are associated with saguaro stem diameter and to determine if Bergmann's rule is applicable to saguaros. Location The northern Sonoran Desert in Arizona, USA. Methods Thirty saguaro populations were sampled (height, diameter, number of branches), and after adjustment for population height structure, mean relative thickness of saguaros was calculated for each plot (population). Fifty‐seven climate variables were calculated for the thirty populations. Regression was run to determine which variables (cold winter, hot summer and precipitation) best predict relative thickness. Previous studies have demonstrated a significant positive relationship between winter precipitation and saguaro branching ( Drezner, 2003 , Ecography, in press). To determine if relative thickness may be an artefact of branching (branches), partial correlation analysis was employed. Results Mean March precipitation best predicts relative thickness. When only winter temperature variables are considered, none are significantly related to relative thickness. Relative thickness is not an artefact of branches. Main conclusions (1) Rainfall, not temperature, best predicts saguaro stem thickness. In addition, despite the focus on summer rains in the literature, winter precipitation is the best predictor of thickness. (2) Bergmann's rule is not applicable to saguaro populations as has been previously suggested [e.g. Niering et al. (1963) Science 142 , 15], as thickness does not increase significantly with latitude. In addition, the suggested mechanism for Bergmann's rule, cold winter temperature, does not significantly predict saguaro stem diameters over the area studied. In some populations that experience high winter rainfall as well as cold temperatures, individuals likely derive thermal benefits from a larger stem diameter; however, the trend is not observed over the area studied and it does not appear to be adaptive.  相似文献   

2.
Questions: We asked several linked questions about phenology and precipitation relationships at local, landscape, and regional spatial scales within individual seasons, between seasons, and between year temporal scales. (1) How do winter and summer phenological patterns vary in response to total seasonal rainfall? (2) How are phenological rates affected by the previous season rainfall? (3) How does phenological variability differ at landscape and regional spatial scales and at season and inter‐annual temporal scales? Location: Southern Arizona, USA. Methods: We compared satellite‐derived phenological variation between 38 distinct 625‐km2 landscapes distributed in the northern Sonoran Desert region from 2000 to 2007. Regression analyses were used to identify relationships between landscape phenology dynamics in response to precipitation variability across multiple spatial and temporal scales. Results: While both summer and winter seasons show increases of peak greenness and peak growth with more precipitation, the timing of peak growth was advanced with more precipitation in winter, while the timing of peak greenness was advanced with more precipitation in summer. Surprisingly, summer maximum growth was negatively affected by winter precipitation. The spatial variations between summer and winter phenology were similar in magnitude and response. Larger‐scale spatial and temporal variation showed strong differences in precipitation patterns; however the magnitudes of phenological spatial variability in these two seasons were similar. Conclusions: Vegetation patterns were clearly coupled to precipitation variability, with distinct responses at alternative spatial and temporal scales. Disaggregating vegetation into phenological variation, spanning value, timing, and integrated components revealed substantial complexity in precipitation‐phenological relationships.  相似文献   

3.
On the subtropical and tropical wintering grounds of migratory birds, variation in moisture levels and habitat can influence the availability of food resources and subsequently impact overwintering birds. Using stable carbon isotopes in blood samples as a measure of moisture, we assessed the interactive effects of rainfall, vegetation, and moisture on the demographics and condition of Prairie Warblers (Setophaga discolor) wintering in The Bahamas. Carbon isotopes in Prairie Warbler blood were more depleted in taller, wetter habitats; we additionally detected novel temporal effects of rainfall on isotope values. During a winter with more rainfall, most birds maintained mass and pectoral muscle regardless of the habitat type occupied. In a winter with less rainfall, birds lost mass and pectoral muscle, and this effect was more pronounced in birds with enriched isotope values and birds that occupied drier, shorter habitat. Prairie Warblers exhibited strong patterns of sexual habitat segregation with males disproportionately observed in areas with taller vegetation and females in shorter vegetation. During the drier winter, older males had better maintenance of pectoral muscle compared to females and younger individuals. Also in the drier winter, daily rainfall patterns explained more of the variation in body condition compared to the date of capture; pectoral muscle was best explained by recent precipitation (during the previous 30 days), while size‐corrected mass was more a function of longer‐term (90‐day) rainfall and habitat moisture. Our findings along with other studies suggest that Prairie Warblers and other migratory birds are sensitive to interactions between annual variation in winter rainfall, within‐season daily rainfall patterns, and habitat quality. Increasing drought and habitat loss in the Caribbean may be having a negative impact on wintering bird populations. To best conserve Nearctic–Neotropical migratory passerines in the region, we recommend prioritizing the protection of the least drought‐prone wintering areas.  相似文献   

4.
Esler  Karen J.  Rundel  Philip W. 《Plant Ecology》1999,142(1-2):97-104
A comparative study of community structure and seasonal growth dynamics in the arid winter rainfall regions of the Succulent Karoo in South Africa and the Mojave Desert of the United States suggests that remarkably divergent patterns of resource use and resultant growth form diversity exist in regions with outwardly similar climatic regimes. An understanding of these divergent patterns in the two winter rainfall deserts allows predictions to be made on vegetation response to global change. Above-ground plant growth in the Succulent Karoo begins with the first significant rains in late summer and continues through winter because moderate minimum temperatures allow continued growth. These communities have low structural diversity above-ground, but also below-ground, where root systems commonly do not exceed 20 cm in depth. These shallow root systems harvest water from upper soil horizons soon after rain falls, and growth declines as rainfall decreases in late spring. In contrast, low temperatures during winter inhibit growth in the Mojave Desert until early spring at a time when a mean 74% of the hydrologic year precipitation (July-June) has already occurred. Thus species in this structurally diverse system rely on deeper stores of water for growth in spring and early summer. A global change scenario of a 2 to 4°C increase in mean annual temperature and increased summer rainfall in the Mojave desert would be expected to produce similar conditions in the Mojave Desert to those that exist in the Succulent Karoo today. Assuming no genetic constraints on phenotypic plasticity, this would suggest increased species diversity and a decline in structural diversity in the Mojave Desert over evolutionary time. Increased summer rainfall in the Succulent Karoo would be expected to lead to invasions of grasses and thus increased competitive pressure reducing community diversity.  相似文献   

5.
We analyzed long-term organic and inorganic nitrogen inputs and outputs in precipitation and streamwater in six watersheds at the H.J. Andrews Experimental Forest in the central Cascade Mountains of Oregon. Total bulk N deposition, averaging 1.6 to 2.0 kg N ha–1 yr–1, is low compared to other sites in the United States and little influenced by anthropogenic N sources. Streamwater N export is also low, averaging <1 kg ha–1 yr–1. DON is the predominant form of N exported from all watersheds, followed by PON, NH4-N, and NO3-N. Total annual stream discharge was a positive predictor of annual DON output in all six watersheds, suggesting that DON export is related to regional precipitation. In contrast, annual discharge was a positive predictor of annual NO3-N output in one watershed, annual NH4-N output in three watersheds, and annual PON output in three watersheds. Of the four forms of N, only DON had consistent seasonal concentration patterns in all watersheds. Peak streamwater DON concentrations occurred in November-December after the onset of fall rains but before the peak in the hydrograph, probably due to flushing of products of decomposition that had built up during the dry summer. Multiple biotic controls on the more labile nitrate and ammonium concentrations in streams may obscure temporal DIN flux patterns from the terrestrial environment. Results from this study underscore the value of using several watersheds from a single climatic zone to make inferences about controls on stream N chemistry; analysis of a single watershed may preclude identification of geographically extensive mechanisms controlling N dynamics.  相似文献   

6.
Studies on mountain streams in the English Lake District   总被引:2,自引:0,他引:2  
Sodium and potassium in streamwater were measured over a period of 2 years, and concentrations of other major ions were occasionally monitored. The supply of sodium, potassium and calcium in rainfall was measured over a period of 1 year. A tentative budget is given for annual income and output of sodium, potassium and calcium in the Duddon catchment. Sodium and potassium concentrations in streamwater were independent of stream discharge; both ions displayed an annual cycle of high concentrations in winter and low concentrations in summer. The input of sodium was dependent on rainfall, wind direction and wind speed; 40% of the annual supply in rain was precipitated in the autumn. Sodium income in rainfall during the remaining 9 months of the year was insufficient to balance the output in streamwater. Annual income of potassium in rainfall was equal to annual output in streamwater. But potassium was retained on the catchment during spring and summer and was slowly released to the streams in autumn and winter. Only about 10% of the annual calcium output was derived from atmospheric sources.  相似文献   

7.
河北省典型区主要作物有效降雨量和需水量特征   总被引:11,自引:0,他引:11  
对作物全生育期内有效降雨量及需水规律的研究是进行合理灌溉及水资源优化配置的重要依据。以河北省鸡泽县为典型区域,利用FAO推荐的Penman-Monteith公式和分段单值平均作物系数法对冬小麦、夏玉米和棉花的有效降雨量及需水规律进行计算,并通过M-K检验法分析近60 a来鸡泽县有效降雨量及作物需水量序列的趋势变化及突变现象。结果表明:近60 a鸡泽县冬小麦在生育中期需水量最大,为210.2 mm,有效降雨量均集中在生育中期,缺水量以13.2 mm/10 a的速率呈显著性减少趋势;夏玉米在初始生长期需水量最大,为157.7 mm,有效降雨量集中在生育中期,缺水量以7.0 mm/10 a的速率呈不显著减少趋势;棉花在快速发育期需水量最大,为227.9 mm,有效降雨量集中在生育中期,缺水量以22.3 mm/10 a的速率呈显著性减少趋势。在一定程度上对河北省对农田灌溉用水效率和效益以及保障农作物科学高效生产具有重要的指导意义。  相似文献   

8.
Differential utilization of summer rains by desert plants   总被引:21,自引:0,他引:21  
Summary Seasonal changes in the hydrogen isotope ratios of xylem waters were measured to determine water sources used for growth in desert plants of southern Utah. While all species used winter-spring recharge precipitation for spring growth, utilization of summer rains was life-form dependent. Annuals and succulent perennials exhibited a complete dependence on summer precipitation. Herbaceous and woody perennial species simultaneously utilized both summer precipitation and remaining winter-spring precipitation, with herbaceous species much more reliant on the summer precipitation component. Several of the woody perennials exhibited no response to summer precipitation. Currently, precipitation in southern Utah is evenly partitioned between winter and summer time periods; however, global circulation models predict that summer precipitation will increase in response to anticipated climate change. Our data indicate that components within the community will differentially responde to the change in precipitation patterns. These results are discussed in relation to impact on competition and possible changes in community structure.  相似文献   

9.
Arizona and New Mexico receive half of their annual precipitation during the summer monsoon season, making this large-scale rain event critical for ecosystem productivity. We used the monsoon rains to explore the responses of soil bacterial and fungal communities to natural moisture pulses in a semiarid grassland. Through 454 pyrosequencing of the 16S rRNA gene and ITS region, we phylogenetically characterized these communities at 22 time points during a summer season. Relative humidity increased before the rains arrived, creating conditions in soil that allowed for the growth of microorganisms. During the course of the study, the relative abundances of most bacterial phyla showed little variation, though some bacterial populations responded immediately to an increase in soil moisture once the monsoon rains arrived. The Firmicutes phylum experienced over a sixfold increase in relative abundance with increasing water availability. Conversely, Actinobacteria, the dominant taxa at our site, were negatively affected by the increase in water availability. No relationship was found between bacterial diversity and soil water potential. Bacterial community structure was unrelated to all environmental variables that we measured, with the exception of a significant relationship with atmospheric relative humidity. Relative abundances of fungal phyla fluctuated more throughout the season than bacterial abundances did. Variation in fungal community structure was unrelated to soil water potential and to most environmental variables. However, ordination analysis showed a distinct fungal community structure late in the season, probably due to plant senescence.  相似文献   

10.
Climate change is expected to impact the amount and distribution of precipitation in the arid southwestern United States. In addition, nitrogen (N) deposition is increasing in these regions due to increased urbanization. Responses of belowground plant activity to increases in soil water content and N have shown inconsistent patterns between biomes. In arid lands, plant productivity is limited by water and N availability so it is expected that changes in these factors will affect fine root dynamics. The objectives of this study were to quantify the effects of increased summer precipitation and N deposition on fine root dynamics in a Mojave Desert ecosystem during a 2‐year field experiment using minirhizotron measurements. Root length density, production, and mortality were measured in field plots in the Mojave Desert receiving three 25 mm summer rain events and/or 40 kg N ha?1 yr?1. Increased summer precipitation and N additions did not have an overall significant effect on any of the measured root parameters. However, differences in winter precipitation resulting from interannual variability in rainfall appeared to affect root parameters with root production and turnover increasing following a wet winter most likely due to stimulation of annual grasses. In addition, roots were distributed more deeply in the soil following the wet winter. Root length density was initially higher under canopies compared to canopy interspaces, but converged toward the end of the study. In addition, roots tended to be distributed more deeply into the soil in canopy interspace areas. Results from this study indicated that increased summer precipitation and N deposition in response to climate change and urbanization are not likely to affect fine root dynamics in these Mojave Desert ecosystems, despite studies showing aboveground plant physiological responses to these environmental perturbations. However, changes in the amount and possibly distribution of winter precipitation may affect fine root dynamics.  相似文献   

11.
We studied the spatial patterns and temporal dynamics of vegetation structural responses to precipitation variation in grassland, transitional, and desertified‐shrubland ecosystems in an 800 km2 region of Northern Chihuahua, USA. Airborne high‐fidelity imaging spectroscopy data collected from 1997 to 2001 provided spatially detailed measurements of photosynthetic and senescent canopy cover and bare soil extent. The observations were made following wintertime and summer monsoonal rains, which varied in magnitude by >300% over the study period, allowing an assessment of ecosystem responses to climate variation in the context of desertification. Desertification caused a persistent increase in both photosynthetic vegetation (PV) and bare soil cover, and a lasting decrease in nonphotosynthetic vegetation (NPV). We did not observe a change in the spatial variability of PV cover, but its temporal variation decreased substantially. In contrast, desertification caused the spatial variability of NPV to increase markedly, while its temporal variation did not change. Both the spatial and temporal variation of exposed bare surfaces decreased with desertification. Desertification appeared to be linked to a shift in seasonal precipitation use by vegetation from mainly summer to winter inputs, resulting in an apparent decoupling of vegetation responses to inter‐annual monsoonal variation. Higher winter rainfall led to decreased springtime spatial variability in the PV cover of desertified areas. Higher summer rainfall resulted in decreased PV cover variation in grassland, transition and desertified‐shrubland regions. The effects of desertification on NPV dynamics were more than three times greater than on PV or bare soil dynamics. Using remotely sensed PV and NPV as proxies for net primary production (NPP) and litter dynamics, respectively, we estimated that desertification decreases the temporal variability of NPP and increases spatial variation of litter production and loss. Quantitative studies of surface biological materials and ecosystem processes can now be measured with high ‘structural’ detail using imaging spectroscopy and shortwave‐infrared spectral mixture analysis.  相似文献   

12.
Moisture inputs drive soil respiration (SR) dynamics in semi-arid and arid ecosystems. However, determining the contributions of root and microbial respiration to SR, and their separate temporal responses to periodic drought and water pulses, remains poorly understood. This study was conducted in a pine forest ecosystem with a Mediterranean-type climate that receives seasonally varying precipitation inputs from both rainfall (in the winter) and fog-drip (primarily in the summer). We used automated SR measurements, radiocarbon SR source partitioning, and a water addition experiment to understand how SR, and its separate root and microbial sources, respond to seasonal and episodic changes in moisture. Seasonal changes in SR were driven by surface soil water content and large changes in root respiration contributions. Superimposed on these seasonal patterns were episodic pulses of precipitation that determined the short-term SR patterns. Warm season precipitation pulses derived from fog-drip, and rainfall following extended dry periods, stimulated the largest SR responses. Microbial respiration dominated these SR responses, increasing within hours, whereas root respiration responded more slowly over days. We conclude that root and microbial respiration sources respond differently in timing and magnitude to both seasonal and episodic moisture inputs. These findings have important implications for the mechanistic representation of SR in models and the response of dry ecosystems to changes in precipitation patterns.  相似文献   

13.
Abstract Chemical knockdown procedures were used to sample canopy arthropods at 3 month intervals over 1 year at two sites, one in eastern Australia and the other in western Australia. Samples were taken from narrow-leaved ironbark, Eucalyptus crebra, and grey box, Eucalyptus moluccana, in the east and from jarrah, Eucalyptus marginata, and marri, Eucalyptus calophylla, in the west. Arthropods were more abundant on trees in eastern Australia and exhibited different seasonal patterns from those in the west. Members of different functional groups exhibited different seasonal patterns, with some herbivorous groups responding to times of leaf production, decomposers and fungus feeders responding to high moisture availability, and predators/parasites responding to the abundance of food items. Seasonal variability was slightly higher in the west, possibly reflecting the greater seasonal amplitude in rainfall. In the eastern forest, proportionately more taxa peaked in spring or summer and declined to minimum numbers in winter. In the western forest several taxa attained peak numbers in autumn, winter or spring, while others declined to minimum values in winter or summer. The phenological patterns of canopy arthropods appear to be linked to the condition of the host plant and/or to climatic factors. Comparison of the western Australian data to those from a second year of sampling at a time when rainfall was greater and fell later into the season indicated that variability in arthropod numbers between years can be as great as that between seasons. Implications of the variability in seasonal and annual patterns of canopy invertebrate communities are discussed in relation to the need for long-term sampling and in relation to evaluating the impact of disturbance on forest communities.  相似文献   

14.
Summary Geographical distribution and annual periodicity of the Australian Zebra Finch(Taeniopygia guttata castanotis) are determined mainly by temperature and precipitation. Low temperatures are the limiting factors in the eastern and southern, rich and long-lasting rainfalls in the northern parts of the continent. A special situation exists in southwestern Australia as well as in the western parts of South Australia: here, the temporal relation between rainfall and low temperatures seems to be essential in limiting breeding success and thus limiting geographical distribution, too. In all regions with summer rains the Zebra Finch is able to exist at lower mean temperatures than it does in regions with winter rains only. A minor limiting factor is the existence of dense woodland which is strictly avoided by the species.Man has influenced geographical districution of the Zebra Finch favourably (by clearing woodland, constructing bores and cattle-troughs, releasing cage-birds) and unfavourably (by introducing superior species of birds).The breeding season of the species is influenced merely by rainfall in the northern and central parts of Australia, merely by temperature in the eastern and southern parts and — as mentioned — by the connection of low temperatures and rainfall in the southwestern parts.  相似文献   

15.
土壤表面干沙层在冬春少雨季节较厚,流动沙地的又明显地大于固和地的。流动沙地地表干沙层以下的土壤含水量一般在于同深化度固定沙地的,特别是在报系密集分布的土层。科季或早春土壤含水量比较低,雨季的明显较高,雨季结束后,土层中的水分在重力的作用下会向深层运动,导致根系层土壤含水量下降。在植物生长季,缺少降雨时,因少地的土土壤水量会因植物的蒸腾消耗而变得很低(1%~2%),而流动沙地的不受或受此影响很小。地  相似文献   

16.
The ecology and distribution of Quercus aegilops ssp. brantii and the associated Pistacia atlantica var. mutica and P. khinjuk are examined as a basis for explaining the Late Quaternary history of the forest in the Zagros mountains. The three species are tolerant of aridity and low temperatures but sensitive to heavy snowfall. In most of the area, summer rain does not occur. Forest expansion is generally limited by the inability of seedlings to survive the four-month summer drought. Pollen diagrams from Lake Zeribar, Kurdistan show the absence of trees during the last glacial period and the migration of forest into the region between 10 000 and 5 500 B.P. This has been interpreted as indicating aridity during the Pleistocene with gradually increasing precipitation in the late glacial and Holocene. However, the sensitivity of these species to snow and their tolerance of low overall precipitation implicate higher snowfall rather than lower precipitation as the cause of absence during the Pleistocene. The inability of seedlings to survive the present summer-dry conditions suggests that summer rainfall, and not higher total precipitation, was the factor that finally allowed migration. These conclusions are supported by independent evidence of increased winter precipitation during the pleniglacial and a period of summer rainfall but low annual precipitation during forest expansion.  相似文献   

17.
The question of how aridity might influence avian clutch size, through the influences of rainfall seasonality and environmental stochasticity (unpredictability), has received little attention. A marked east-west gradient in aridity across South Africa provides a unique opportunity to test for such influences. Using an extensive collection of nest records for 106 terrestrial bird species from the South African Nest Record Card Scheme, we tested three predictions related to rainfall seasonality and stochasticity. Analyses were conducted at two levels, the first examining each species independently, and the second grouping species into five dietary guilds. The first prediction, that clutch size should generally increase with higher rainfall seasonality (i.e. higher seasonal fluctuation of food availability), was supported, particularly in the most arid environments where food abundance is more closely linked to rainfall. Controlling for rainfall seasonality, the second prediction, that clutch size should generally decrease as a bet-hedging strategy in arid, stochastic environments, was also supported. Although the timing of the rainy season differs among regions in South Africa (winter, early summer, later summer, year-round), birds primarily nest during spring. The relative timing of rainfall and breeding is expected to have different consequences for seasonal variation in clutch size among rainfall regions. The third prediction, of different patterns of seasonal variation in clutch size between rainfall regions, was also supported. In the winter and early-summer rainfall regions, early-nesting birds (breeding with or soon after the rains) generally had a larger clutch size than late-nesting birds. In the late-summer rainfall region, early-nesting birds (breeding well before the rains) had a smaller clutch size than late-nesting birds.  相似文献   

18.
Several aspects of terrestrial ecosystems are known to be associated with the North Atlantic Oscillation (NAO) through effects of the NAO on winter climate, but recently the winter NAO has also been shown to be correlated with the following summer climate, including drought. Since drought is a major factor determining grassland primary productivity, the hypothesis was tested that the winter NAO is associated with summer herbage growth through soil moisture availability, using data from the Park Grass Experiment at Rothamsted, UK between 1960 and 1999. The herbage growth rate, mean daily rainfall, mean daily potential evapotranspiration (PE) and the mean and maximum potential soil moisture deficit (PSMD) were calculated between the two annual cuts in early summer and autumn for the unlimed, unfertilized plots. Mean and maximum PSMD were more highly correlated than rainfall or PE with herbage growth rate. Regression analysis showed that the natural logarithm of the herbage growth rate approximately halved for a 250 mm increase in maximum PSMD over the range 50-485 mm. The maximum PSMD was moderately correlated with the preceding winter NAO, with a positive winter NAO index associated with greater maximum PSMD. A positive winter NAO index was also associated with low herbage growth rate, accounting for 22% of the interannual variation in the growth rate. It was concluded that the association between the winter NAO and summer herbage growth rate is mediated by the PSMD in summer.  相似文献   

19.
A modern analogue technique is applied to two high-resolution pollen sequences from NW Romania to provide the first quantitative evidence for winter, summer and annual temperatures and for precipitation across the Holocene in this region.

The pollen-based climate reconstructions allow the identification of four main intervals: i) an early, less stable period between 11,700 and 11,200 cal. yr BP; (ii) generally stable conditions between 11,200 and 8300 cal. yr BP with winter and annual temperatures and precipitation higher than at present, and summer temperatures about the same; (iii) lower winter and annual temperatures, and higher summer temperatures and precipitation between 8000 and 2400 cal. yr BP; (iv) warmer winter and annual temperatures and lower precipitation for the last 2400 years, whereas summer temperatures became cooler at Steregoiu and remained stable at Preluca Tiganului.

The pollen-based climate reconstructions at the two sites show similar patterns in annual and winter temperatures and precipitation changes during the Holocene, but the trends appear to be less consistent for summer temperatures.

Our pollen-based reconstructions revealed several short-term climatic oscillations during the Holocene, the strongest of which occur between 10,300–10,100, 8300–8000, 6800–6400, 5100–4900, 4000–3600 and 3200–3000 cal. yr BP.  相似文献   


20.
降雨和土壤湿度对贵州旱田土壤N2O释放的影响   总被引:15,自引:0,他引:15  
以南方亚热带代表性旱田土壤-贵州玉米-油菜轮作田、大豆-冬小麦轮作田和休耕地为观测对象,研究土壤N2O释放通量季节变化与降雨和土壤湿度的关系,同时,采用DNDC模型定量探讨了未来降雨量变化对土壤N2O释放的潜在影响,结果表明,降雨与N2O释放峰间存在明显的驱动-响应关系,N2O释放通量与降雨量和土壤湿度间存在正相关性,模型检验结果表明,夏秋季土壤N2O释放通量与降雨量变化呈正相关,而降雨量的大幅度增加或下降将引起冬春季土壤N2O释放通量的微弱下降。  相似文献   

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