New Asellota from the Antarctic deep sea (Crustacea, Isopoda, Asellota), with descriptions of two new genera

Two new genera, Reductosoma gen.n. (Desmosomatidae) and Desmostylis gen.n. (Desmosomatidae), and five new species are described from the Maud Rise and the Gunnerus Ridgc, Antarctica. Two new species of the Desmosomatidae (Eugerdellatinae and Desmosomatinae) are described: Reduetosoma gunnera gen. et sp.n. and Desmosoma hesslera sp.n.; one species of the Munnopsidae (Eurycopinae), Disconectes colernuni sp.n.; and two species of the Macrostylidae from the Antarctic deep sea, Desmostylis obscurus gen. et sp.n. and Macrostylis sarsi sp.n. Comments on the phylogeny and zoogeography are given.     

Assessment of monophyly of species-groups within Afrotropical Empidini (Diptera: Empididae: Empidinae), with a cladistic analysis of the Empis setitarsus-group

Abstract.  The Afrotropical Empidini are reviewed from a phylogenetic point of view. Seven monophyletic groups are recognized within the tribe, namely Empis ( Coptophlebia ) bivittata , E. ( C ) chrysocera , E. ( C ) hyalea , E. ( C ) namwamba , E. ( C ) poecilosoma , E. ( Disneyempis ) and E. setitarsus . Their monophyly is assessed by comparison with the remaining Empidini. The E. setitarsus -group is cladistically analysed to study the evolution of the length of vein M1 by optimization on the cladogram, and the status of subgenus Coptophlebia Bezzi is discussed. A key to the seven Afrotropical species-groups of Empidini is provided, genus Rhamphomyia is no longer recognized for the Afrotropical Region, and four new species belonging to the E. setitarsus -group, namely E. cinerarius sp.n., E. pseudosetitarsus sp.n., E. spinosa sp.n and E. spungaberaensis sp.n., are described and included in a key to all known species of this group.     

A revision of the Oriental stonefly genus Phanoperla (Plecoptera: Perlidae)

The genus Phanoperla (=Dyaperla Banks, 1939) (Plecoptera: Perlidae: Perlini) is revised and generic diagnoses are provided for adults and larvae. Diagnostic and constitutive characters of the tribe Neoperlini are discussed, and Chinoperla Zwick, 1980, is shown to be the closest relative of Phanoperla. Many past misidentifications of Phanoperla species have been corrected by the use of characters recently recognized as important, namely the structure of the internal genitalia of male and female specimens and particularly the complex pattern of spines on the male penial sac made visible by eversion of this structure, and details of sculpturing of the egg chorion.
The following species of Phanoperla are recognized: amorpha sp.n.; anomala (Banks, 1939); bakeri (Banks, 1924); ceylonica Kawai, 1975; comuta sp.n.; flaveola (Klapálek, 1921), comb.n. ( =clarissa (Banks, 1913), syn.n.), (=Neoperla hageni Banks, 1920, syn.n.), (= N.consimilis Banks, 1924, syn.n.); guttata sp.n.; himalayana Zwick, 1977 (= siwalika Harper, 1977); limosa (Hagen, 1858); maculata sp.n.; maindroni (Navas, 1926), comb.n.; malayana sp.n.; minutissima (Enderlein, 1909); nana sp.n.; nervosa Banks, 1939; nuwara Kawai, 1975; omega sp.n.; pallipennis (Banks, 1938); parva sp.n.; pumilio (Klapálek, 1921), comb.n.; peniculus Kawai, 1969a; schmidi sp.n.; sertispina Jewett, 1975; srilanka sp.n.; sumatrae sp.n.; testacea (Hagen, 1858); wedda sp.n. P.claggi (Banks, 1938) is a nomen nudum.
All species are (re-)described and figured; all primary types have been examined. A key to species is provided. Most species can be assigned to one of seven species groups which are defined.
Phanoperla is endemic to the Oriental Region. Species groups are generally widespread, but individual species are in most cases known only from restricted areas.     

Revision of the Australian Coccinellidae (Coleoptera). Part 3. Tribe Sukunahikonini

Abstract  The Australian members of the circumtropical coccinellid tribe Sukunahikonini are revised and placed in three genera: Scymnomorphus Weise (five species), Pharellus Sicard (two species) and Paraphellus Chazeau (two species). Scymnus rostratus Lea, 1929 is transferred to Paraphellus as P. rostratus (Lea) comb. n. The following new species are described from Australia: Pharellus glabratus sp. n., P. popei sp. n.; Paraphellus magnopunctatus sp. n.; Scymnomorphus fulvus sp. n., S. hirtus sp. n., S. ker sp. n., S. luteus sp. n., S. storeyi sp. n. Nomenclatural history, diagnoses and distribution are provided for each species. Keys to the genera and species are included.     

The taxonomy and diversity of Platerodrilus (Coleoptera,Lycidae) inferred from molecular data and morphology of adults and larvae

The Oriental neotenic net-winged beetles attracted attention of biologists due to conspicuous large-bodied females; nevertheless phylogenetic relationships remain contentious and only a few species are known in both the fully metamorphosed males and neotenic females. The phylogenetic analyses and morphology of larvae and adults provide data for investigation of relationships and species delineation. Platrilus Kazantsev, 2009, Platerodriloplesius Wittmer, 1944, and Falsocalochromus Pic, 1942 are synonymized to Platerodrilus Pic, 1921. Platrilus hirtus (Wittmer, 1938) and Pl. crassicornis (Pic, 1923) are transferred to Platerodrilus Pic, 1921. Platerodrilus hoiseni Wong, 1996 is proposed as a junior subjective synonym of Falsocalochromus ruficollis Pic, 1942. Platerodrilus is divided in three species-groups: P. paradoxus, P. major, and P. sinuatus groups defined based on the shape of genitalia and molecular phylogeny. The following species are described: Platerodrilus foliaceus sp. n., P. wongi sp. n. (P. paradoxus group); P. ngi sp. n., P. wittmeri (P. major group), P. ijenensis sp. n., P. luteus sp. n., P. maninjauensis sp. n., P. montanus sp. n., P. palawanensis sp. n., P. ranauensis sp. n., P. sibayakensis sp. n., P. sinabungensis sp. n., P. talamauensis sp. n., and P. tujuhensis sp. n. (P. sinuatus group). P. korinchiana robinsoni Blair, 1928 is elevated to the species rank as P. robinsoni Blair, 1928, stat. n. The conspecific semaphoronts are identified using molecular phylogeny for P. foliaceus sp. n., P. tujuhensis sp. n., P. montanus sp. n., P. maninjauensis sp. n.; additional female larvae are assigned to the species-groups. Diagnostic characters are illustrated and keys are provided for P. paradoxus and P. major groups.     

The pterolichoid feather mites (Acarina, Astigmata) of the Megapodiidae (Aves, Galliformes)

The pterolichoid feather mites of megapodes are reviewed. Named taxa are briefly discussed and most are figured. The Pterolichidae (Pterolichinae) are: Ascetohchus Perez & Atyeo, three species; Echinozonus Atyeo & Perez, six species; Pereziella Atyeo, two species; and Phycoferus Atyeo & Perez, two species. New pterolichine taxa, with the type species listed first, include: Botryaspis cordiforma gen. et sp.n. and B. cordata gen. et sp.n.; Cycloprotarsus lineatus gen. et sp.n., C. centralis gen. et sp.n. and C. monacrotrichus gen. et sp.n.; Eurypterolichus gen.n. for Pterolichus navicula comb.n. Trouessart & Neumann and E. coniger gen. et sp.n.; Goniodurus gen.n. for Pterolichus ( Pseudalloptes ) quadratus comb.n. Trouessart and G. bilobatus gen. et sp.n.; Haptepigynus gen.n. for Pterolichus ( Pseudalloptes ) tridentiger comb.n. Trouessart and H. holonotus gen. et sp.n.; Heliaspis ventralis gen. et sp.n.; Leipobius ocellatus gen. et sp.n.; Maleolichus maleo gen. et sp.n.; Mayracarus gen.n. for Pterolichus (P.) tritilobus comb.n. Trouessart; Megapodobius arcuatus gen. et sp.n. and M. striatus gen. et sp.n.; Oxygynurus brevissimus gen. et sp.n., O. longicaulis gen. et sp.n., O. mediocaulis gen. et sp.n. and O. parvicaulis gen. et sp. n.; Prionoturus amembranatus gen. et sp. n.; Talegallobius bidentatus gen. et sp.n.; and Tanysomacarus imperfectus gen. et sp.n. and T. brachymeles gen. et sp.n. A new taxon of the Thoracosathesidae is: Thoracosathes caudiculata sp.n. Keys are provided and host- commensal associations are discussed. All taxa are restricted to the Megapodiidae. Cheylabis fuscina Trouessart is assigned to Pereziella and has as a synonym P. dupilcata Atyeo.     

The nematode fauna in the sulphide–rich brine seep and adjacent bottoms of the East Flower Garden, NW Gulf of Mexico. II. Monhysterida

Eight nematode species of the order Monhysterida are described from the East Flower Garden at 72 m depth in the north–western Gulf of Mexico. The material is from sandy and rocky bottom samples influenced or beyond the influence o f a sulphide–rich brine seep. Seven species are new to science: Monhystera anoxybiotica sp.n., Gonionchus intermedius sp.n., Theristus (Theristus) coplatus sp.n., Theristus (Theristus) rezaki sp.n., Xyala oxybiotica spa., Desmolaimoides thiobioticus gen. et sp.n. and Linhomoeus gittingsi sp.n. Specimens of a Linhomoeus species are also described. The species is probably new to science, but the available material only includes females and juveniles.     

Phylogenetic revision of the Taenogera Kröber genus-group (Diptera: Therevidae), with descriptions of two new genera

The phylogenetic relationships of the Taenogera genus-group, consisting of nine genera, are examined using cladistic methods. Twenty-one species representing nine genera and one outgroup were compared in a cladistic analysis across 99 states in 44 characters. The genus Taenogera Kröber is revised to contain only T. longa (Schiner), T. nitida (Macquart) and T. notatithorax Mann. Two new genera are described and figured: Actenomeros gen. n., with two species, A. corniculaticaudus sp. n. and A. onyx sp. n.; and Taenogerella gen. n., with four species, Ta. elizabethae sp. n., Ta. platina sp. n., Ta. schlingeri sp. n. and Ta . nigrapicalis (Mann) comb. n. Phylogenetic analysis also supports the removal of Nanexila gracilis (Mann) comb. n. from Taenogera .     

Rhachicreagra (Acrididae, Ommatolampinae): forest grasshoppers from Central america with unique aedeagal asymmetry

Abstract. Rhachicreagra species are confined to Central American wet tropical forests, both lowland and montane. The asymmetric aedeagus with the left penis valve suppressed is unique in the Acridoidea. Keys are provided to both sexes of the seventeen recognized species: achrosta sp.n., anchidiphalara sp.n., astytophallus sp.n., brachysphagiocerca sp.n., drymocnemensis sp.n., gracilis Bruner, Haematodes sp.n., Khayachorsa sp.n., himantocerca sp.n. maya sp.n., melanota sp.n., sp.n., mexicana Hebard, nothra Rehn (= pallipes Bruner, = aeruginosa Bruner syn.n), obsidian sp.n., olmeca sp.n., pomatiaphallus sp.n., sphagicerca sp.n.; the small geographical ranges of these species argue for great care in the choice of representative forest blocks for purposes of faunal conservation in Central America.     

Fern sporophagy in Coleoptera from the Juan Fernandez Islands,Chile, with descriptions of two new genera in Cryptophagidae and Mycetophagidae

Abstract. Fern sporophagy is reported in species of Cryptophagidae, Mycetophagidae and Anthribidae occurring on the Juan Fernandez Islands, Chile. The following taxa are described and/or discussed: Cryptophagidae: Cryptothelypterus gen.n., with five species: C.obrieni sp.n., C.pteropilosus sp.n., C.selkirki (Bruce) comb.n., C.skottsbergi (Bruce) comb.n., and C.splendens (Bruce) comb.n.; Mycetophagidae: Filicivora gen.n., with one species, F.chilensis (Philippi & Philippi) comb.n.; and Anthribidae: Opisolia lenis Jordan. A key to the species of Cryptothelypterus is provided. Convergent adaptations for fern sporophagy in these groups are discussed, hypotheses are given for the evolution of this habit, and comments are made on wing atrophy.     

Revision of the Malagasy genus Trichoteleia Kieffer (Hymenoptera, Platygastroidea, Platygastridae)

The species of the genus Trichoteleia Kieffer (Hymenoptera: Platygastridae) are revised: 42 species are recognized, of which two were previously named and are redescribed: Trichoteleia afo Talamas, sp. n., Trichoteleia albidipes Kieffer, Trichoteleia bicolor Talamas, sp. n.; Trichoteleia bidentata Talamas sp. n.; Trichoteleia carinata Talamas, sp. n.; Trichoteleia cincta Talamas & Masner, sp. n.; Trichoteleia delilah Talamas, sp. n.; Trichoteleia eburata Talamas, sp. n.; Trichoteleia echinata Talamas, sp. n.; Trichoteleia fisheri Talamas & Masner, sp. n.; Trichoteleia funesta Talamas, sp. n.; Trichoteleia halterata Talamas & Masner, sp. n.; Trichoteleia hemlyae Talamas & Masner, sp. n.; Trichoteleia irwini Talamas & Masner, sp. n.; Trichoteleia janus Talamas, sp. n.; Trichoteleia jiro Talamas, sp. n.; T. ketrona Talamas, sp. n.; Trichoteleia levii Talamas & Johnson, sp. n.; Trichoteleia longiventris Talamas & Masner, sp. n.; Trichoteleia minima Talamas, sp. n.; Trichoteleia nify Talamas & Masner, sp. n.; Trichoteleia oculea Talamas, sp. n.; Trichoteleia orona Talamas & Masner, sp. n.; Trichoteleia parvipennis Talamas & Masner, sp. n.; Trichoteleia pauliani (Risbec); Trichoteleia picturata Talamas, sp. n.; Trichoteleia prima Talamas, sp. n.; Trichoteleia prolixa Talamas, sp. n.; Trichoteleia quazii Talamas, sp. n.; Trichoteleia ravaka Talamas, sp. n.; Trichoteleia rugifrons Talamas & Masner, sp. n.; Trichoteleia solocis Talamas, sp. n.; Trichoteleia sphaerica Talamas, sp. n.; Trichoteleia subtilis Talamas & Masner, sp. n.; Trichoteleia tahotra Talamas & Masner, sp. n.; Trichoteleia takariva Talamas, sp. n.; Trichoteleia tezitra Talamas, sp. n.; Trichoteleia tigris Talamas, sp. n.; Trichoteleia tonsa Talamas, sp. n.; Trichoteleia warreni Talamas & Masner, sp. n.; Trichoteleia xantrox Talamas, sp. n.; Trichoteleia zuparkoi Talamas & Masner, sp. n. A neotype is designated for Trichoteleia albidipes and a lectotype is designated for Trichoteleia pauliani.     

A review of the Agabus affinis group (Coleoptera: Dytiscidae), with the description of a new species from Siberia and a proposed phylogeny

Abstract. Agabus yakutiae sp.n., from Siberia, is placed in the A.affinis (Paykull) species-group, characterized by the unique structure of the male stridulatory organ formed by sternum 3 and the metafemora. The group is defined, the two Nearctic and four Palaearctic species included are reviewed, and a key is given for their identification. Phylogenetic relationships with other Agabus species groups are discussed, and a phylogeny for the species in the affinis-group is suggested.     

Two new rhabditida species (Nematoda: Rhabditidae) parasites of Cyclocephala signaticollis (Coleoptera: Scarabaeidae) in Argentina

Two different Rhabditida species (Nematoda: Rhabditidae) Parasitorhabditis platidontus n. sp. and Cruznema campestris n. sp. are described and illustrated from the larvae of Cyclocephala signaticollis (Coleoptera: Scarabaeidae) from Buenos Aires province, Argentina. Parasitorhabditis platidontus n. sp. is characterized by having 4 odontoplates in the stoma and the vulva is at 80% of the length of the body. Cruznema campestris n. sp. can be distinguished by the presence of 4 odontoplates and the arrangement of the male genital papillae, 9 pairs, of which 4 are preanal and 5 postanal.     

Monograph of the Afrotropical species of Scelio Latreille (Hymenoptera,Platygastridae), egg parasitoids of acridid grasshoppers (Orthoptera,Acrididae)

The genus Scelio is a cosmopolitan and speciose group of solitary parasitoids of the eggs of short-horned grasshoppers (Orthoptera: Acrididae). A number of these hosts are important pests, including plague locusts of the genus Schistocerca. Species of Scelio are recognized as potentially important biological control agents, but this possibility has yet to be fully realized, in part because the species-level taxonomy is still incompletely developed. The species of the pulchripennis group have been recently revised. As a continuation of this effort, here we revise the Afrotropical species of Scelio, excluding the pulchripennis species group. Sixty two (62) species are treated, 48 of which are new. Species are classified into the following species groups: ernstii (12 species, 9 new), howardi (23 species, 19 new), ipomeae (6 species, 5 new), irwini (4 species, 3 new), simoni (3 new species) and walkeri (12 species, 9 new). Keys to species groups and to the species within each group are provided. New species described are: S. albatus Yoder, sp. n., S. aphares Yoder, sp. n., S. apospastos Yoder, sp. n., S. ardelio Yoder, sp. n., S. aurantium Yoder, sp. n., S. balo Valerio & Yoder, sp. n., S. bayanga Yoder, sp. n., S. bubulo Yoder, sp. n., S. cano Yoder, sp. n., S. clypeatus Yoder, sp. n., S. concavus Yoder, sp. n., S. copelandi Yoder, sp. n., S. crepo Yoder, sp. n., S. destico Yoder, sp. n., S. dupondi Yoder, sp. n., S. effervesco Yoder, sp. n., S. erugatus Yoder, sp. n., S. exophthalmus Yoder, sp. n., S. fremo Valerio & Yoder, sp. n., S. gemo Yoder, sp. n., S. grunnio Yoder, sp. n., S. harinhalai Yoder, sp. n., S. igland Yoder, sp. n., S. impostor Yoder, sp. n., S. irwini Yoder, sp. n., S. janseni Yoder, sp. n., S. latro Yoder, sp. n., S. memorabilis Yoder, sp. n., S. modulus Yoder, sp. n., S. mutio Yoder, sp. n., S. ntchisii Yoder, sp. n., S. parkeri Yoder, sp. n., S. phaeoprora Yoder, sp. n., S. pilosilatus Yoder, sp. n., S. pipilo Yoder, sp. n., S. quasiclypeatus Yoder, sp. n., S. retifrons Yoder, sp. n., S. ructo Yoder, sp. n., S. scomma Yoder, sp. n., S. simoni Yoder, sp. n., S. simonolus Yoder, sp. n., S. somaliensis Yoder, sp. n., S. susurro Yoder, sp. n., S. tono Yoder, sp. n., S. transtrum Yoder, sp. n., S. tritus Yoder, sp. n., S. ululo Yoder, sp. n., S. vannoorti Valerio & Yoder, sp. n. The following species are redescribed: S. afer Kieffer, S. chapmani Nixon, S. howardi Crawford, S. ipomeae Risbec, stat. n., S. mauritanicus Risbec, S. philippinensis Ashmead, S. remaudierei Ferrière, S. striatus Priesner,S. taylori Nixon, and S. zolotarevskyi Ferrière. The genus Lepidoscelio Kieffer is treated as a junior synonym of Scelio Latreille, syn. n.; its type species, Lepidoscelio fuscipennis Kieffer, 1905 is transferred to Scelio, renamed Scelio obscuripennis Johnson, nom. n. (preoccupied by Scelio fuscipennis Ashmead, 1887), and redescribed. The following additional species are transferred from Lepidoscelio to Scelio: S. cayennensis (Risbec), comb. n., S. insularis Ashmead, rev. comb., S. luteus (Cameron), comb. n., S. thoracicus Ashmead, rev. comb. Lectotypes are designated for S. africanus Risbec, S. ipomeae Risbec, S. mauritanicus Risbec, S. remaudierei Ferrière, S. sudanensis Ferrière, and S. zolotarevskyi Ferrière. Scelio gaudens Nixon is a junior synonym of Scelio striatus Priesner, syn. n.; Scelio africanus Risbec and Scelio clarus Fouts are both junior synonyms of Scelio afer Kieffer, syn. n.; Scelio sudanensis Ferrière and Scelio cheops Nixon are both junior synonyms of Scelio zolotarevskyi Ferrière, syn. n.; Scelio cahirensis Priesner is a junior synonym of Scelio mauritanicus Risbec, syn. n. The name Scelio chapmanni Nixon is an incorrect original spelling, requiring an emendation to S. chapmani. Digital versions of the identification keys are available at http://www.waspweb.org/Platygastroidea/Keys/index.htm     

New species and new record of Tetranychus Dufour from Australia, with a key to the major groups in this genus based on females (Acari: Prostigmata: Tetranychidae)

A new species of spider mite, Tetranychus bunda sp. n., is described and illustrated from Australia. It was found damaging the foliage of Desmodium tortuosum (Sw.) DC. (Fabaceae) in Darwin, Northern Territory. In addition, the geographical range of Tetranychus fijiensis Hirst is extended to include Australia. This species was found in the Northern Territory feeding on frangipani ( Plumeria sp., Apocynaceae), betel palm ( Areca catechu L., Arecaceae) and Macarthur feather palm ( Ptychosperma macarthurii [H. Wendl. ex Veitch] (H. Wendl. ex Hook. f., Arecaceae)). Details of the biology of T. bunda sp. n. and T. fijiensis are given. A key to the major groups of Tetranychus Dufour of the world, based on females, is presented and species known to occur in Australia are outlined.     

New species of the genus <Emphasis Type="Italic">Parasa</Emphasis> (Lepidoptera,Limacodidae) from Southeastern Asia

Five new species of the genus Parasa Moore, 1859 from Southeastern Asia are described. At present, this genus comprises 94 species in Southeastern Asia. P. actiacus Solovyev, sp. n. is included in the lepida group, P. fidea Solovyev, sp. n. and P. anchutka Solovyev, sp. n., in the bimaculata-group, and P. pygmy Solovyev, sp. n. and P. martini Solovyev, sp. n., in the undulata group.     

Definition and Revision of the Orthrius-group of genera (Coleoptera, Cleridae, Clerinae)

An "Orthrius-group" of genera is proposed, and defined to include Aphelochroa Quedenfeldt, 1885; Caridopus Schenkling, 1908; Dozocolletus Chevrolat, 1842; Gyponyx Gorham, 1883; Languropilus Pic, 1940; Orthrius Gorham, 1876; Pieleus Pic, 1940; Xenorthrius Gorham, 1892; plus three new genera Neorthriusgen. n., Nonalatusgen. n. and Pseudoastigmusgen. n. A phylogeny of the 11 constituent Orthrius-group genera (analysis of 22 morphological characters using Clerus Geoffroy as the out-group taxon was performed with TNT v1.1) is proposed. Four genera are synonymised: Burgeonus Pic, 1950, syn. n. (with Aphelochroa Quedenfeldt, 1885); Brinckodes Winkler, 1960, syn. n. and Quasibrinckodes Winkler, 1960, syn. n. (both with Dozocolletus Chevrolat, 1842); and Dedana Fairmaire, 1888, syn. n. (with Orthrius Gorham, 1876). The genera Falsoorthrius Pic, 1940 and Mimorthrius Pic, 1940 are transferred from Clerinae to the subfamily Tillinae.     

Further contributions to the aleocharine fauna of the Yukon Territory, Canada (Coleoptera, Staphylinidae)

The aleocharine beetles of the Yukon Territory, Canada are reviewed based on material studied since the most recent survey of the territory in 2008. The present contribution recognizes a fauna of 125 species, of which 9 are new to science, 20 represent new territorial records and one represents a new Canadian record. Seventeen species are considered Holarctic, 6 introduced, and 2 species are of undetermined status (Holarctic or adventive). The Yukon fauna is classified in 32 genera and 8 tribes. The new species are: 1) Acrotona horwoodae Klimaszewski & Godin, sp. n.; 2) Atheta (Microdota) microelytrata Klimaszewski & Godin, sp. n.; 3) Atheta (Microdota) riparia Klimaszewski & Godin, sp. n.; 4) Atheta (Datomicra) whitehorsensis Klimaszewski & Godin, sp. n.; 5) Ocyusa yukonensis Klimaszewski & Godin, sp. n.; 6) Philhygra pseudolarsoniKlimaszewski & Godin, sp. n.; 7) Philhygra terrestris Klimaszewski & Godin, sp. n.; 8) Boreophilia davidgei Klimaszewski & Godin, sp. n.; and 9) Boreophilia herschelensis Klimaszewski & Godin, sp. n.