湖南零陵晚泥盆世叶虾类一新种——Echinocaris hunanensis sp. nov.

描述了湖南零陵普利桥晚泥盆世锡矿山组叶虾类一新种——Echinocaris hunanensis sp.nov..晚古生代叶虾类化石较为稀少,Echinocaris在亚洲是首次发现.     

Biostratigraphical value of some Strunian (Devonian, uppermost Famennian) Productidina, Rhynchonellida, Spiriferida, Spiriferinida brachiopods

The space of Devonian time considered in this paper corresponds to the uppermost part of the Famennian or the latest Famennian as a fourfold subdivision of the stage, usually called by many authors Strunian in neritic facies and Wocklumeria Stufe or Wocklumian in pelagic facies. Here, we examine the biostratigraphical value of certain brachiopod genera and species as bio-markers of the uppermost Famennian throughout the world (Europe, the former USSR, Middle East, Asia, North America, North Africa, and Australia). We have focused our study for species which stratigraphical range has been firmly established in correlation with the conodont biozones (Upper expansa, Early, Middle and Late praesulcata), the ammonoid biozones (do VI = “Wocklumeria Stufe”), or the foraminifera biozones (kobeitusana Zone). Other data for which the stratigraphical range is not based on standard biozonation are provided in the annex. The brachiopod bio-markers discussed in this paper belong to 14 Productidina genera (Acanthatia, Araksalosia, Ericiata, Hamlingella, Mesoplica, Nigerinoplica, Orbinaria, Ovatia, Rugauris, Semiproductus, Sentosia, Spinocarinifera, Steinhagella, Whidbornella), 7 Rhynchonellida genera (Araratella, Centrorhynchus, Hadyrhyncha, Megalopterorhynchus, Novaplatirostrum, Rozmanaria, Tchanakhtchirostrum), 13 Spiriferida genera (Brachythyris, Cyrtospirifer, Dichospirifer, Eochoristites, Imbrexia, Parallelora, Prospira, Rigauxia, Sphenospira, Tenisia, Toryniferella, Tylothyris, Voiseyella) and one Spiriferinida genus (Syringothyris). Other orders have not been studied in this paper. The main features of the uppermost Famennian brachiopod taxa represented in this paper are commented. All these taxa are listed as completely as possible throughout the world (with complements in the annex). A quick summary of the geographic distribution of the more represented taxa is given in conclusion.     

Trilobites Asteropyginae dévoniens d’Iran

Ten species of trilobites belonging to 4 genera of the sub-family Asteropyginae DELO are described from the Devonian (Givetian-Frasnian) of Iran. Three of which are new: Neocalmonia lutensis nov. sp., Neocalmonia yazdii nov. sp. and Radiopyge sardarensis nov. sp. The study of this material have required the revision of Heliopyge sharudensis(PILLET, 1974) from the Givetian of the Eastern Alborz Mountains and of Neocalmonia quadricosta PILLET, 1969, type-species of the genus Neocalmonia, from the Frasnian of Afghanistan. The recognition of genera and species in common with the Givetian and the Frasnian of Afghanistan emphasizes the close relations between these two regions. The genera Bradocryphaeus, Heliopyge, Neocalmonia, Radiopyge, represented by related species, are likewise present in Iran, Afghanistan and Western Europe. Their presence confirms that these regions belong, during the Devonian, to the North Gondwanian domain.     

Morphogenesis and evolution of Kutorgina Billings, 1861 (Brachiopoda, Kutorginida)

A comparative morphological analysis of all known members of the Lower-Middle Cambrian genus Kutorgina, one of the brachiopod genera with a carbonate shell that are oldest and richest in species. The evolution of the genus is discussed, and the borders between the Kutorgina species are emended. A new species, Kutorgina elanica sp. nov., is described.     

GENERIC HOMES FOR BRITISH SILURIAN LINGULOID BRACHIOPODS

Abstract: Since the brachiopod Order Lingulida has been greatly revised in recent years, with particular emphasis on Cambrian and Ordovician genera, many well‐established Silurian linguloid species from the British Isles have been left without an appropriate genus in which to place them. This is rectified here by allocating the various species, mostly erected in the nineteenth or early twentieth centuries, to a variety of genera which have been mainly erected within the last forty years. Two new genera are erected, Mergliella, with type species Lingula bechei Davidson, 1866 , and Striatilingula, with type species Lingula? striata J. de C. Sowerby, 1839 .     

New Devonian punctate rhynchonellids (Brachiopoda) from Transcaucasia

New genus and species of punctate rhynchonellids Sharovaella mirabilis are described from the Famennian (Upper Devonian) of Transcaucasia.     

A new delthyridoid spiriferid genus from the Dra Valley, Morocco, and its phylogenetic affinities (Brachiopoda, Lower Devonian)

Multiplicatispirifer, a new delthyridoid brachiopod genus with a gently divergent capillate micro-ornamentation and bifurcating costae on flanks, sulcus and fold, has been identified in the Mdâouer-el-Kbîr Formation of the Dra Valley (southern Anti-Atlas, Morocco). Its type speciesMultiplicatispirifer foumzguidensis n. gen., n. sp., at present the only species of this genus, is described. This form has hitherto been determined asFimbrispirifer orStruveina by various authors. In this work, the new genus is compared with the multiplicate and capillate spiriferid genera with bifurcating costae which also have costae in the sulcus and on the fold:Costispirifer andPerryspirifer from North America,Elymospirifer from South China,Borealispirifer from Asia andMultispirifer from Central Europe. The relation ofMultiplicatispirifer to the fimbriateFimbrispirifer, Struveina andVandercammenina, from which it is distinguished by its capillate micro-ornamentation, is discussed. The new genus is hitherto only known from the Moroccan southern Anti-Atlas. Its colsest relatives, taxa ofCostispirifer, occur in the Eastern American Realm (Gaspé, Canada and the eastern United States) and the Nevada Province (western United States).Multiplicatispirifer occurrences are restricted to the Lower/Upper Emsian boundary interval (upper Lower Devonian). The reasons for a possible migration corridor between North America and North Africa before Emsian time are discussed.     

Les vertebres devoniens de l'Iran central. III: Antiarches

The Upper Devonian of the Kerman region (Central Iran) has yielded but a few remains of antiarchs. Some ofthem may be referred to the almost world-wide distributed genus Asterolepis, whereas others are suggestive of the genera Byssacanthus and Stegolepis, previously recorded only from the Middle and Upper Devonian of Soviet Union.     

First steps to restructuring the problematic genus Lasiotocus Looss, 1907 (Digenea: Monorchiidae) with the proposal of four new genera

Lasiotocus Looss, 1907 is the largest genus within the Monorchiidae Odhner, 1911, with 52 species currently considered valid. Species belonging to this genus exhibit wide morphological variation and it is likely that many of them belong in other genera; however, testing the validity of the group has been hampered by a lack of molecular sequence data, especially for the type-species. Here, we report the first DNA sequences for Lasiotocus mulli (Stossich, 1883) Odhner, 1911, the type-species of the genus, and only the sixth Lasiotocus species to be sequenced. Sequences were generated for three ribosomal DNA markers, the ITS2, 18S and 28S regions, and for one mitochondrial DNA marker, the cox1 region. Phylogenetic analyses show that the six sequenced species form four clearly distinct clades, each of which we argue require separate genera. On the basis of these relationships and review of the morphology of all species in the genus, we propose a system of six genera. Four genera, Lasiotocus, the resurrected concept of Ancylocoelium Nicoll, 1912, Infundiburictus n. gen., and Sinistroporomonorchis n. gen., represent the four molecular clades. Two genera, Paralasiotocus n. gen. and Alloinfundiburictus n. gen., are proposed on the basis of morphology only. Ancylocoelium is resurrected for Ancylocoelium typicum Nicoll, 1912, Infundiburictus is erected for Lasiotocus arrhichostoma Searle, Cutmore & Cribb, 2014, Sinistroporomonorchis is erected for Lasiotocus glebulentus Overstreet, 1971and Lasiotocus lizae Liu, 2002, Paralasiotocus is erected for Lasiotocus okinawaensis Machida, 2011, and Alloinfundiburictus is erected for Lasiotocus cacuminatus Nicoll, 1915. Of the 52 species presently recognised in Lasiotocus, 43 are distributed among these genera; nine are considered species inquirendae or are transferred to other genera. We think it likely that more genera will be required for the species reviewed here but more sequence data are essential to further refine the classification.     

Biostratigraphie des thecidees (Brachiopodes) donnees recentes ou inedites

Numerous thecideid faunules have been discovered since the last years in the world. An especially stratigraphic or phylogenic interest is shown by some of these. For instance: thecideiform species from Upper Permian which are precious records on the group origin (genus FalaferGrant and AnsehiaTermier & Termier), or Upper Trias (genus HungarithecaDagis); the first triassic thecidean species, collected in Norian of Romania (Thecidella sp., Patr. & Paj.) and several species from Middle Oxfordian of Poland (genus NeothecidellaPaj., AgerinellaPatr. and RioultinaPaj.); the first species discovered in Berriasian from Crimea (Rioultina robustaSmirn.) and Algeria (Praelacazella sp., Rev. & Paj.). We can mention again some discoveries which concern different species from Afghanistan (Campanian), France (Maestrichtian), Pyrénées (Eocene) and West-Algeria (Miocene).These new data, sometimes unpublished, give precise paleoecologic (separate publication, Pajaud, 1974), phylogenic and stratigraphic informations.     

Taxonomical and biostratigraphical remarks on the rhynchonellide “Paurorhyncha” (Brachiopoda) in the Yidade Formation of eastern Yunnan,China

Additional specimens of rhynchonellide brachiopods from the marly limestones in the Yidade Formation at the Panxi section in eastern Yunnan have been ascribed to the species “Paurorhyncha” squamosa Wang, 1956 and “P.” depressa Wang, 1956. The two species have been used frequently as index fossils for the Frasnian (Upper Devonian), but their taxonomic assignments are problematic because their internal structures remain unknown. In this study, detailed systematical examinations on both external characters and internal structures revealed by serial sections suggest that the two species are more appropriately assigned to the genus Hadrotatorhynchus Sartenaer, 1986. Based on the stratigraphical distributions of Hadrotatorhynchus and the conodonts in the upper unit of the Yidade Formation, the Hadrotatorhynchus-bearing horizons are re-considered as the uppermost Givetian (Middle Devonian) rather than Frasnian stage, although the precise position of the Middle/Upper Devonian boundary still depends on further investigations of high-resolution biostratigraphy.     

Stenolaemate bryozoans from the Geirud Formation (Upper Devonian/Lower Carboniferous) of Central Alborz (Iran)

Two bryozoan species are described from the Geirud Formation (Upper Devonian/Lower Carboniferous) of Central Alborz (Iran). Trepostome Schulgina mutabilis Troizkaya, 1975 is known from the Upper Devonian (Famennian) of Central Kazakhstan. The new species Ascopora geirudensis n. sp. is the earliest known representative of the rhabdomesine genus Ascopora Trautschold, 1876.     

桂东、桂东南晚泥盆世一个弗拉斯期放射虫动物群

本文系统描述了广西玉林、贺县等地晚泥盆世榴江组硅质岩中的放射虫化石4属14种,其中有1新属2新种。这个放射虫动物群的多数分子曾见于苏联南乌拉尔地区和澳大利亚的弗拉斯阶。     

New data on the Devonian and Carboniferous Graptolithina (Dendroidea) from Belgium with notes on possible occurrences of Rhabdopleuridae in the Belgian Carboniferous

An old find of a graptolite from the Eifelian Jemelle Formation, combined with the recent discovery of an exceptionally well-preserved specimen from the Upper Frasnian (Matagne Formation), allow documentation of the geologically oldest and youngest dendroid fossils from the Devonian of Belgium. Both are ascribed to the long-ranging genus Callograptus (Acanthograptidae). These records enlarge the sparse knowledge of graptolite diversity and occurrences along the southern margin of Laurussia during the mid-Paleozoic. In Belgium, Devonian dendroids are exclusively known from the Dinant Synclinorium. The Devonian dendroid faunas previously reported from the Eifelian, Givetian and Frasnian of the Ardenne–Rhenish massifs (Belgium and Germany) are of low diversity (one to five species per locality). The specimens, generally inedaquatelly preserved, were ascribed to the genera Callograptus, Dictyonema, Palaeodictyota?, and Ruedemannograptus? In these massifs, Carboniferous dendroids are only known from the Viséan (Moliniacian) Denée Konservat-Lagerstätte. Other but undocumented reports of dendroids (and rhabdopleurids) are from the historical type area of the Tournaisian Stage (Tournai area, Brabant Parautochthon). Rhabdopleuridae from the Viséan–Serpukhovian of the Campine Basin (Turnhout borehole) remain incompletely known in the absence of the previously described material.     

Early Devonian (Lochkovian) trilobites and brachiopods from the west Balkhash Region,Kazakhstan and their significance for regional terrane dynamics

A moderately diverse Early Devonian (Lochkovian) trilobite and brachiopod association has been recovered from the lower part of the volcano-sedimentary Shakshagaily Formation, exposed on the western side of Lake Balkhash in central Kazakhstan. Its discovery demonstrates the presence of Lower Devonian marine deposits in the region. The trilobite assemblage includes representatives of six genera: the dalmanitid Kasachstania alperovichi nov. sp. is the dominant taxon, with other components being Ananaspis?, Leonaspis?, Maurotarion, Trimerus (Edgillia), and Warburgella. Associated brachiopods include species of Glossoleptaena, Leptaena, Protolepstostrophia, Pseudostrophochonetes, and Resserella. Whilst a significant proportion of these taxa show distinct links with the contemporaneous trilobite and brachiopod faunas of the north Balkhash Region, the occurrence of Trimerus (Edgillia) aff. kinglakensis Gill, 1949 suggests a possible link with the Lochkovian trilobite fauna of the Australasian sector of Gondwana. The west Balkhash Region preserves an extensive record of late Silurian to Early Devonian island arc volcanism, slope-rise sedimentation and olistostrome formation, which has invariably been ignored in models favouring early amalgamation of the Kazakhstanian terranes into a single continent by the end of the Ordovician.     

New Upper Devonian Rhynchonellids (Brachiopoda) from Transcaucasia

Anew genus, Zezinia gen. nov., and two new species, Zezinia multicostata sp. nov. and Zaigunrostrum nakhichevanense sp. nov., are described from the Upper Devonian of Transcaucasia. The first genus is one of the last Frasnian representatives of the family Uncinulidae. The Famennian Z. nakhichevanense sp. nov. belongs to punctate rhynchonellids of the family Trigonirhynchiidae.     

Cancrinella and Costatumulus (Brachiopoda) from the Permian of South Mongolia and South China: Their morphology,biostratigraphy and distribution

Two brachiopod genera, Cancrinella Fredericks and Costatumulus Waterhouse, are studied from the Permian of South Mongolia and South China. Many of the specimens previously described as Cancrinella are revised and assigned to Costatumulus. The new data presented in this paper indicate that these two genera are similar to each other, but are distinguishable by different features in relation to their morphology. Several species of both genera are described here, providing critical new information on the morphology and taxonomy of Cancrinella and Costatumulus. The reported biostratigraphical information reveals that species of Cancrinella are present from the Lower Carboniferous (Tournaisian) to Middle Permian (Capitanian), whereas species of Costatumulus are mostly restricted to the Lower Permian (Sakmarian) to Upper Permian (Changhsingian). An analysis of the palaeogeographical distribution of Cancrinella and Costatumulus reveals that Cancrinella was more commonly distributed in the Northern Transitional Zone and the Boreal Realm, in contrast to Costatumulus, which tended to be more common in the Southern Transitional Zone and Gondwanan Realm.     

Devonian (Eifelian) disparid crinoids from the Zeravashan range,southern Tien Shan,Uzbekistan

No Devonian disparid crinoids have been described from Uzbekistan, although parahexacrinid thecae and assorted columnals have been described in several papers in the past 50 years. Discovery of a cup of Pisocrinus and a thecae of Haplocrinites are the first of these genera known from Uzbekistan. The stratigraphic range of Haplocrinites is revised to be late Early Devonian (Emsian) to Early Carboniferous (Tournaisian) because Silurian species assigned to the genus are based on loose ossicles that morphologically do not belong to the genus. The paleogeographic ranges of Pisocrinus and Haplocrinites are extended into Uzbekistan. Haplocrinites uzbekistanensis n. sp. is described.     

Sphenophyllum Brongniart (Sphenopsida) from the Upper Devonian of South China

Sphenophyllum Brongniart constitutes a common and important element in the Carboniferous and Permian floras around the world, but its records in the Devonian Period are limited and remain to be investigated with respect to their morphology, anatomy, and distribution. Diversity of Sphenophyllum during Late Devonian in South China was significant with the presence of the well-known species including S. lungtanense Gothan and Sze, S. pseudotenerrimum Sze, and S. changxingense Huang et al. In this article, we describe a new species of the genus, S. fanwanense n. sp., from the Fanwan section of Changxing County, Zhejiang Province, characterized by having at least two orders of axes, with six spoon-shaped leaves attached at the node. A comprehensive review of previous records of Sphenophyllum from the Upper Devonian of South China shows that at least four species (S. fanwanense, S. lungtanense, S. pseudotenerrimum and S. changxingense), each with distinct leaf morphology, occur in South China in the Late Devonian, mostly from the Wutong Formation (Famennian). Given the lack of concrete evidence for anisophylly or heterophylly among the Devonian Sphenophyllum species (and more broadly, the Devonian sphenopsids), we suggest that the development of these features later in the Carboniferous and Permian periods was likely related to an increase in complexity of forest communities through time.     

Gastropod mollusks from the Hauterivian of Ulyanovsk (Volga Region): 2. Genera Khetella Beisel, 1977 and Cretadmete gen. nov.

The diagnosis of the genus Khetella Beisel, 1977 is emended, and a new genus, Cretadmete, is described. Two new species belonging to these genera, K. trautscholdi and C. neglecta, are established from the Upper Hauterivian of Ulyanovsk (Volga Region).