Trace fossils of the arthropod Camptophyllia from the Westphalian (Carboniferous) rocks of Lancashire, UK and their palaeoenvironmental context

Two arthropod trace fossils are described and analysed from the Carboniferous Lower Westphalian (C. communis and basal A. modiolaris chronozones) coal-bearing strata of Lancashire. The biserial trackway Diplichnites triassicus consists of five overlapping en echelon sets of 7–9 tracks preserved as epichnia and hypichnia in lacustrine siltstones. The trackway suggests subaqueous in-phase walking by a multi-segmented producer with a body length of 35–40 mm, width 17–22 mm, and 7–9 appendages. Curved, clustered, or laterally repeated, hypichnial lobes with transverse striations on the base of ripple cross-laminated sandstone are identified as Rusophycus versans. This trace fossil is interpreted as shallow resting or furrowing burrows of a homopodous arthropod, 30–60 mm long, 15–30 mm wide, and probably the same kind of arthropod as produced D. triassicus.A review of contemporary arthropod body fossils from Lagerstätten in Lancashire favours the onisciform, or Arthropleura like arthropod Camptophyllia as a potential producer of both of these trace fossils in a lacustrine palaeoenvironment.This study integrates the analysis of sediments, trace fossils and body fossils for reconstructing the arthropod biota and ecology in Westphalian lacustrine and crevasse splay fluvial palaeoenvironments.     

Refinements in the Upper Permian to Lower Jurassic stratigraphy of Karakorum, Pakistan

New sampling on critical intervals of the uppermost Permian and Triassic successions of the Northern Karakorum Terrain in the Karakorum Range (Pakistan) has refined the stratigraphy. Two types of successions may be distinguished in the Karakorum Range: a carbonate platform succession, spanning the whole interval from Upper Permian to Upper Triassic, possibly with several gaps; and a basinal succession, deposited from the Middle Permian to Early Carnian (Late Triassic), when the carbonate platform prograded into the basin. With the approaching and later docking of the Karakorum Block against the Asian margin closing the Paleo-Tethys, a portion of Karakorum emerged while another part subsided as a fore-deep, receiving clastics from the emerging Cimmerian Range. Molassic sediments filled the basin, whilst shallow-water carbonates transgressed over the emerged carbonate platform sometime between the latest Triassic and the Pliensbachian (Early Jurassic), with Cimmerian deformation occurring to the north. The age control is provided by conodonts, with assemblages of late Wuchiapingian, Changhsingian, Induan (Griesbachian and Dienerian), late Olenekian, early Anisian, late Ladinian, and early Carnian ages, respectively. Some information on the section around the P/T boundary is provided by palynology and isotopic C¹³ values. The dating of the Norian/Rhaetian platform is provided by foraminifers.     

Extremely distal fluvial sandstone within the playa system of Arnstadt Formation (Norian,Late Triassic), Central Germany

The Wachsenburg Sandstone of Thuringia (Central Germany) occurs within playa deposits of the Arnstadt Formation (Late Triassic, Norian) and furnishes an example of ephemeral river metamorphosis under dryland conditions. Characterized by high flow regime features, the sand-dominated lithofacies constitution exhibits sedimentation by channel processes under the influence of recurring flash floods. Bearing signatures of subaerial exposures, the fining-upward lithofacies cycles are bound by low-angle lateral accretion elements suggesting deposition in a meandering stream. Channel migration in response to point bar expansion and active bank erosion, led to the development of four laterally shifting point bar events. Unimodal palaeocurrent patterns with low variance and azimuthal dispersion support the point bar origin of the Wachsenburg Sandstone. With reduced water budget under largely semi-arid climate, the river progressively became smaller, highly sinuous and ultimately abandoned. The resulting point bar succession was finally covered with sheet flow deposits of over bank origin. The sandstone was deposited during a period of low base-level when the playa system temporarily fell dry.     

Indications for a humid climate in the Western Desert of Egypt 11–10 Myr ago: evidence from Galagidae (Primates, Mammalia)

The timing of desertification of the Sahara Desert is poorly understood, with recent estimates indicating an onset of hyper-aridity during the Latest Miocene. Field work in Egypt in 2005 has led to the discovery of evidence that indicates that 11–10 Ma the Western Desert was covered in woodland. Fossiliferous cave breccia at Sheikh Abdallah, Western Desert, Egypt, has yielded a Late Miocene (11–10 Ma) microvertebrate fauna, which contains Galagidae, Microchiroptera, Macroscelididae, Soricidae, Erinaceidae, and Rodentia. The locality also yielded the remains of frogs, snakes, lizards, and birds. The fauna indicates a mean annual rainfall in excess of 500 mm and perhaps as much as 1,200 mm. This palaeoclimatic information is important because it reveals that the Sahara Desert, which is today the largest in the world, was either considerably smaller during the Late Miocene than it is today, or that it did not yet exist as a continuous hyper-arid belt right across the continent. This data accords with estimates of a Latest Miocene (8–7 Ma) increase in aridity in the Sahara. To cite this article: M. Pickford et al., C. R. Palevol 5 (2006).     

Late Pleistocene and Holocene environments of the Southeastern United States from the stratigraphy and pollen content of a peat deposit on the Georgia Coastal Plain

A 5.4 m peat core from the Sandy Run Creek (SRC) valley in the upper Coastal Plain of Georgia shows that before 30 ka the valley had an aggradational sandy floor with shallow swales and low ridges of 1–2 m amplitude suggesting a braided stream pattern and a low water table. The climate from 30–25 ka was cool and dry and the vegetation open grassland with stands of pine and spruce that produced few fires. At ca. 16 ka a warmer, wetter climate caused SRC to meander and incise the valley fill removing some previously deposited sediment at the site of our peat core so that sediments of 13 ka rest directly on sediments deposited at 25 ka. After ca. 13 ka higher groundwater levels initiated vertical accretion of floodplains allowing peat accumulation in shallow depressions across the valley. Pollen from the Younger Dryas period (ca. 13–11 ka) indicates a cool, moist environment of open oak woodland, mesic trees, riparian populations of alder, and reduced levels of pine. By the early to mid Holocene, tupelo and oak dominated over pine indicating wetter conditions, while sand stringers in peat record periodic heavy rains, an elevated water table, and intervals of substantial runoff. Cooling after ca. 4.5 ka brought drier and more variable conditions. Fires increased and southern pine replaced tupelo and oak. The absence of sand stringers in peat younger than 4.5 ka indicates fewer intense rainfall events.     

Lacustrine sedimentation in the diapir-controlled Miocene Bicorb Basin, eastern Spain

The Miocene Bicorb Basin is a small elongated basin developed by normal faulting of a thick Jurassic–Cretaceous carbonate succession and subsequent diapirism of Upper Triassic mudstones and evaporites. The basin fill comprises a sequence over 650 m thick formed by two units. The lower, alluvial unit consists of a complex alternation of conglomerates, sandstones, mudstones, and minor lacustrine limestones. The upper unit comprises marginal alluvial and lacustrine deposits in which seven lacustrine facies associations have been distinguished. The inner lacustrine deposits comprise mudstones and carbonates with minor evaporitic deposits. At the northeast and southwest basin edges, alluvial inputs led to the development of delta and fan-delta environments where sandstone and conglomerate deposits dominate. The latter stages of the lake evolution are represented by an expansive thick limestone sequence which close to the NW fault-bounded margin pass laterally to breccias. The hierarchical arrangement of the upper unit shows five different orders of sequences. The first-order sequence defines a major vertical trend of lake expansion whereas three second-order sequences are linked to major flooding-expansion episodes. All these sequences are related to the tectonic evolution of the diapir. Metre-scale, third- and fourth-order sequences and fifth-order sequences, which consist of finely laminated rhythmite deposits probably record climatically forced processes. The diapir–graben system evolution and the climatic variations have exerted major controls on the sequential arrangement and evolution of the lacustrine system. Both factors have also strongly influenced the changes in the gastropod assemblages.     

Integrated stratigraphy of the Early Miocene lacustrine deposits of Pag Island (SW Croatia): Palaeovegetation and environmental changes in the Dinaride Lake System

An integrated stratigraphic study of a Neogene lacustrine succession on the Pag Island (Croatia), combining quantitative pollen analysis, magnetostratigraphy, cyclostratigraphy, biostratigraphy and gamma-ray measurements, provides new insights into orbitally controlled variations in palaeo-vegetation and depositional patterns in the Dinaride Lake System. The quantitative palynological record shows a cyclical pattern of vegetation changes that closely corresponds to sedimentological patterns. The intervals with a high abundance of thermophilous and xeric indicators, suggesting a warm and dry climate, generally coincide with intervals of frequent lignite deposition and shallow lake facies. This suggests that both records are dominantly controlled by variations in past climatic conditions and lake level. Our data show two large-scale warming and shallowing-upward cycles, which are interpreted to be forced by the ~ 100 kyr eccentricity cycle of the Earth's orbit. Magnetostratigraphic data of the examined section reveal a long (113 m) reversed polarity interval, followed by a 7 m thick interval of normal polarity at the top. The inferred depositional rate of ~ 0.3 mm/yr, combined with biostratigraphic constraints by mollusks, suggests that the most logical correlation of the reversed interval is to chron C5Cr. This indicates that the Pag succession was deposited between 17.1 and 16.7 Ma and that it corresponds to the Burdigalian Stage of the Early Miocene, and the regional Karpatian Stage of the Central Paratethys. The high relative percentage of thermophilous pollen taxa, Engelhardia and Taxodium-type being the most prominent, generally indicates a subtropical humid climate for the SW Croatian part of the Dinaride Lake System. The observed warming trend is possibly related to the onset of the Miocene Climatic Optimum.     

Improved purification of the membrane-bound hydrogenase–sulfur-reductase complex from thermophilic archaea using ε-aminocaproic acid-containing chromatography buffers

A hydrogenase–sulfur reductase (SR) complex was purified from membrane preparations of the extremely thermophilic, acidophilic archaeon Acidianus ambivalens using a combination of sucrose density gradient centrifugation and column chromatography (FPLC). All chromatographic steps were performed in the presence of 0.5% ε-aminocaproic acid resulting in the elution of the SR complex as a sharp peak. In contrast, chromatography using buffers without ε-aminocaproic acid, or in the presence of detergents, were not successful. The purified A. ambivalens SR complex consisted of at least four subunits with relative molecular masses of 110 000, 66 000, 39 000 and 29 000, respectively. A similar procedure was applied to purify the membrane-bound hydrogenase from Thermoproteus neutrophilus, a non-related extremely thermophilic but neutrophilic archaeon, which consisted of only two subunits with relative molecular masses of 66 000 and 39 000, respectively.     

The temnospondyl amphibian Cyclotosaurus from the Upper Triassic of Poland

Abstract:  A gap in the Late Triassic fossil record of the capitosaur amphibian Cyclotosaurus is filled by new material from lacustrine deposits at Krasiejów, Poland, corresponding in age to the Lehrberg Beds (late Carnian) of Germany. The skull of the Polish cyclotosaur is intermediate in several respects between that of Cyclotosaurus robustus from the middle Carnian Schilfsandstein of Germany and the younger C. mordax from the early Norian Stubensandstein. It shows a decrease in the width of the skull and in the degree of concavity of the posterior margin of the skull roof. The differences are significant enough to warrant erection of a novel species, the name Cyclotosaurus intermedius sp. nov. being proposed. The pectoral girdle, identified for the first time in Cyclotosaurus , suggests the genus was more fully adapted to an aquatic mode of life than was Paracyclotosaurus .     

Utilisation préhistorique de la technique minière d’abattage au feu dans le district cuprifère de Cabrières (Hérault)Prehistoric use of the fire-setting method in the copper-mining district of Cabrières (Hérault, France).

Fire-setting to open up mines has been used on hard rock since prehistoric times. In the copper-mining district of Cabrières, the existence of metre-sized spherical or sub-spherical cavities, sometimes spaced along the same vertical in an ore seam, has usually been ascribed to this method 12; 13 and 20. Two AMS ¹⁴C dating of micro-charcoal found in dolomite and of burnt ore breccias related to extraction in these cavities give the ages 3830 ± 40 BP, cal BC 2340–2130 and 3900 ± 40 BP, cal BC 2480–2280, which is the first evidence in France of the use of fire-setting in prehistoric mines.

Résumé

L’abattage, ou l’ouverture des mines, par le feu, a été pratiqué dans le cas de roches dures depuis la Préhistoire. Dans le district minier cuprifère de Cabrières, l’existence de cavités sphériques ou sub-sphériques métriques, parfois échelonnées sur une même verticale le long des filons minéralisés, a été classiquement attribuée à cette technique 12; 13 and 20. Deux datations ¹⁴C par AMS des micro-charbons de bois récoltés dans les brèches de dolomie et de minerais brûlés liés au dépilage minier de ces cavités ont donné les âges de 3830 ± 40 BP, cal BC 2340–2130 et de 3900 ± 40 BP, cal BC 2480–2280, confirmant, pour la première fois en France, l’utilisation dans les mines préhistoriques de l’abattage au feu.     

Taxonomical and palaeoecological investigations of the chondrichthyan and osteichthyan fish remains from the Middle-Late Triassic deposits of the Villány Hills (Southern Hungary)

Two sites in the Villány Hills, Hungary, have yielded rich fish assemblages from Middle to Late Triassic shallow marine deposits. The collected material comes from the Ladinian Templomhegy Dolomite Member and from the Carnian Mészhegy Sandstone Formation. The ichthyofauna is composed of both chondrichthyans (Hybodontidae indet., Palaeobates angustissimus, ‘Polyacrodus’ sp., ?Lissodus sp.) and osteichthyans (Gyrolepis sp., Birgeria sp., and further indeterminate actinopterygians). Despite the large sample size, no remains of neoselachians have been found. The Ladinian Templomhegy Dolomite is dominated by durophagous hybodontiforms (Palaeobates angustissimus, ?Lissodus sp.), but the piscivorous hybodontid and the generalist ‘Polyacrodus’ sp. are missing, while in the fish fauna collected from the Carnian Mészhegy Formation indeterminate piscivorous hybodontids are the most common elements and durophagous forms are much less abundant. The dominance of piscivorous hybodontids in the Carnian Mészhegy Sandstone could be related to the global decrease of diversity of marine fish-eating reptiles (e.g., nothosaurs) or to a change of paleoenvironmental conditions. The present study improves our knowledge on the poorly known Triassic vertebrate faunas of the Tisza Mega-unit, which formed a segment of the passive Neotethys margin of the European Plate and shows an important example of a potential vertebrate faunal shift during the Middle to Late Triassic.     

Chirothere Footprint Sites from the Otter Sandstone Formation (Middle Triassic,late Anisian) of Devon,United Kingdom

Three chirothere footprint sites are documented from the Middle Triassic (Anisian) fluvial Otter Sandstone Formation of Sidmouth, Devon, UK. One site, on the foreshore below Peak Hill, has revealed numerous tracks, including a probable manus imprint, which are recorded here. Another site, also beneath Peak Hill, has produced a single probable manus track from a higher horizon. The third site, on the foreshore below Salcombe Hill Cliff, has yielded a small number of pes tracks similar to those from Peak Hill. The footprints at all sites are interpreted as belonging to a large Chirotherium ichnospecies, perhaps C. barthii Kaup. Unusually, the tracks are preserved in mudstone, often in convex epirelief, and are inferred to have been generated subaqueously. The apparent restriction of the footprints to the higher part of the Otter Sandstone Formation supports other evidence, such as a greater abundance of vertebrate skeletal material, subhorizontal rather than vertical rhizocretions, and a greater frequency of lacustrine facies, for a less arid climate and higher water table than lower in the sequence.     

Palynological dating of the Alturaia Arkose (Balagne, northern Corsica): geological implications

In Alpine Corsica, the Balagne Nappe displays the best-developed sedimentary succession associated with an ophiolite sequence. This sedimentary succession includes the Alturaia Arkose, whose age is still unknown. Several shale horizons cropping out in the Cima di Alturaia area were studied for palynological analyses. In this paper, a new palaeontological find in the Alturaia Arkose is reported and the related geological implications are discussed. The collected data indicate the occurrence of a palynological assemblage of Late Barremian to Middle Aptian age. The Alturaia Arkose can be regarded as a clastic deposit of Late Barremian–Middle Aptian age derived from rocks cropping out in Hercynian Corsica. To cite this article: M. Marroni et al., C. R. Palevol 3 (2004).

Résumé

Datation palynologique de l’arkose de l’Alturaia (Balagne, Corse septentrionale) : conséquences géologiques. En Corse alpine, la nappe de Balagne montre la meilleure succession sédimentaire associée à une séquence ophiolitique. Cette succession inclut l’arkose de l’Alturaia, dont l’âge est encore inconnu. Plusieurs horizons de shales ont été étudiés en vue d’analyses palynologiques, dans la zone de la Cima di l’Alturaia. Nous y indiquons une découverte paléontologique, et nous en discutons les implications géologiques. Les données nouvelles montrent la présence d’un assemblage palynologique d’âge Barrémien supérieur–Aptien moyen. L’arkose de l’Alturaia peut ainsi être considérée comme un dépôt détritique de cet âge, alimenté par les roches affleurant dans la Corse hercynienne. Pour citer cet article : M. Marroni et al., C. R. Palevol 3 (2004).     

Characteristics of peritidal facies and evidences for subaerial exposures in Dachstein-type cyclic platform carbonates in the Transdanubian Range,Hungary

In the Late Triassic, an extremely large carbonate platform system (Dachstein-type platforms) developed on the margin of the Neotethys. On the wide inner platform cyclic peritidal, lagoonal successions were deposited. In the Transdanubian Range (Hungary), the lower part of the 1.5–2-km-thick cyclic succession (Upper Tuvalian–mid-Norian) is pervasively dolomitised, the upper part (Upper Norian–Rhaetian) is non-dolomitised; there is a transitional interval between them made up of partially dolomitised cycles. The peritidal–lagoonal cycles are commonly bounded by well-developed disconformity surfaces reflecting subaerial erosion that punctuated the marine carbonate accumulation. Truncation of the cycles was preceded by pervasive cementation of the previously deposited cycle. In the early stage of the platform evolution, tidal flat dolomitisation under semi-arid conditions led to the consolidation of the previously deposited sediments. The truncation surfaces were commonly covered by dolocretes. During the more humid Late Norian–Rhaetian period, the early cementation was followed by karstification, accumulation of wind-blown dust and pedogenesis. Erosion during regularly recurring subaerial exposure that commonly reached the previously deposited subtidal beds suggests eustatic control of the cyclicity and supports the application of an allocyclic model, even if the Milankovitch signal is imperfect.     

Neritic carbonate for six submerged coral reefs from northern Australia: Implications for Holocene global carbon dioxide

Total contribution of six recently discovered submerged coral reefs in northern Australia to Holocene neritic CaCO₃ and CO₂ is assessed to address a gap in global budgets. CaCO₃ production for the reef framework and inter-reefal deposits is 0.26–0.28 Mt. Holocene CO₂ production is 0.14–0.16 Mt. Coral and coralline algae are the dominant sources of Holocene CaCO₃ although foraminifers and molluscs are the dominant constituents of inter-reefal deposits. The total amount of Holocene neritic CaCO₃ produced by the six submerged coral reefs is several orders of magnitude smaller than that calculated using accepted CaCO₃ production values because of very low production, a ‘give-up’ growth history, and presumed significant dissolution and exports. The contribution of submerged coral reefs to global Holocene neritic CaCO₃ is estimated to be 0.26–0.62 Gt, which yields 0.15–0.37 Gt of CO₂. This amount of CO₂ is 0.02–0.05% of the 780 Gt added to the atmosphere since 18 kyr BP. Contributions from Australian submerged coral reefs are estimated to be 0.05 Gt CaCO₃ and 0.03 Gt CO₂ for an emergent reef area of 47.9 × 10³ km². Based on the growth history of the submerged coral reefs in the Gulf of Carpentaria, maximum global Holocene CaCO₃ fluxes could have attained 0.3 Gt yr^− 1 between 11 and 7 ka BP. This additional CaCO₃ would have culminated in a maximum CaCO₃ production from all (emergent and submerged) coral reefs of 1.2 Gt yr^− 1 and neritic CaCO₃ production of 2.75 Gt yr^− 1. The dilemma remains that the global area and CaCO₃ mass of submerged coral reefs are currently unknown. It is inevitable that many more submerged coral reefs will be found. Our findings imply that submerged coral reefs are a small but fundamental source of Holocene neritic CaCO₃ and CO₂.     

A Re-evaluation of Small Tetrapods from the Middle Triassic Otter Sandstone Formation of Devon, England

Material of small sauropsids from the Otter Sandstone Formation of east Devon (Sherwood Sandstone Group; Middle Triassic; Anisian) includes remains that were formerly attributed to a primitive procolophonid. In the light of new specimens, this material is instead found to contain remains of a diapsid and a procolophonine procolophonid. Among these fossils, the medium-sized procolophonine, Kapes bentoni sp. nov., is the first record of this Russian genus in the British Triassic. Coartaredens isaaci gen. et sp. nov. is a small diapsid tentatively assigned to Lepidosauromorpha. The heterodont lower dentition of Coartaredens comprises a row of large, conical posterior teeth and tightly packed, procumbent incisiforms. Two additional specimens are distinguished on the basis of distinctive dentary remains. One of these is of possible procolophonid affinity, while the dentition of the second resembles that of the aberrant Early Triassic parareptilian genus Sclerosaurus .     

Invertébrés remarquables du Callovien inférieur de la Voulte-sur-Rhône (Ardèche, France)Remarkable Invertebrates from the Lower Callovian of la Voulte-sur-Rhône (Ardèche, France)

Remarkable Invertebrates from the Lower Callovian of la Voulte-sur-Rhône (Ardèche, France). The four-excavating field expeditions from 1983 to 1986 led to collect a considerable amount of new data from this exceptional paleontological site, which was neglected, and unexplored for the essential, until now. This paper presents a synthesis of the main available results. For the first time, the succession of the lithofacies is precised from all preserved deposits thick about 12 m. These observations reveal, on the whole, a great homogeneity of lithologic, paleobiologic or taphonomic characteristics, which denote a rather great constancy of the global environmental conditions. The lithofacies are schistous marls revealing former uncemented paleosubstrates, with clay very dominating (montmorillonite for 60–70%, calcium carbonate, siderite and limonite for less than 30% or 40%). The faunas are nearly exclusively composed of pelagic, nectopelagic or mesopelagic organisms (swimming in full water or dependent on free algal environments); they include very few benthic organisms. The marine water depth in this area would be estimated to have reached roughly 200 m. The surface and middle water strata, where lived the major part of organisms, must have been usually agitated, well oxygenated and illuminated. At the opposite side, the bottom water must have been very quiet and anoxic, in aphotic zona, which is responsible for the soft tissue mineralisation.     

Altered regulation of renal sodium transporters and natriuretic peptide system in DOCA–salt hypertensive rats

Although deoxycorticosterone acetate (DOCA)–salt hypertension is a volume dependent model of hypertension, it shows polyuria and natriuresis. It is expected that dysregulation of aquaporin water channels (AQPs) and sodium transporters associated with natriuretic peptide (NP) system may play an escape role in sodium retaining state. One week after left unilateral nephrectomy, rats were subcutaneously implanted with silastic DOCA (200 mg/kg) strips. Physiologic saline was supplied as a drinking water to all animals. 4 weeks after operation, the protein expression of AQPs, sodium transporters, and endopeptidase (NEP) was determined in the kidneys by semiquantitative immunoblotting and immunohistochemistry. The mRNA expression of NP system was determined by real-time polymerase chain reaction. The amount of urinary ANP excretion was measured by radioimmunoassay. In DOCA–salt rats, urine osmolality was decreased while urinary excretion of sodium was increased. The expression of AQP1-3 as well as that of α-1 subunit of Na,K–ATPase, NHE3, NKCC2 and NCC was decreased in the kidney. The mRNA expression of ANP, brain natriuretic peptide (BNP), C-type natriuretic peptide (CNP) was increased in the kidney. The expression of NEP was decreased, and urinary ANP excretion was increased. Downregulation of AQPs and sodium transporters may contribute to mineralocorticoid escape in DOCA–salt hypertension. Increased expression of natriuretic peptides associated with downregulation of NEP may play a role in natriuresis.