Two new genera of cuckoo wasps (Chrysididae: Amiseginae) from Rovno and Baltic ambers

Cuckoo wasps of the subfamily Amiseginae, Eosega heterotarsata gen. et sp. nov. and Foveorisus kilimniki gen. et sp. nov., from the Late Eocene Baltic (Russia) and Rovno ambers (Ukraine), respectively, are described. Primitive Eosega is close to the extinct genus Palaeobethylus Brues in Kieffer, 1914 and represents the so-called American series of genera of Amiseginae. Foveorisus is found to be morphologically close to a number of extant oriental genera. All known fossil Amiseginae are listed. Stick insects of the family Archipseudophasmatidae, possible hosts of the Eocene amisegines, are recorded in the Rovno amber for the first time.     

Gall midges of the supertribe stomatosematidi (Diptera,Cecidomyiidae) in the Palaearctic region,with descriptions of new taxa from the late Eocene Rovno Amber. 2. The genera <Emphasis Type="Italic">Didactylomyia</Emphasis> and <Emphasis Type="Italic">Rovnodidactylomyia</Emphasis> gen. n.

One new genus (Rovnodidactylomyia Fedotova et Perkovsky, gen. n.) and four new species (Didactylomyia dlusskyi sp. n., Rovnodidactylomyia zosimovichi gen. n. et sp. n., R. sidorenkoi sp. n., and R. iconica sp. n.) are described from an amber sample of the late Eocene Period (Rovno District, Ukraine). Two species from a Baltic amber sample of the late Eocene Period, described earlier in the genus Bryocrypta, are attributed to the genus Rovnodidactylomyia (R. girafa (Meunier, 1904) comb. n. and R. capitosa (Meunier, 1904)). The first fossil species of the genus Didactylomyia is described for the first time; two contemporary Palaearctic species are recorded. Keys to the species of the genus Didactylomyia are given.     

Ants of the Genus <Emphasis Type="Italic">Fallomyrma</Emphasis> Dlussky et Radchenko (Hymenoptera,Formicidae, Myrmicinae) from Late Eocene European Ambers

The extinct monotypic ant genus Fallomyrma was described by Dlussky and Radchenko in 2006 based on workers from Late Eocene Rovno (Ukraine), Scandinavian (Denmark), and Bitterfeld (Germany) ambers. Three new Fallomyrma species from the Rovno amber are here described based on workers: F. anodonta sp. nov., F. marginata sp. nov., and F. robusta sp. nov. A key to all known species of this genus is compiled. The quantity ratios (of the numbers of species and specimens) of Fallomyrma and other amber Myrmicinae genera are considered. The opinion of the autochthonous origin of the Rovno amber is confirmed.     

A new species of derelomine weevils (Coleoptera,Curculionidae, Curculioninae: Acalyptini) from the Rovno amber

Protoceletes hirtus sp. nov. (Curculionidae: Curulioninae: Acalyptini) from the Late Eocene Rovno amber is described. This is the second fossil of the genus. The new species is close to P. wolfschwenningerae Rheinheimer, 2007 from Baltic amber and differs it in the separated coxae, asymmetrically convex eyes, longer and clearly semierect hairs on the surface, and the longer body.     

A new species of Micromalthidae (Coleoptera) from the Rovno amber: 1. Adult morphology

Micromalthus priabonicus sp. nov. from the Late Eocene Rovno amber (Ukraine) is described. The new species is readily distinguished from M. eocenicus Kirejtshuk et al., 2010 from the Early Eocene amber of Oise in France by the shorter body, unequal length of antennomeres 3 and 4, ovate terminal antennomere, less transverse head, transverse scutellum, and by the considerably shortened adsutural elytral margin.     

A new caddisfly of the fossil genus <Emphasis Type="Italic">Archaeotinodes</Emphasis> (Insecta: Trichoptera: Ecnomidae) from the Baltic amber

A new caddisfly species of the fossil genus Archaeotinodes, A. igneusaper sp. nov., is described from the Upper Eocene Baltic amber. The new species is close to A. pauper Ulmer, 1912 and A. lanceolata Ulmer, 1912 from the Baltic amber, differing from these in the structure of the male genitalia.     

New long-horned caddisflies in Eocene Baltic amber (Insecta,Trichoptera)

We describe two new extinct caddisflies, Electroadicella kuenowi sp. nov. and Triaenodes simoni sp. nov., embedded in Baltic amber. Both species belong to the tribe Triaenodini and enlarge the family Leptoceridae to a remarkable number of 18 extinct Eocene species from Baltic amber. The extinct Electroadicella and the extant Triaenodes are possibly closely related, by a distinct synapomorphy, a recurved process from the basal plate of the inferior appendages of the ninth abdominal segment in the male genitalia.     

New taxa of soft-winged flower beetles (Coleoptera,Malachiidae) in Baltic and Rovno amber

Two new genera and three new species of soft-winged flower beetles (Coleoptera, Malachiidae) are described from Rovno and Baltic Eocene amber. Protocephaloncus gen. nov. with the type species P. perkovskyi sp. nov. was found in the Rovno amber. It differs from its congeners in the tribe Troglopini in the primitive sculpture of the male facial area as a longitudinal narrow depression between the eyes lacking an appendage inside, and two trapezoid impressions on both sides of the depression closer to its lower part. A transverse pronotum, slightly rounded, but not sinuate posteriorly, is also a diagnostic feature of the new genus. A new genus of the tribe Attalini, Palpattalus gen. nov., was found in the Baltic amber. The tarsal comb on the anterior second tarsomere of the male, complicatedly curved pygidium, and cylindrical apical palpomere allow the definite attribution of the genus to the tribe Attalini. The characteristically enlarged, rounded and slightly flattened 1st palpomere, possessing a long seta directed outward, of the new genus resembles similar structures found in members of the tribe Colotini, all males of which have modified and enlarged 3rd and 4th palpomeres. Probably, Palpattalus gen. nov. represents an ancestral form for these two tribes. This allows the establishment of independent status for Antholinus Mulsant and Rey, 1867 stat. resurr., with anterior tarsi in males lacking comb on the second tarsomere, in contrast to all other subgenera of Attalus Erichson, 1840. Males of two different species, Palpattalus baltiensis sp. nov. and P. eocenicus sp. nov., were found as inclusions in Baltic amber. They differ in the coloration of the elytra, yellow with black spots in Palpattalus baltiensis sp. nov. and unicolorous black in P. eocenicus sp. nov.; the shape of pygidia, longitudinal and strongly elongated in P. baltiensis and transverse and rounded in P. eocenicus, and in a number of other characters.     

<Emphasis Type="Italic">Rovnodiplosis eduardi</Emphasis> gen. et sp. nov., the first record of a fossil gall midge of the supertribe Mycodiplosidi (Diptera,Cecidomyioidea, Cecidomyiidae) in the Late Eocene of the Rovno amber

Higher gall midges of the supertribe Mycodiplosidi, subfamily Cecidomyiinae, are recorded for the first time in Late Eocene Rovno amber. In other fossil resins, Mycodiplosidi have not yet been recorded. A male Rovnodiplosis eduardi gen. et sp. nov. (Mycodiplosidi), with distinctive binodose flagellomeres having long loopy sensilla is described. Species of Cecidomyiinae known from fossil resins are listed.     

Hepatics from Rovno amber (Ukraine), 6. Frullania rovnoi,sp. nov.

A fossil species of the extant liverwort genus Frullania Raddi is described and illustrated, based on a single inclusion in a piece of Rovno amber (Ukraine) that shares its age with Late Eocene Baltic amber, its northern contemporary. Frullania rovnoi is characterised by leaves with a rounded dorsal lobe and the absence of ocelli. The ventral lobe is inflated and forms a saclike lobule, which is bell-shaped and somewhat constricted above the mouth. The bifid underleaves have several blunt teeth or angulations along the shoulder. The Rovno fossil differs sufficiently from morphologically similar species preserved in Baltic and Bitterfeld amber as to be described as new to science. The shape of the lobules and underleaves, as well as the absence of ocelli, indicate an affiliation to F. sect. Australes, hitherto represented in Eocene amber inclusions solely by F. schumannii (Casp.) Grolle. The Rovno fossil is distinguished from extant species of F. subg. Australes and from F. schumannii by having roughly and irregularly dentate-angulate underleaf margins.     

New species of caddisflies (Insecta: Trichoptera) from the Paleogene resins of Europe

Two new species of caddisflies of the families Philopotamidae and Polycentropodidae, Wormaldia pobeda sp. nov. and Nyctiophylax leningrad sp. nov. are described from the Eocene Baltic and Rovno ambers.     

Have ladybird beetles and whiteflies co-existed for at least 40 Mya?

The beetle family Coccinellidae is rarely recorded from fossils. Most of the records come from the nineteenth and beginning of the twentieth century. A complete list of Coccinellidae records from Baltic amber is presented and discussed. Extensive literature research provided a surprising conclusion that not a single species of Coccinellidae has been formally described from Baltic amber till now. The first two species of ladybird beetles from Eocene Baltic amber are described and placed in the genus Serangium, namely S. twardowskii sp. nov. and S. gedanicum sp. nov. Their phylogenetic placement in the subfamily Microweiseinae is provided. A key to the fossil species of Serangium is given. Extant representatives of the genus are distributed mainly in tropical areas of Asia and Oceania, and are specialised predators of whiteflies. Current discovery shows that during the Middle Eocene, the genus Serangium was distributed wider in the Northern Hemisphere and the evolution of these ladybird beetles was probably influenced by the evolution of whiteflies which are also found in the Baltic amber.     

A new genus and species of gall midges of the supertribe Heteropezidi (Diptera,Cecidomyiidae) found in Eocene amber from Sakhalin

A new genus and species of pedogenetic gall midge, Estoperpetua sakhalinica gen. et sp. nov., of the tribe Heteropezini (subfamily Lasiopterinae) is described. Gall midges have not been recorded in Eocene amber of Sakhalin. A female with 4-segmented tarsi, large transparent triangular sensoria on flagellomeres, short wings and legs, and long 2 + 9-segmented antennae is described. Heteropezidi shows the greatest generic and species diversity in the Late Eocene Rovno amber. Representatives of this supertribe were also found in the Late Eocene Baltic amber, Early Eocene Oise amber, Miocene amber from Mexico and Dominican Republic, and African copal. Heteropezidi from different amber faunas are reviewed.     

Gall midges of the supertribe Stomatosematidi (Diptera,Cecidomyiidae) in the Palaearctic Region,with descriptions of new taxa from the Late Eocene Rovno amber: 1. <Emphasis Type="Italic">Stomatosema,Vanchidiplosis, Clarumreddera</Emphasis> gen. n.

Three recent genera are included in the supertribe Stomatosematidi: Stomatosema (16 species), Vanchidiplosis (4 species), and Didactylomyia (3 species). Seven species of the genus Stomatosema and two species of the genus Vanchidiplosis occur in the Palaearctic Region. No reliable findings of representatives of the supertribe were known in fossilized condition. One new genus and 3 new species are described from amber of the Late Eocene age (Rovno Region): Stomatosema iljieteugeniae sp. n., Clarumreddera korneyevi gen. n. et sp. n., and C. conceptiva sp. n.     

A new species of the genus <Emphasis Type="Italic">Nalassus</Emphasis> Mulsant (Coleoptera: Tenebrionidae: Helopini) from the Baltic amber

The first Eocene species of the tribe Helopini, Nalassus klebsi sp. nov., from the Baltic amber is described. The new species is included in the nominative subgenus and similar to the extant species N. dermestoides (Illiger) and N. laevioctostriatus (Goeze), from which it differs in the shape of eyes (eyes are considerably less transverse than in living Palearctic species of the genus Nalassus Mulsant), the shape of the pronotum with acute posterior angles (other Palearctic Nalassus have obtuse or straight pronotal angles) as well as convex elytral intervals. Species of the less specialized nominative subgenus are widespread in forests and (partly) alpine landscapes of Europe (eastwards up to the Urals) and eastern Asia.     

<Emphasis Type="Italic">Dencyrtus</Emphasis> gen. nov. (Hymenoptera,Chalcidoidea, Encyrtidae) from the Late Eocene Danish Amber

Based on two females from the Danish amber collection of the Zoological Museum of University of Copenhagen, Dencyrtus vilhelmseni gen. and sp. nov. is described. The antennae of the new genus are very similar in structure to that of Eocencyrtus Simutnik, 2001 from the Rovno and Baltic ambers. However, the presence of the filum spinosum on the linea calva of the forewing disc indicates that it belongs to a different subfamily.     

On the taxonomic position of <Emphasis Type="Italic">Issus reticulatus</Emphasis> Bervoets, 1910 (Hemiptera: Fulgoroidea: Issidae) from Baltic amber

A new genus Bolbossus gen. n. is erected in the tribe Parahiraciini for Issus bervoetsi Gnezdilov et Bourgoin, nom. n. pro Issus reticulatus Bervoets, 1910 (nom. praeocc., non Issus reticulatus Herrich-Schäffer, 1835) described from the Baltic amber of Kaliningrad Province in Russia. The genus Issites Haupt, 1956 is assigned to the nominotypical subtribe of the tribe Issini Spinola. Bolbossus bervoetsi (Gnezdilov et Bourgoin) is the first species of the Oriental tribe of the family Issidae discovered in Europe. This finding implies the dispersal of taxa from the Oriental Region to the Western Palaearctic during the Eocene. Other fossil Issidae are briefly discussed.     

On finding a single-clawed aphid, <Emphasis Type="Italic">Germaraphis ungulata</Emphasis> (Homoptera,Aphidinea), in a syninclusion with the ant <Emphasis Type="Italic">Monomorium mayrianum</Emphasis> (Hymenoptera,Formicidae) in the Saxonian amber

The first syninclusion of the myrmicine Monomorium mayrianum Wheeler (two workers) and the aphid Germaraphis ungulata Heie (one nymph) is described from the Bitterfeld amber. Additionally, G. ungulata is recorded in a syninclusion with Ctenobethylus goepperti (Mayr) in the Rovno amber for the first time.     

First Record of the Late Eocene Ichneumon Fly <Emphasis Type="Italic">Rasnitsynites tarsalis</Emphasis> Kasparyan (Ichneumonidae,Townesitinae) in Ukraine Confirms Correlation of the Upper Eocene Lagerstätten

Rasnitsynites tarsalis Kasparyan (Ichneumonidae, Townesitinae) described from the Baltic amber is recorded for the first time in the Rovno amber (Gulyanka, Zhitomir Region); therefore, this is the southernmost and easternmost find of Townesitinae. The presence of numerous extinct taxa of Townesitinae and tribes of Hybrizoninae Blanchard 1845 = Paxylommatinae Förster, 1862 = Hybrizontinae Marshall, 1872 confirms close relation of the Bembridge Marls fauna (Upper Eocene of England) and that of the succinites (Baltic, Rovno, and other Late Eocene European ambers); they are common in the succinites and in the Bembridge Marls and are only known from these localities.     

<Emphasis Type="Italic">Protocopidosoma</Emphasis> gen. nov. (Hymenoptera,Chalcidoidea, Encyrtidae) from the Late Eocene Danish amber

Protocopidosoma kononovae gen. et sp. nov. from Danish amber is described. The specimen is very similar to representatives of the extant genus Copidosoma Ratzeburg, 1844 and differs from them in the long and wide veins in the forewing, as in most encyrtids from Late Eocene European ambers.