亚洲西部和欧洲东南部芍药属的一个亚种

根据存于欧洲一些大标本馆的大量标本的观察,分布于希腊爱琴海东部、土耳其南部、塞浦路斯、黎巴嫩、叙利亚、伊拉克北部的Paeonia mascula与欧洲中部至巴尔干半岛的均不相同,可确认为一个独立的亚种。绝大多数个体总有一些小叶全裂,因此小叶及裂片数为(9)12～18(23);小叶宽椭圆形至卵圆形,通常两面无毛或背面疏生柔毛。     

Paeonia daurica Andrews or P. mascula ssp. triternata (Pall. ex DC.) Stearn & P. H. Davis (Paeoniaceae)?

The peony in the Crimea of Ukraine and its allied populations have been variously taxonomically treated, as Paeonia daurica Andrews or P. mascula ssp. triternata (Pall. ex DC.) Stearn & P. H. Davis. Supported by the National Geographical Society, we have conducted extensive field observations and population sampling of this group in Turkey. In addition, relevant herbarium specimens from the herbaria B, BEO, BM, BUCA, E, G, GZU, K, P, SA, SOM, UPA, and WU were critically examined. Principal coordinate analysis was performed using MVSP-Version 3.13b analysis software. As a result, P. daurica was shown to be clearly differentiated from P. mascula in the number of leaflets/segments of the lower leaves and the shape of the terminal leaflets. P. daurica is diploid, except for three local tetraploids in the Caucasus, whereas P. mascula is consistently tetraploid. The two units were not found growing together, even in southern Turkey, where they are sympatric. P. daurica is considered to be a good species, which ranges from Croatia to Iran through Turkey and the Caucasus, and comprises six subspecies.  © 2007 The Linnean Society of London, Botanical Journal of the Linnean Society , 2007, 154 , 1–11.     

紫斑牡丹及其一新亚种

本文回顾了有关紫斑牡丹的调查和分类历史。它曾被混同于 P．suffruticosa,P．papaveracea 及P．suffruticosa var．papaveracea。它以叶2至3回羽状复叶,小叶17～33,花瓣白色,基部有大紫斑, 花丝黄色,花盘和柱头淡黄色区别于近缘种。种下分化为两个异域的亚种;秦岭北坡的紫斑牡丹小叶全 部或大部分分裂,是一个新亚种,P．rockii subsp．taibaishanica,而P.rockii subsp．linyanshanii T.Hong et G．L.ostii则是P．rockii subsp．rockii的多余名。     

Paeonia rockii and Its One New Subspecies from Mt. Taibai,Shaanxi of China

The history of exploration and classification of Paeonia rockii is reviewed in the present paper. Two allopatric subspecies in this species are recognized. The new subspecies, subsp, taibaishanica Hong, found on northern slope of the Qingling Range has leaflets ovate or rounded, totally or mostly lobed. P. rockii subsp, linyanshanii T. Hong et G. L. Ostiis found to be a superfluous name of P. rockii subsp, rockii.     

中国野生牡丹研究 二：芍药属牡丹组新分类群

本文发表一个新亚种和一个新等级。新亚种林氏牡丹和紫斑牡丹同属一个种系,花瓣基部均具紫斑,花丝,花盘及柱头均为白色;但林氏牡丹的小叶多为披针形或窄卵形,全缘,而紫斑牡丹的小叶多为卵形或卵圆形,常１－３深裂。     

Karyotypes of four wild Paeonia species from Greece

The karyotypes of eight wild Greek Paeonia taxa were studied, viz. P. clusii ssp. clusii (2n = 10, 20), P. clusii ssp. rhodia (2n = 10), P. mascula ssp. mascula (2n = 20), P. mascula ssp. hellenica (2n = 20), P. mascula ssp. icarica (2n = 20), P. mascula ssp. russi (2n = 10), P. peregrina (2n = 20) and P. parnassica (2n = 20). P. mascula and P. clusii have a very similar haploid chromosome complement, x = 5, including three metacentric chromosomes, one submetacentric and one subtelocentric. In the chromosome complements of the tetraploid P. peregrina and P. parnassica the same chromosome types are included but their karyotype differ from those of the former two species by a number of characters. The main karyotype differences are concerned with the size differentiation of the chromosomes which in the case of P. peregrina has resulted in the diploidization of the tetraploid karyotype.
From the individuals examined, six were found to have abnormal karyotypes suggesting chromosomal exchanges. These structural aberrations in which mainly submetacentric and subtelocentric chromosomes are involved have also been described and discussed.     

牡丹一新种——中原牡丹,及银屏牡丹的订正

分子证据和新增加的形态证据显示,银屏牡丹Paeonia suffruticosa ssp．yinpingmudan作为新亚种发表时所依据的两份标本实为两个实体,产自安徽巢湖的（潘开玉和谢中稳9701,模式）实为凤丹户P.ostii的成员,而产自河南嵩县的（洪德元等H97010）实为一个新分类群。本文把P.suffruticosa ssp．yinpingmudan处理为P．ostii的异名,并依据河南的标本描述了一个新种——中原牡丹P.cathayana D．Y Hong＆K．Y.Pan。中原牡丹与风丹和矮牡丹P.jishanensis近缘,区别在于前者下部叶有ll-15片小叶,花白色：后者小叶背面有毛,裂片多,萼片顶端圆钝,花白色。     

Paeonia rotundiloba (D.Y.Hong) D.Y.Hong: A new status in tree peonies (Paeoniaceae)

Paeonia decomposita Hand.-Mazz.subsp,rotundiloba D.Y.Hong is here raised to the specific rank,P.rotundiloba (D.Y.Hong) D.Y.Hong,based on its distinctiveness of four diagnostic characters:number of carpels;height of disk; number of leaflets of the lower leaves,and shape of the terminal leaflets; and unpublished molecular data.Paeonia rotundiloba differs distinctly from P.decomposita in having carpels mostly 3,less often 2 or 4,very rarely 5 (vs.almost always 5,very occasionally 4 or 3),disk 8-15 mm high (vs.4-9.6 mm),leaflets mostly 19-39(vs.29-63) in number,and ratio of length to width of the terminal leaflets 1.09-1.93 (vs.1.81-2.99).     

Paeonia (Paeoniaceae) in the Caucasus

The taxonomy of the genus Paeonia in the Caucasus has been controversial, with recognized species varying in number from one to 13. The taxonomic history of Paeonia in this area is reviewed (including an analysis of the characters used by previous authors) based on extensive field observations, population sampling and critical examination of a large number of herbarium specimens. The results show that Paeonia may be divided into three groups. The P. intermedia group is known from only a single population. In the P. tenuifolia group, all the characters used for distinguishing the three previously recognized species were found to be polymorphic. In the P. daurica group, petal colour, shape and size of leaflets, and indumentum of leaflets and carpels were used to distinguish nine species, but these characters were found to be polymorphic or continuous in variation, and thus can only be used for infraspecific classification. Thus, three species are recognized: P. intermedia , P. tenuifolia and P. daurica . The last species is further divided into five subspecies: sspp. coriifolia , wittmanniana , mlokosewitschii , macrophylla and tomentosa stat. nov .  © 2003 The Linnean Society of London, Botanical Journal of the Linnean Society, 2003, 143 , 135–150.     

Paeonia rotundiloba (D. Y. Hong) D. Y. Hong: A new status in tree peonies (Paeoniaceae)

Abstract Paeonia decomposita Hand.‐Mazz. subsp. rotundiloba D. Y. Hong is here raised to the specific rank, P. rotundiloba (D. Y. Hong) D. Y. Hong, based on its distinctiveness of four diagnostic characters: number of carpels; height of disk; number of leaflets of the lower leaves, and shape of the terminal leaflets; and unpublished molecular data. Paeonia rotundiloba differs distinctly from P. decomposita in having carpels mostly 3, less often 2 or 4, very rarely 5 (vs. almost always 5, very occasionally 4 or 3), disk 8–15 mm high (vs. 4–9.6 mm), leaflets mostly 19–39 (vs. 29–63) in number, and ratio of length to width of the terminal leaflets 1.09–1.93 (vs. 1.81–2.99).     

Lathyrus nivalis subsp. sahinii subsp. nov. (sect. Platystylis, Leguminosae) from Turkey

The new subspecies Lathyrus nivalis Hand.-Mazz. subsp. sahinii H. Genç (sect. Platystylis , Leguminosae) from Turkey is described and illustrated with diagnostic characters and taxonomic comments. The characters of the new subspecies are compared with those of the type subspecies, from which it differs mainly in the shape, length, width and venation of leaflets, length and width of the stipules, flower color, legume, and style length. In addition, the geographic distribution of the new subspecies is outlined.     

Simaba guianensis subsp. huberi, a new Venezuelan taxon of Simaroubaceae

Simaba guianensis subsp.huberi (Simaroubaceae), restricted to the savannas of the Venezuelan State of Amazonas, is described and illustrated. The new subspecies is distinguished from the other two subspecies ofS. guianensis by its smaller size, characteristics of the leaflets (size, texture, apex shape, and margin curvature), and fruit color.     

芍药属牡丹组分类新注

自我们近年发表一系列牡丹组分类文章以来, 国内外基本上赞同我们8个种的分类系统,但对一些问题仍有不同见解。本文进一步申述牡丹Paeonia suffruticosa Andrews 是一个独立的种而不是人工杂种, 以及银屏牡丹P. suffruticosa ssp. yinpingmudan是牡丹P. suffruticosa的野生类型而不是逸生的P. ostii的理由。上述论点也得到了分子树的支持。P. jishanensis T. Hong &; W. Z. Zhao是一个合法名称, 而P. spontanea (Rehder) T. Hong &; W. Z. Zhao则确实是一个多余名。太白山紫斑牡丹的学名应是P. rockii ssp. atava (Brühl) D. Y. Hong &; K. Y. Pan, 因此P. moutan Sims ssp. atava Brühl不应是可疑的分类群。本文还对Halda的6个组合和两个杂交种名作了处理。结果, 本文包括了5个新异名和一个新组合。     

Taxonomical history and revision of Paeonia sect. Moutan (Paeoniaceae)

The taxonomical history of the woody group(peonies) in the genus Paeonia is reviewed in the present paper. The group is endemic to China, but Paeonia suffruticosa is commonly cultivated throughout the north temperate region and has long been known as “the King of Flowers” in China. However, the taxonomy of the group had been neglected before the 1990' s. Since 1990, a number of new species and subspecies have been published. With the support from the National Geographic Society, the senior author of the present paper and his coworkers have made expeditions to all parts of the distribution area of the group, visited all the type localities and sampled a total of 64 populations. Based on the character analysis and examination of the type specimens or photos, each taxon is reviewed with its name checked nomenclaturally, and finally the taxonomy of the whole group is revised. As a result, eight species, three of which each contains two subspecies, are recognized. They are Paeonia suffruticosa Andrews subsp. suffruticosa and subsp. yinpingmudan D. Y. Hong, K. Y. Pan et Z. W. Xie, P. jishanensis T. Hong et W. Z. Zhao, P. qiui Y. L. Pei et D. Y. Hong, P. ostii T. Hong et J. X. Zhang, P. rockii (S. G. Haw et L. A. Lauener )T. Hong et J. J. Li subsp. rockii and subsp. taibaishanica D. Y. Hong, P. decomposita Hand.-Mazz. Subsp. decomposita and subsp. rotundiloba D. Y. Hong, P. delavayi Franeh. and P. ludlowii (Stern et Taylor)D. Y. Hong. P. baokangensis Z. L. Dai et T. Hong and P. yananensis T. Hong et M. R. Li are considered as hybrids between P. rockii and P. qiui and between P. rockii and P. jishanensis respectively. In addition, the reduction of a number of names recently published to synonyms is explained.     

Yinpingmudan,the Wild Relative of the King of Flowers,Paeonia suffruticosa Andrews

Paeonia suffruticosa Andrews subsp. spontanea (Rehder) S. G. Haw et L. A. Lauener ( ＝P. spontanea ＝ P. jishanensis) has been considered as the wild form of widely cultivated P. suffruticosa. However, the subspecies differs from the cultivated one in shape, division and indumentum of leaves and reproductive biology. As a result of extensive field work, a form with two individuals in Yinping Shan, Caohu, Anhui, and Songxian, Henan, are found to be extremely similar to P. suffruticosa, but apparently different from P. jishanensis. The form is considered to be the wild one of P. suffruticosa and described as new: P. suffruticosa subsp, yinpingmudan Hong, K. Y. Pan et Z. W. Xie.     

Plant regeneration in <Emphasis Type="Italic">Arachis stenosperma</Emphasis> Krapov. and W. C. Gregory from roots and calluses derived from leaflets of <Emphasis Type="Italic">in vitro</Emphasis> plants

Seed explants of A. stenosperma were cultured on MS medium supplemented with 6-benzylaminopurine with the aim of rescuing nonviable accessions stored in seed bank conditions. The regeneration potential of leaf explants from in vitro plants derived from embryonic axes was studied by using whole leaflets and leaflet segments. Explants were cultured on Murashige and Skoog (MS) medium supplemented with different concentrations of 6-benzylaminopurine and naphthalene acetic acid. Indirect organogenesis was observed in response to 6-benzylaminopurine, either alone or in association with naphthalene acetic acid, in both explant types. Media supplemented with naphthalene acetic acid as the sole growth regulator induced rhizogenesis in whole leaflets and leaflet segments, with subsequent shoot production directly from the roots.     

银屏牡丹——花王牡丹的野生近亲

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Leydigia louisi louisi Jenkin, 1934 in the Neotropics, L. louisi mexicana n.subsp. in the Central Mexican highlands

Leydigia louisi Jenkin, 1934 was recently redescribed by Kotov (2003). We found in Guatemala and South of Mexico populations of this species, similar with the typical African form. A re-examination of Daday's (1902) specimens from Patagonia demonstrated that his material, determined as L. leydigi, was also L. louisi. At the same time, we describe a new subspecies Leydigia louisi mexicana n.subsp. from four water bodies on the central Mexican plateau (State of Mexico). Parthenogenetic females, ephippial females, juvenile and adult males are described. SEM illustrations are given for L. louisi mexicana n.sp. It differs from the nominotypical subspecies by (1) ocellus of irregular shape; (2) chitinous insertions within distal segments of swimming setae far from bases of segments; (3) on IDL, 3 bunches of very long setules; (4) setules on distal lobe of limb II long; (5) longest seta of exopodite III with long, fine setules; (6) seta 1 of exopodite IV relatively short; (7) a two-lobed body on inner margin of limb V. Features 2–6 were never used in the genus previously, but were applied in the systematics of other genera of Chydoridae. The ranges of these two subspecies are, probably, the Neotropical and Holarctic regions.     

Cytogeography and taxonomy of the Paeonia obovata polyploid complex (Paeoniaceae)

 The taxonomy of the Paeonia obovata polyploid complex in E-Asia has been controversial with treatments ranging from one species and two varieties to five species and numerous infraspecific taxa. Extensive observations in the field, analysis of morphological characters based on 32 populations sampled, and the examination of a large amount of herbarium specimens show that all the characters used by previous authors for distinguishing the species are variable between or even within populations: Leaf position, petal colour, stamen number (from 21 to 110 in the red-flowered and from 58 to 239 in the white-flowered form), filament and anther colour, style length as well as follicle number and position. Therefore, all these characters are of no value for species delimitation. However, an extensive chromosome survey and the analysis of morphological characters show that the ploidy level is basically correlated with the geographical distribution and with the indumentum on the lower leaf surface. Thus, the recognition of one species with two subspecies is proposed: Paeonia obovata subsp. obovata, mostly diploid, widely distributed in the north, east and south of the distribution range and glabrous to sparsely pubescent or hirsute on the lower leaf surface, and subsp. willmottiae, tetraploid, limited to the west and with leaves densely hirsute or pubescent below. Received October 30, 1998 Accepted June 21, 2000