On the finding of ornithomimid dinosaurs (Saurischia, Ornithomimosauria) in the Upper Cretaceous beds of Tajikistan

Fragmentary remains of ornithomimids (Ornithomimidae indet.) from the Lower Santonian (Upper Cretaceous) Kansai locality in northwestern Fergana (Tajikistan) are described, and the composition of its dinosaurian assemblage is updated.     

A mosasaur from the Cenomanian of Russia

The premaxilla of a mosasauroid lizard from the Lysaya Gora 3 locality in Saratov (Upper Cretaceous, Cenomanian) is described. To date, the oldest known mosasaur remains have been dated Turonian. The premaxilla is most similar in morphology to that of Russellosaurus from the Turonian of the United States; however, the absence of sufficient diagnostic characters only allows its attribution to Mosasauridae indet.     

Découverte d’une nouvelle faune d’âge cénomanien dans la région de Forthassa (Atlas saharien occidental,Algérie) : Implications biostratigraphiques et paléoenvironnementales

The upper Cretaceous of the eastern part of the Ksour Mountains (Western Saharan Atlas, Algeria) is classically subdivided into three fossiliferous lithostratigraphic formations: El Rhelida (lower Cenomanian), Mdaouer (lower-middle Cenomanian) and Rhoundjaïa (upper Cenomanian-lower Turonian). The western part of the Ksour Mountains, which is less documented and poorly known, is here the subject of a detailed sedimentological and palaeontological study. Near Hassi Djeifa locality (Forthassa) the upper Cretaceous is represented by the upper part of Mdaouer Formation and by the lowermost part of Rhoundjaïa Formation. The deposits attributed to the Mdaouer Formation are composed of reddish or green marlstone intercalated by biolaminated limestone, bioclastic limestone with gastropods and bivalves, and sandstone. The selachians (Mafdetia tibniensis, Baharipristis bastetiae), pycnodontiform fishes, and ostracod fauna (Damonella spp., Ilyocypris? sp., Cypridea sp., Paracypria? sp.) coming from marly beds are studied in detail. The two selachian species (Order of Rajiformes) were so far restricted to the Early Cenomanian of Egypt. Our sedimentological and palaeontological data support a probable Early Cenomanian age of the studied deposits and indicate a coastal plain palaeoenvironment continually influenced by the action of tide.     

Mid-cretaceous Ornithocheirids (Pterosauria,Ornithocheiridae) from Russia and Uzbekistan

Ornithocheirid remains from 16 localities of Russia and Uzbekistan are reviewed. New material of ornithocheirids from the localities of Mogoito (Barremian-Aptian), Stoilo quarry (Albian-Cenomanian), and Khodzhakul (Cenomanian) is described. The Khodzhakul locality has yielded Ornithocheirus (the first record of this genus in Asia).     

First definitive stegosaur from the Middle Jurassic of Kyrgyzstan

Four articulated posterior dorsal vertebrae from the Middle Jurassic (Callovian) Balabansai Svita at Sarykamyshsai locality in the northern Fergana Valley represent the first record of Stegosauria in Kyrgyzstan. The vertebrae cannot be attributed to any known stegosaurian genus. This is one of the oldest records for the Stegosauria.      

Historical distribution patterns of trigonioidids (non-marine Cretaceous bivalves) in Asia and their palaeogeographic significance

The non-marine trigonioidid bivalves show five phases of radiation in the Cretaceous of Pal-Asia: pre-Aptian (?Valanginian/Hauterivian–Barremian), Aptian, Albian, Cenomanian and Turonian–Maastrichtian. Their distribution patterns show two distinct palaeo-river systems feeding trigonioidids. Before the Cenomanian, the river system occupied the southwestern–southern–southeastern Pal-Asian continental margin areas. During the Turonian–Maastrichtian, it extended along the line of southcentral China−eastern China−northeastern China−northern China and Mongolia−northwestern China–eastern Fergana Basin of Kyrgyzstan−western Tajikistan Basin of Tajikistan–Tashkent area of Kazakhstan−central Kyzylkum of northern Uzbekistan–Aral Sea area of Kazakhstan. Furthermore, the general trigonioidid distribution pattern demonstrates that Japan was probably attached to part of eastern China and/or Korea during the ?Valanginian/Hauterivian–Cenomanian stages.     

Ammonites diartianus d'Orbigny, 1850,Vascoceratidae du cénomanien supérieur de saint-calais (Sarthe)

Ammonites diartianus d'Orbigny is a Vascocerasoof late Cenomanian (Sciponoceras gracile Zone) age, occuring in both Sarthe (France) and southern England. It is the earliest vascoceratid known from France, and its recognition is of great significance, lending support to recent suggestions that the classic Vascoceras gamai - mundaeChoffat group are of Upper Cenomanian age.     

Ulphaceaster sarthacensis,nouveau genre et nouvelle espèce d’échinide Archiaciidae du Cénomanien (Sarthe,France)

A new genus and a new species of the echinoid family Archiaciidae from Cenomanian deposits in Sarthe (Paris Basin, France) are described. The genus Ulphaceaster nov. gen. can be distinguished from other archiaciids by its anterior ambulacrum, which is located in a deep groove running from the peristome to the apex where the test enlarges in a tea-spoon shape. The apical rostrum is taller and more slender than for species of the close genus Claviaster. The species Uphaceaster sarthacensis sp. nov. is known from the Saint-Ulphace locality only (Sables et Grès de Lamnay Formation, La Garenne quarry), where it is found in shallow marine sandy limestones, from infralittoral deposit environments.     

Actinocamax primus arkhangelsky (belemnitellidae; upper cretaceous) biometry,comparison and biostratigraphy

A sample ofActinocamax primus Arkhangelsky, 1912 from the Lower Middle Cenomanian limestones of the Wunstorf quarry west of Hannover (NW Germany) is studied by univariate and bivariate biometric methods in order to analyse the variation of critical characters.A. primus is closely related toA. plenus (Blainville, 1825) but differs from that species by being smaller and more slender.A. primus appears in the Lower Cenomanian and continues into the Lower Middle Cenomanian. It is mainly distributed in the northern part of the North European Palaeobiogeographic Province.A. plenus is recorded from the Middle Cenomanian-lower Lower Turonian of the Russian Platform, but only from the Middle Upper Cenomanian in NW Europe. It is widespread in the North European Province.The primus event in the Lower Middle Cenomanian and theplenus event in the Middle Upper Cenomanian are briefly discussed.     

A new pterosaurian record from the Cenomanian of the Volga Region

Abstract-A rostral fragment of the dentary symphysis from the Cenomanian Melovatka-3 locality (Volgograd Region) is referred to the genus Lonchodectes based on the presence of elevated jaw borders, longitudinal groove on the dorsal surface of the dentary symphysis, and small alveoli of teeth, almost uniform in size and widely spaced. Lonchodectes sp. from Melovatka-3 is closely similar to L. platystomus from the Albian of England. This is the first record of the family Lonchodectidae in Russia and outside Great Britain.     

The Cenomanian–Turonian boundary in Jordan: Ammonite biostratigraphy and faunal turnover

Well-exposed fossiliferous Upper Cenomanian–Lower Turonian marine sedimentary rocks are present in west-central Jordan. Ammonites serve as an important faunal marker for this interval and can be used to subdivide the Cenomanian–Turonian transition into two upper Cenomanian biozones (Neolobites vibrayeanus and Vascoceras cauvini) and two lower Turonian biozones (Vascoceras proprium and Choffaticeras segne). A revised stratigraphic range of the Vascoceras cauvini Zone in the study area is proposed, consisting of the Metoicoceras geslinianum and Neocardioceras juddii zones of the standard zonation. Based on intercontinental biostratigraphic correlation, a minor unconformity appears to be present around the Cenomanian–Turonian boundary, and a part of the lower Turonian is probably missing. In addition, a faunal turnover is recorded in the uppermost Cenomanian, marked by the disappearance of most of the Cenomanian taxa, including Costagyra olisiponensis (Sharpe), Ceratostreon flabellatum (Goldfuss), Ilymatogyra africana (Lamarck), Rhynchostreon suborbiculatum (Lamarck), Harpagodes nodosus (Sowerby), and Heterodiadema libycum (Desor). This bioevent is thought to be an effect of the Oceanic Anoxic Event OAE 2; the dramatic shifts in species richness and diversity spanning the Cenomanian–Turonian boundary in the study area occurred in response to the major paleoclimatic and paleoenvironmental perturbations prevailing at that time. The stratigraphic and paleontological patterns studied in Jordan are very similar to those recorded in Egypt in terms of litho- and biostratigraphy, event stratigraphy, and macroinvertebrate content, suggesting the presence of uniform triggering mechanisms and bio-sedimentary responses in the Upper Cretaceous basins of the Middle East and providing clues for a high-resolution correlation between the two areas.     

A giant ornithomimosaur from the Early Cretaceous of China

Ornithomimosaurs (ostrich-mimic dinosaurs) are a common element of some Cretaceous dinosaur assemblages of Asia and North America. Here, we describe a new species of ornithomimosaur, Beishanlong grandis, from an associated, partial postcranial skeleton from the Aptian-Albian Xinminpu Group of northern Gansu, China. Beishanlong is similar to another Aptian-Albian ornithomimosaur, Harpymimus, with which it shares a phylogenetic position as more derived than the Barremian Shenzhousaurus and as sister to a Late Cretaceous clade composed of Garudimimus and the Ornithomimidae. Beishanlong is one of the largest definitive ornithomimosaurs yet described, though histological analysis shows that the holotype individual was still growing at its death. Together with the co-eval and sympatric therizinosaur Suzhousaurus and the oviraptorosaur Gigantraptor, Beishanlong provides evidence for the parallel evolution of gigantism in separate lineages of beaked and possibly herbivorous coelurosaurs within a short time span in Central Asia.     

<Emphasis Type="Italic">Ardea sytchevskayae</Emphasis> sp. nov., a new heron species (Aves: Ardeidae) from the Middle Miocene of Mongolia

A new heron species, Ardea sytchevskayae sp. nov., from the Middle Miocene Sharga locality is described. The new species is represented by an almost complete coracoid and a few partial coracoids, partial scapula, and caprometacarpus. A. sytchevskayae was a relatively large heron, of the same size as A. alba, and intermediate in morphology between A. alba and other species traditionally included in this genus. In addition, the Sharga locality has yielded a large bittern and a medium-sized heron of uncertain taxonomic position.     

Probable Ankylosaur Ossicles from the Middle Cenomanian Dunvegan Formation of Northwestern Alberta,Canada

A sample of six probable fragmentary ankylosaur ossicles, collected from Cenomanian deposits of the Dunvegan Formation along the Peace River, represent one of the first dinosaurian skeletal fossils reported from pre-Santonian deposits in Alberta. Specimens were identified as ankylosaur by means of a palaeohistological analysis. The primary tissue is composed of zonal interwoven structural fibre bundles with irregularly-shaped lacunae, unlike the elongate lacunae of the secondary lamellar bone. The locality represents the most northerly Cenomanian occurrence of ankylosaur skeletal remains. Further fieldwork in under-examined areas of the province carries potential for additional finds.     

Late Cretaceous (Cenomanian–Maastrichtian) calcareous nannofossils from Goban Spur (DSDP Sites 549, 551): Implications for the palaeoceanography of the proto North Atlantic

The early late Cretaceous (Cenomanian–early Turonian) is thought to have been one of the warmest periods of the Phanerozoic. This period was characterised by tropical sea surface temperatures of up to 36 °C and a pole-to-equator-gradient of less than 10 °C. The subsequent Turonian–Maastrichtian was characterised by a continuous climatic cooling, peaking in the Maastrichtian. This climatic cooling and the resulting palaeoceanographic changes had an impact on planktic primary producer communities including calcareous nannofossils. In order to gain a better understanding of these Cenomanian–Maastrichtian palaeoceanographic changes, calcareous nannofossils have been studied from the proto North Atlantic (Goban Spur, DSDP Sites 549, 551). In order to see potential differences between open oceanic and shelf dwelling nannofossils, the data from Goban Spur have been compared to findings from the European shelf (northern Germany).A total of 77 samples from Goban Spur were studied for calcareous nannofossils revealing abundant (mean 6.2 billion specimens/g sediment) and highly diverse (mean 63 species/sample) nannofossil assemblages. The dominant taxa are Watznaueria spp. (mean 30.7%), Prediscosphaera spp. (mean 18.3%), Zeugrhabdotus spp. (mean 8.3%), Retecapsa spp. (mean 7.2%) and Biscutum spp. (mean 6.6%). The Cenomanian assemblages of both Goban Spur (open ocean) and Wunstorf (shelf) are characterised by elevated abundances of high fertility taxa like Biscutum spp., Zeugrhabdotus spp. and Tranolithus orionatus. Early Turonian to Maastrichtian calcareous nannofossil assemblages of Goban Spur are, however, quite different to those described from European sections. Oceanic taxa like Watznaueria spp., Retecapsa spp. and Cribrosphearella ehrenbergii dominate in Goban Spur whereas the fertility indicators Biscutum spp. and T. orionatus are more abundant in the European shelf assemblages. This shift from a homogeneous distribution of calcareous nannofossils in the Cenomanian towards a heterogeneous one in the Turonian–Maastrichtian implies a change of the ocean circulation. The “eddy ocean” system of the Cenomanian was replaced by an oceanic circulation similar to the modern one in the Turonian–Maastrichtian, caused by the cooling. The increased pole-to-equator-gradients resulted in an oceanic circulation similar to the modern one.     

Cenomanian-Turonian rudists from Western Sinai, Egypt: Systematic paleontology and paleoecology

The Cenomanian-Turonian sequence is well exposed in western central Sinai and contains a considerable number of rudist species. The identified rudists belong to Radiolitidae, Hippuritidae and Requieniidae. Fifteen species are described, belonging to ten genera: Requienia, Toucasia, Apricardia, Radiolites, Eoradiolites, Praeradiolites, Sphaerulites, Sauvagesia, Hippurites, and Vaccinites. Eleven species are reported in the Cenomanian and four species in the Turonian. Within the Cenomanian, two new species are described: Requienia tortuosi and Eoradiolites lenisexternus. The genus Requienia and the species Apricardia carentonenis d’Orbigny, Eoradiolites syriacus (Conrad), Sphaerulites agariciformis Delamétherie, Sphaerulites depressus Blanckenhorn and Vaccinites cf. grossouvrei (Douvillé) are reported for the first time from Egypt. The Cenomanian and Turonian rudists in the western Sinai show either elevator or clinger morphotypes, with the predominance of the former type; recumbent mode of life is rarely represented by some Requieniidae. The elevators are of isolated and clustered occurrences and more represented in the Cenomanian sequence. Rudists of the study area are mainly of parautochthonous fabrics with low to dense packing. Autochthonous fabrics are also achieved by some species, mainly in the Turonian. The disappearance of rudists from the middle part of the sequence and above the Cenomanian/Turonian boundary is due to a deeper setting that resulted from sea-level rise or seafloor subsidence (shelf drowning). The occurrence of rudists with oysters and other benthic fossils in the siliciclastic and carbonate sediments of the Cenomanian and Turonian sedimentary rocks in the Sinai indicate that the sequence was deposited on a broad, shallow shelf. Although the first marine transgression invaded the central Sinai in the late Cenomanian, transgressive deepening conditions continued until the Turonian. The rudists of central Sinai are of Tethyan affinity with significant relation with North Africa, the Middle East, and Southern Europe.     

Cenomanian Cephalopods of Bellezma-Aures mountains,NE Algeria: Taxonomy and biostratigraphy

The Cenomanian Smail Marl Formation in the Bellezma and Aures mountains of northeastern Algeria is well exposed and rich in macroinvertebrates, including cephalopods. Twenty-five ammonite species and two nautiloid species from the Thénièt El Manchar, Djebel Metrassi and Djebel Bouarif sections are described and seven assemblage zones proposed as provisional working units: Mantelliceras mantelli–Sharpeiceras laticlavium Assemblage Zone (lower Cenomanian), Mantelliceras dixoni Assemblage Zone (uppermost lower Cenomanian), Cunningtoniceras inerme Assemblage Zone (lower middle Cenomanian), Acanthoceras rhotomagense and Acanthoceras cf. rhotomagense Assemblage Zone (middle middle Cenomanian), Acanthoceras amphibolum Assemblage Zone (upper middle Cenomanian), Acanthoceras jukesbrownei Assemblage Zone (uppermost middle Cenomanian), and Neolobites vibrayeanus Assemblage Zone (lower upper Cenomanian). The lower Cenomanian Mantelliceras mantelli–Sharpeiceras laticlavium Zone is subdivided into the Submantelliceras aumalense, Sharpeiceras schlueteri and Mantelliceras saxbii subzones. The base of the middle Cenomanian is marked by the disappearance of species of Mantelliceras and the appearance of the scleractinian coral Aspidiscus cristatus (Lamarck, 1801), associated with Cunningtoniceras inerme (Pervinquière, 1907). The middle Cenomanian contains abundant A. cristatus and species of Acanthoceras, i.e., A. cf. rhotomagense (Defrance in Brongniart, 1822), A. jukesbrownei (Spath, 1926) and A. amphibolum Morrow, 1935. An ammonite zonal scheme is then applied to the Cenomanian exposed in the Bellezma-Aures mountains located on the southern Tethys margin in Algeria.     

Cenomanian (Upper Cretaceous) bivalves from the Hameimat Massifs,north of Tebessa,Algeria: Systematics,biostratigraphy, palaeoecological and taphonomical remarks

In the eastern Saharan Atlas, particularly in the northern area of Tebessa Province (NE Algeria), the widely outcropping Cenomanian strata display a highly diversified macrofauna, among which bivalves are prominently represented. Twenty-eight bivalve species are here reported for the first time from the Cenomanian of Hameimat Massifs. Based on the stratigraphic distribution of these bivalves, five bivalve zones were recognized, i.e., Costagyra olisiponensis - Gyrostrea delettrei, Rhynchostreon suborbiculatum - Exogyra conica, Ceratostreon flabellatum, Ilymatogyra africana, and Pycnodonte vesicularis vesiculosa - Rastellum carinatum zones. Correlation to the ammonite biozones of the same region as follows: the Costagyra olisiponensis - Gyrostrea delettrei and the Rhynchostreon suborbiculatum - Exogyra conica zones occur respectively in the Sharpeiceras schlueteri and Mantelliceras saxbii subzones of the lower Cenomanian Mantelliceras mantelli Zone. The Ceratostreon flabellatum Zone is correlated with the middle Cenomanian Acanthoceras rhotomagense Zone. The Ilymatogyra africana Zone is correlated with the upper Cenomanian Calycoceras naviculare and the Metoicoceras geslinianum zones. Finally, the Pycnodonte vesicularis vesiculosa - Rastellum carinatum Zone represents the uppermost Cenomanian. Detailed analysis of biometrical and morphological features of these bivalve specimens provides the most reliable tool within the scope of palaeo-environmental reconstitution and the many palaeo-ecological variables that had driven the development and distribution of these macro-invertebrates. Comparison of these new data to those of adjacent south Tethyian areas supports the homogeneity of the Cenomanian bivalve faunas. Such an affinity underlines more vividly the favorable marine communications and currents driving the geographic dispersal of these bivalves during the Cenomanian.     

Elaphrosaurus,an ornithomimid dinosaur from the upper jurassic of north America and Africa

A humerus from the Morrison Formation (Upper Jurassic) of western North America is referred toElaphrosaurus which is represented by a skeleton from the Tendaguru Beds of Tanzania, East Africa. The referral ofCoelurus agilis Marsh (Morrison Formation, western U.S.A.) andTeinurosaurus sauvagei (Huene) (Kimmeridgian, France) toElaphrosaurus is incorrect. The recognition ofElaphrosaurus from North America provides additional evidence for a Morrison-Tendaguru land connection sometime during the Late Jurassic.Elaphrosaurus is referred to the family Ornithomimidae because the form of the humerus, slender and straight with a low deltopectoral crest, is diagnostic of this family.Deinocheirus from the Upper Cretaceous of Mongolia is a giant ornithomimid.