Soft anatomy and the affinities of conodonts

Recent claims that conodonts are members of the Craniata or Vertebrata are based in part upon soft tissue features that have been preserved in a small number of specimens. These features include what appear to be radials in the caudal fin and paired structures that have been identified as eye remnants. The evidence for radials is limited, but credible. However, the anatomy of extant cyclostomes suggests that the paired structures are more reasonably interpreted as otic capsules than the remnants of sclerotic eye capsules. Moreover, even if these structures are the remnants of eyes, conodonts might equally well be a sister group to the craniates as a member of that group. Aside from these paired structures, conodont fossils exhibit no features that are suggestive of a cartilaginous skeleton. Given that cyclostome fossils sometimes show evidence of the cartilages of the head, the apparent absence of a similar skeleton in conodont animals calls into question the claim that they are craniates. The simple single chevron shape of conodont myomeres also suggests that they lie outside of the Craniata. All living craniates have double-chevron myomeres as adults, whereas simple myomeres of the conodont type are found in the non-craniate cephalochordates. Thus the available soft tissue evidence suggests that conodonts are best regarded as the sister group of the craniates.     

基于深度残差神经网络迁移学习的牙形刺图像识别

为了探索基于深度神经网络模型的牙形刺图像智能识别效果,研究选取奥陶纪8种牙形刺作为研究对象,通过体视显微镜采集牙形刺图像1188幅,收集整理公开发表文献的牙形刺图像778幅,将图像数据集划分为训练集和测试集。通过对训练集图像进行旋转、翻转、滤波增强处理,解决了训练样本不足的问题。基于ResNet-18、ResNet-34、ResNet-50、ResNet-101、ResNet-152五种残差神经网络模型,采用迁移学习方法,对网络模型进行训练以获取模型参数,五种模型测试Top-1准确率分别为85.37%、85.85%、83.90%、81.95%、80.00%, Top-2准确率分别为94.63%、94.63%、94.15%、93.17%、93.66%,模型对牙形刺图像具有较好的识别效果。通过对比研究发现,ResNet-34识别准确率最高,说明对于特征简单的牙形刺属种,增加网络深度并不一定能提升准确率,而确定合适深度的模型则不仅可以提高识别准确率,还可以节约计算资源。通过ResNet-34模型的迁移学习训练和重新训练效果对比可以看出,迁移学习不仅可以获得较高的准确率,而且可以较快获取模型参...     

广西桂林泥盆纪牙形刺组合序列与海平面变化

本文以桂林灵川岩山圩中,晚泥盆世界线剖面,桂林沙河能源疗养院背后信都组－唐家湾组剖面及桂林额头村Ｄ／Ｃ界线剖面的牙形刺资料为基础,结合前人在桂林地区所做的牙形刺生物地层工作,识别出该区中,晚泥盆世牙形刺化石的１６个组合序列,并以此为尺度结合层序地层格架,层序界面及层序中的微相叠加形式,讨论该区中,晚泥盆世高分辨的海平面变化。     

Trilobites d'affinité bohémienne dans l'Éodévonien des Pyrénées centrales françaises

In the upper Adour valley, a series attributed to the Carboniferous yield conodonts and a variegated trilobite fauna with exclusively bohemian affinity. The trilobite fauna is represented by six families, the most of them being described in the Pyrénées for the first time. Trilobites indicate the Praguian whereas conodonts upper Lochkovian-Praguian age. A new unnamed subspecies of Crotalocephalus sternbergi (Boeck, 1827) is described.     

华南中泥盆世艾菲尔期浅海相四射珊瑚的组合特征和古生物地理区系

虽然在华南中泥盆世艾菲尔期(Eifelian)较深水或斜坡相地层中产有许多国际标准的牙形类带化石,但在广阔的浅海相地层中却很难寻觅到其踪影。华南浅海相沉积地区是否缺失艾菲尔阶?在野外众多剖面的实地考察中表明,从下泥盆统埃姆斯阶到中泥盆统吉维特阶都是连续沉积,中间并没有发现地层的缺失或间断现象。浅海相的艾菲尔阶与其上覆的吉维特阶和下伏的埃姆斯阶都是连续沉积,说明可能是由于海水太浅,不太适合那些国际标准分子的生存而已。艾菲尔期中期末发生一次生物灭绝事件(Mid-Eifelian event),favositids,heliolitids和许多古老类型的珊瑚在地球上灭绝。华南艾菲尔期四射珊瑚可以划分成下、上两个完全不同的组合:1)Utara-tuiasinen sis-Sociop hyllum minor组合(牙形类partitus带—costatus带);2)Columnaria spinosa-Dendrostella praerhenana组合(牙形类australis带—kocklianus带)。     

湘西北晚二叠世-早三叠世早期牙形石古生态

通过对牙形石器官的对称性、相分布和地球化学特征的研究,探讨湘西北晚二叠世至三叠纪初牙形动物的生态分布.按器官所包括不同对称型分子的比例,牙形动物器官的对称性可分为4级,器官的对称性越高,动物运动能力越强.湘西北晚二叠世一三叠纪初地层可分为6种沉积相,即盆地相、盆缘-台坡相、开阔台地相、边缘浅滩相、局限台地相和泥坪相.具不同对称级器官的属种在6种沉积相中的差异分布,反映了牙形动物的3种生态类型:自由游泳型、浮游型和底栖型.牙形石地球化学组分的变化与当时环境演变相吻合.随水深增加,化石中的P含量升高,而Ca、s、sr的含量降低;Al、Fe和Ba在近岸浅水环境的牙形石中富集;而Mg的富集指示高盐环境.基于上述研究,划分了4个牙形石相,建立了一个组合横向替代的牙形动物生态分布模式.     

Conodont element function

There are close similarities between conodont elements and teeth both in general shape and in that they possess pointed tips and have expanded bases with more porous tissue. A number of examples of conodont elements which parallel specific kinds of tooth shapes and organizations are added to earlier known similarities. Both in teeth and in conodont elements the surface structures include cutting edges, striations, and barbs. The change in strength of the conodont denticles caused by the evolution of white matter is also shown to agree with a tooth function. On the other hand, the elements grew throughout the life of the animal by lamellae added to the surface. The solution of this paradox is found in the elements alternating between a growth phase and a functional phase. During growth the oral surface of the elements was enveloped in folds of secreting soft tissue. Structures henceforth termed burrs were formed at the contacts between the folds. Parts of the burrs evolved into cutting edges. Three different bite types occurred among the conodont elements. Many (all?) conodonts were predators which used their elements to seize and to process the food mechanically. The shape of the conodont elements cannot be used for conclusions regarding the affinities of the conodonts. Similarly, an identification of a fossil as belonging to the conodonts must be supported by other evidence than just shape.     

Conodont anatomy, chordate phylogeny and vertebrate classification

Interpretations of conodont anatomy and affinity continue to generate controversy. Fossilized soft-tissue evidence indicates that conodonts possessed eyes, extrinsic eye muscles, a notochord, myomeres, a differentiated tail with fin radiais, possible otic capsules and possible branchial structures. Indirect evidence suggests a differentiated brain and cartilaginous head skeleton. The multi-component phosphatic tissue complexes of the conodont feeding apparatus cannot be compared to the amorphous apatite of extant agnathan otoliths. By limiting cladistic analysis to a restricted selection of these characters the hypothesis that conodonts are a sister group of the clade comprising extant hagfish, lampreys and gnathostomes can be supported. However, exhaustive analysis of a more complete character-set strongly supports the hypothesis that conodonts are more derived than hagfish. From a taxonomic perspective, these two hypotheses have no effect on how conodonts should be classified. Whether they are a stem group (the former hypothesis) or part of the crown group (the latter), conodonts are clearly part of the total group Vertebrata (=Craniata).     

Conodont affinity and chordate phylogeny

Current information on the conodonts Clydagnathus windsorensis (Globensky) and Promissum pulchrum Kovács‐ Endrödy, together with the latest interpretations of conodont hard tissues, are reviewed and it is concluded that sufficient evidence exists to justify interpretation of the conodonts on a chordate model. A new phylogenetic analysis is undertaken, consisting of 17 chordate taxa and 103 morphological, physiological and biochemical characters; conodonts are included as a primary taxon. Various experiments with character coding, taxon deletion and the use of constraint trees are carried out. We conclude that conodonts are cladistically more derived than either hagfishes or lampreys because they possess a mineralised dermal skeleton and that they are the most plesiomorphic member of the total group Gnathostomata. We discuss the evolution of the nervous and sensory systems and the skeleton in the context of our optimal phylogenetic tree. There appears to be no simple evolution of free to canal‐enclosed neuromasts; organised neuromasts within canals appear to have arisen at least three times from free neuromasts or neuromasts arranged within grooves. The mineralised vertebrate skeleton first appeared as odontodes of dentine or dentine plus enamel in the paraconodont/euconodont feeding apparatus. Bone appeared later, co‐ordinate with the development of a dermal skeleton, and it appears to have been primitively acellular. Atubular dentine is more primitive than tubular dentine. However, the subsequent distribution of the different types of dentine (e.g. mesodentine, orthodentine), suggests that these tissue types are homoplastic. The topology of relationships and known stratigraphic ranges of taxa in our phylogeny predict the existence of myxinoids and petromyzontids in the Cambrian.     

Phylogenetic reconstruction of weevil superfamily Curculionoidea (coleoptera) using the SYNAP method

Phylogenetic relationships within the superfamily Curculionoidea were reconstructed. Autapomorphies of the superfamily Curculionoidea include more or less pronounced snout, clubbed antennae, and partially sclerotized or completely membranous male tergite 9. Weevil families can be divided into three groups. The first group includes the most primitive family Nemonychidae. The second group includes nine families (Anthribidae, Belidae, Oxycorynidae, Eccoptarthridae, Allocorynidae, Rhynchitidae, Attelabidae, Ithyceridae, and Brentidae). The third (“higher”) group includes six families (Brachyceridae, Cryptolaryngidae, Dryophthoridae, Curculionidae, Scolytidae, and Platypodidae).     

深圳市大鹏半岛蕨类植物区系及其生态特点

深圳市大鹏半岛的蕨类植物共有105种,它们隶属于35科、65属。其中含5种以上的科有水龙骨科(Dryopteridaceae)、金星蕨科(Thely pteridaceae)、凤尾蕨科(Pteridaceae)、卷柏科(Selaginellaceae)、鳞毛蕨科(Dryopteridaceae)、蹄盖蕨科(Athy riaceae)、铁角蕨科(Aspleniaceae)、鳞始蕨科(Lindsaeaceae)。其区系具有较强的热带性质,科、属、种的分布区类型均以热带至亚热带为主,分别占总科数的54.26%,总属数的92.86%以及总种数的70.87%。单属科和单种属的比例较高,分别占总科数的54.26%,总属数的69.23%。按生长基质的不同,将大鹏半岛的蕨类植物划为土生、石生、水生和附生四种生态类型,其中以土生类型为主。     

Allometry in Anisian (Middle Triassic) segminiplanate conodonts and its implications for conodont taxonomy

Conodonts are a clade of chordates and are valuable indicator fossils for biostratigraphy. The segminiplanate (neogondolelliform) conodonts represent a major morphological group ranging from upper Carboniferous to Upper Triassic marine sediments. However, the morphological similarity of segminiplanate P₁ elements generates problems for taxonomy, especially in the Permian and Triassic clades. This paper represents the first study of morphological variation in Triassic segminiplanate conodonts using a geometric morphometric approach. The laminar microstructures observed in conodont cross‐sections indicate that, within our analysed specimens, smaller conodonts with fewer laminae are generally from an earlier ontogenetic stage while larger conodonts with more laminae are from a later stage of ontogeny. Using linear regressions between relative warp scores from both upper and lateral views and conodont length, we demonstrate strongly allometric growth patterns for the species Paragondolella bifurcata Budurov & Stefanov. Our results indicate that the species‐group taxon Pg. praeszaboi bystrickyi (Kovacs et al.) is an early growth stage of Pg. bifurcata and thus synonymous. We suggest that the allometry of conodonts should be considered seriously, especially when there are numerous transitional morphologies between large‐ and small‐sized conodonts. Reconstructing the ontogenetic series and using larger‐sized conodonts within the numerous transitional morphologies in the population of a rock sample for the definition of new species are suggested for future studies.     

beta-fructosidase superfamily: homology with some alpha-L-arabinases and beta-D-xylosidases

Comparison of the amino acid sequences of four families of glycosyl hydrolases reveals that they are homologous and have several common conserved regions. Two of these families contain beta-fructosidases (glycosyl hydrolase families GH32 and GH68) and the other two include alpha-L-arabinases and beta-xylosidases (families GH43 and GH62). The latter two families are proposed to be grouped together with the former two into the beta-fructosidase (furanosidase) superfamily. Several ORFs can be considered as a fifth family of the superfamily on the basis of sequence similarity. It is shown for the first time that a glycosyl hydrolase superfamily can include enzymes with both inversion and retention mechanism of action. Composition of the active center for enzymes of the superfamily is discussed.     

胶东海岸的沙生植被

胶东半岛位于山东省东部,地处黄、渤海之滨,陆地海岸线全长1668.58km,占山东省陆地海岸线总长的一半以上。海岸地理位置约当北纬36°15′42″—38°24′00″,东经119°33′00″—122°42′18″之间,属暖温带生     

Pathologies of Devonian Conodont Elements of the Voronezh Anteclise

Paleontological Journal - Pathological phenomena in Devonian conodonts are separated into five genetic groups: (1) mechanical breakdown of hard tissues with and without subsequent...     

武陵山区蕨类植物研究

武陵山区蕨类植物资源丰富,现知有644种(包括变种),分属116属, 46科;蕨类区系属热带-亚热带山地类型;属的地理成份有12种类型,与日本、印度-马来西亚有密切关系;武陵山区蕨类区系属我国西南区系,与华中华东区关系密切,华南、华北区次之;蕨类垂直分布分四带。     

内蒙古巴特敖包地区早泥盆世牙形刺

内蒙古达茂旗(达尔罕茂明安联合旗)巴特敖包地区以碳酸岩相古生代地层发育而引人注意。经多年研究,特别是李文国等(1985)的研究,使本区珊瑚、腕足类、层孔虫、鹦鹉螺等大化石的研究工作已有一定基础,并依据牙形刺首先确定了在阿鲁共剖面有泥盆纪地层的存在。但本区泥盆纪地层仍存在很多问题,必须通过对牙形刺的研究加以解决。在王成源研究员的精心安排和亲自指导下,作者等在巴特敖包地区的七个剖面进行了系统取样研究。经对158个样品(重667kg)的分析,在本区发现了大量的原定为志留纪的早泥盆世地层,包括西别河组命名剖面。分析结果同时证明,本区不存在志留系温洛克统的沉积。巴特敖包剖面断层发育,志留—泥盆纪地层交替出现,巴特敖包组应取消,原定的阿鲁共剖面层序颠倒。本区地层应厘订为志留系的西别河组(罗德洛统—普里道利统)和下泥盆统的阿鲁共组(Lochkovian阶),前者以包尔汉图剖面为标准剖面,后者应寻找更好的标准剖面。本文重点总结巴特敖包地区早泥盆世的牙形刺。本区共建立了4个牙形刺带,文中共描述了8牙形刺属,21种,其中包括4亚种、1新种(Belodellataeniocuspidatasp.nov.)和1未定种。     

四川盆地下三叠统牙形刺及其油气意义

作者对四川盆地２４条以海相碳酸盐岩地层为主的下三叠统剖面作了系统的研究,在取得了丰富的牙形刺资料的基础,对下三叠统牙形刺进行了区域对比,划分了７个牙形刺带。并对牙形刺色变与油气的关系等进行了研究。编制了ＣＡＩ、古地温等值线图及其与油气分布关系图,并提出了该盆地最有利的油气远景区ＣＡＩ＝２－３;有利的产油气区ＣＡＩ＝２．５－３的新看法。     

Préparation des Conodontes par manipulationen milieu liquide

Described processing for preparation of conodonts improves the well-preservation of very fragile specimens. In classical procedures freed conodonts are airy moved without any technical care of rubbing and falling. In the new exposed technique freed conodonts are moved under distiled water; the previous inconveniences are avoided and the well-preserved conodonts number is increased.