New insects (Insecta: Trichoptera,Eoblattida) from the Lower Permian of Russia

The insect fauna of the Lower Permian Tyulkino locality in the Perm Region of Russia is considered. New members of the orders Trichoptera and Eoblattida are described. Members of the primitive family Microptysmatidae (suborder Protomeropina) are typical of the Lower and Middle Permian deposits of Eurasia and are represented by the two genera: Microptysmella Kukalová-Peck et Willmann, 1990 and Kamopanorpa Martynov, 1928. Kamopanorpa tyulkiana sp. nov. and K. tamarae sp. nov. (Trichoptera, Microptysmatidae) and Parakhosara kamica sp. nov. (Eoblattida, Megakhosaridae) are described.     

New Eoblattida from the Permian of Russia and the United States and the origin of earwigs (Insecta: Eoblattida,Forficulida)

New taxa of the order Eoblattida are described from the Lower Permian of the United States and Russia. Pryg absurdus gen. et sp. nov. of the family Prygidae fam. nov. is described from the Leonardian of Elmo (Kansas, United States). Parapryg alogus gen. et sp. nov. of the same family is described from the Vyatkian of Nedubrovo (Vologda Region, Russia). Kamamica promota gen. et sp. nov. of the family Tillyardembiidae is described from the Ufimian of Tyul’kino (Perm Region, Russia). The similarity of the new family with Tillyardembiidae and with basal earwigs, especially Bardacoleidae, is discussed. A hypothesis is proposed about the monophyletic origin of all three groups from spanioderids or other eoblattids close to them.     

New and little known blattogryllidae (Insecta: Grylloblattida) from the Permian and Triassic of Russia,Kazakhstan, and Kyrgyzstan

New members of the family Blattogryllidae are described, including Permoblattogryllus praecox gen. et sp. nov. from the Middle Permian Soyana locality in Russia, Protoblattogryllus nedubrovensis from the Lower Triassic Nedubrovo locality in Russia, Madygenocephalus micropteron gen. et sp. nov. (the second known case of brachyptery in grylloblattids), and Costatoviblatta similis sp. nov. from the Middle Triassic Madygen locality in Kyrgyzstan. Protoblattogryllus zajsanicus Storozhenko, 1990 from the Upper Permian Karaungir II locality in Kazakhstan is redescribed. The genus Microblattogryllus Storozhenko, 1990 from Madygen is considered to be a synonym of Protoblattogryllus Storozhenko, 1990. Protoblattogryllus abruptus Storozhenko, 1990 from the Madygen locality is transferred to the genus Mesoblattogryllus Storozhenko, 1990.     

New and little-known Eoblattida (Insecta) from the Paleozoic of Russia

New taxa of the insect order Eoblattida from the Upper Carboniferous of Russia are described, including Narkemina kata sp. nov., Narkeminopsis inversa sp. nov., Carbonokata storozhenkoi gen. et sp. nov., Tshunoptera ampla gen. et sp. nov., and Evenkiophlebia collucata gen. et sp. nov. from the Chunya locality (Krasnoyarsk Region), Narkemulla sibirica gen. et sp. nov. from the Chunya and Izykhskie Kopi localities, Khakassia (all Cnemidolestidae), and Izykhia tridentis gen. et sp. nov. from Izykhskie Kopi (?Spanioderidae). Narkeminuta permiana gen. et sp. nov. (Cnemidolestidae) is described from the Kedrovka locality (Kemerovo Region; Lower Permian), and Issadische maximum gen. et sp. nov. (Eoblattida incertae familiae) is descriobed from the Isady locality (Vologda Region; Upper Permian). Permeoblatta borealis Rasnitsyn et Aristov, 2010 (Idelinellidae) from Isady is redescribed.     

New Grylloblattida (Insecta) from the Upper Permian and Lower Triassic of European Russia and Kazakhstan

New grylloblattid insects are described from the Upper Permian and Lower Triassic of European Russia and Kazakhstan: Kuplya minutissima gen. et sp. nov. (Tshekardominidae) from the Severodvinian locality Novo-Aleksandrovka (Orenburg Region); Parachauliodites orthopteroides gen. et sp. nov. (Chaulioditidae), Permofossilis commasticatus gen. et sp. nov. (Permotermopsidae), Megakhosarodes tensilis sp. nov. (Megakhosaridae), and Baharellinus dilaceratus sp. nov. (Blattogryllidae) from the Severodvinian locality Isady (Vologda Region); Dvinopedes salariovensis gen. et sp. nov. (Chaulioditidae) from the Vyatkian locality Aristovo (Vologda Region); Klyazmia karasevi gen. et sp. nov. (Chaulioditidae) from the Vyatkian locality Sokovka (Vladimir Region); Megakhosarodes borealis sp. nov. (Megakhosaridae) from the Vyatkian locality Balymotikha (Vladimir Region); Sigmophlebia rugulosa sp. nov. (Tshekardominidae) from the Upper Permian locality Karaungir (East Kazakhstan Province); and Chauliodites kitshmengensis sp. nov. and C. nedubrovensis sp. nov. (Chaulioditidae) from the Induan locality Nedubrovo (Vologda Region).     

New beetles (Insecta,Coleoptera) from the Lower Triassic of European Russia

Fossil remains of beetles are described from two Lower Triassic localities: Entala (Induan) and Tikhvinskoe (Olenekian). Only one beetle fossil was previously known from the Lower Triassic of Tikhvinskoe. The fossils are rather few and poorly preserved, but they are worth describing as finds rare for the Lower Triassic. Five fossils from Entala most probably belong to beetles of the same species of the formal genus Pseudochrysomelites. Beetles of this genus are especially abundant in deposits close to the Permian–Triassic boundary and can be considered “disaster taxa.” There are no known cases, either in the Permian or in the Middle–Upper Triassic, of a random sample of five specimens belonging to a single species. This suggests that in the Entala oryctocenosis the species diversity of beetles is extremely low. All three beetle fossils found in Tikhvinskoe belong to beetles of different species, showing that diversity had already started to increase. However, it remained low, and all fossils belong to the formal family Schizocoleidae, and two of the three belong to the same genus, Pseudochrysomelites. The Khei-Yaga locality, which immediately follows Tikhvinskoe in time (topmost Olenekian or early Anisian), already contains beetles of the families Asiocoleidae and Permosynidae. In the Lower Anisian of the Buntsandstein, such typical Mesozoic beetles as Cupedidae and Coptoclavidae have been recorded. The appearance of such advanced beetles as early as the Lower Anisian suggests that the famous Permian–Triassic crisis was not as deep as it is usually believed, and many beetles survived it, disappearing, however, from the fossil record in the Early Triassic.     

New taxa of Mesorthopteridae (Insecta: Grylloblattida) from the Permian of Russia

New taxa of grylloblattids of the family Mesorthopteridae are described from the Permian of Russia: Parastenaropodites aquilonius sp. nov. from the Soyana locality (Middle Permian); P. circumhumatus sp. nov. from the Kul’chumovo locality; and Tshermyaninus biforis gen. et sp. nov., Permorthopteron foliaceus gen. et sp. nov., Parastenaropodites pannea sp. nov., P. exossis sp. nov., Mesoidelia procera sp. nov., and M. riphaea sp. nov. from the Isady locality (Upper Permian). Two species of the genus Khosaridelia Storozhenko, 1992 (family Permotermopsidae) and one species of the genus Megakhosarina Storozhenko, 1993 (family Megakhosaridae) are transferred to the genus Parastenaropodites Storozhenko, 1996 (family Mesorthopteridae); one species of the genus Parastenaropodites is transferred to the genus Austroidelia Riek, 1954; as a result, four new combinations are established: Khosaridelia rigida Aristov, 2005 =Parastenaropodites rigidus (Aristov, 2005), comb. nov. (Vorkuta locality, Lower Permian of Russia); Khosaridelia vyatica Aristov, 2009 =Parastenaropodites vyaticus (Aristov, 2009), comb. nov. (Karaungir locality, Middle Permian of Kazakhstan); Megakhosarina intricata Aristov, 2008 =Parastenaropodites intricatus (Aristov, 2008), comb. nov. (Soyana locality, Middle Permian of Russia); and Parastenaropodites nervosa Storozhenko, 1996 =Austroidelia nervosa (Storozhenko, 1996), comb. nov. (Madygen locality, Middle Triassic of Kyrgyzstan). An identification key to species of the genus Parastenaropodites is given.     

Revision of the family Idelinellidae,with a review of the Permian Eoblattida (Insecta)

Permian Eoblattida, which include the families Tillyardembiidae, Protembiidae (=Sylvardembiidae), Soyanopteridae, and Idelinellidae, are reviewed. The latter family is revised; it includes Idelinella macroptera Storozhenko, 1997 from the Middle Permian (Lower Kazanian Substage) of northern European Russia, Permostriga augustalis Novokshonov, 1999, Sylvastriga miranda Aristov, 2004, Strigulla cuculiophora (Aristov, 2002) comb. nov., Cucullistriga cucullata Aristov et Rasnitsyn, gen. et sp. nov., Scutistriga scutata Aristov et Rasnitsyn, gen. et sp. nov. from the Lower Permian (Kungurian Stage) of the Urals, Permeoblatta borealis Rasnitsyn et Aristov, 2010 from the Upper Permian (Upper Severodvinian Substage) of northern European Russia, and probably also Rasstriga americana Aristov et Rasnitsyn, gen. et sp. nov. from the Upper Carboniferous (Desmoinesian Stage) of Illinois.     

New insects (Insecta: Mecoptera,Grylloblattida) from the Middle Permian Chepanikha locality,Udmurtia

New scorpionflies, Asiachorista europaea sp. nov. and Petromantis udmurtica sp. nov. (Mecoptera: Permochoristidae), and new grylloblattids, Tshepanichoptera lacera gen. et sp. nov. (Grylloblattida: Aliculidae) and Miralioma urzhumica sp. nov. (Liomopteridae), are described from the Urzhumian of Udmurtia (Chepanikha locality). Liomopterites novissimus Aristov, 2004 (Liomopteridae) is redescribed.     

New Insects (Insecta: Trichoptera,Reculida, Eoblattida) from the Mesozoic of Asia

New caddisflies, reculid and eoblattid insects from the Mesozoic of Asia are described. Caddisflies of the families Philopotamidae (Mesoviatrix paradoxa gen. et sp. nov. and Kempia piotri gen. et sp. nov.) and Polycentropodidae (Polylongevus eskovi gen. et sp. nov.) from the Upper Jurassic–Lower Cretaceous of the Kempendyai locality in Yakutia are described. A short review and comparison of fossil members in these families are provided. New Gryllones insects, Shurabia tanga sp. nov., Sauk batkenicus gen. et sp. nov. from the Sauk Tan’ga locality (Lower Jurassic of Kyrgyzstan), Say kirgizicus gen. et sp. nov. from the Shurab III locality (Reculida: Geinitziidae; Lower or Middle Jurassic of Kyrgyzstan) Griphopteron iya sp. nov. (Eoblattida: Blattogryllidae) from Iya locality (Middle Jurassic of Russia) are described.     

New grylloblattids of the family Lemmatophoridae (Insecta: Grylloblattida) from the Permian of Russia

New grylloblattids of the family Lemmatophoridae (Insecta: Grylloblattida), Artinska sojanensis sp. nov., Sylvaprisca colorata sp. nov., and Uraloprisca lisca gen. et sp. nov., from the Soyana locality (Lower Kazanian, Arkhangelsk Region) and Paraprisca solikamskensis sp. nov. from the Tyul’kino locality (Ufimian, Perm Region) are described. Two species are transferred from the genus Paraprisca to Uraloprisca gen. nov.: Uraloprisca uralica (G. Zalessky, 1952), comb. nov. and Uraloprisca causaria (Novokshonov, 2000), comb. nov. (both from the Kungurian of the Perm Region). The evolution of the family is discussed.     

New Grylloblattids of the family Megakhosaridae (Insecta: Grylloblattida) from the Permian of Russia

New Megakhosaridae (Insecta: Grylloblattida) are described from the Permian of Russia: Ivakhosara pectinimedia gen. et sp. nov., Megakhosarina intricata sp. nov. and M. minuscula sp. nov. from the Lower Kazanian of Soyana (Arkhangelsk Region), M. magna sp. nov. from the Urzhumian of Chepanikha (Udmurtia), Kargalokhosara terraefossa gen. et sp. nov. from the Urzhumian of Kargala (Orenburg Region), and Alekhosara reticulata gen. et sp. nov. from the Severodvinian of Novo-Aleksandrovka (Orenburg Region).     

New and Little-known Nymphs of Gryllones (Insecta) from the Lower Permian of Russia

New nymphs of the gryllones insects (Insecta; Gryllones) Iblatta attrepida gen. et sp. nov. (Eoblattida incertae sedis) and Tshekardushka artenatis gen. et sp. nov. (Reculida incertae sedis) from the Lower Permian Chekarda locality (Kungurian Stage of Russia) are described. The nymph of Czekardia blattoides Martynov, 1940 (Eoblattida incertae sedis) from Chekarda is redescribed.     

New beetles (Insecta, Coleoptera) from Vyazniki locality, terminal Permian of European Russia

Recently collected fossil beetles from the families Asiocoleidae, Rhombocoleidae, Schizocoleidae, and Permosynidae are described from the locality Vyazniki, terminal for the Permian of European Russia. The findings are represented by isolated elytra, the position of which in the natural system it is often impossible to establish; they are, therefore, described in a formal system. The oryctocenosis is dominated by schizocoleids and permosynids, which start to play an important role at the very end of the Permian. They probably include members of Adephaga and Polyphaga, the principal groups of Mesozoic-Cenozoic beetles. Each of the other two families is represented by a single specimen.     

纺足目昆虫研究进展

纺足目昆虫又被称为足丝蚁,因为其具有独特的纺丝习性而被人所熟知;纺足目是昆虫纲中一个较小的类群,目前只有400余种被描述。本文简述了足丝蚁的形态特征和生物学特性;回顾了足丝蚁现生类群与化石研究历史及进展,概述了近年来足丝蚁的系统发育学研究进展,并提出研究该类群有待解决的问题,展望了纺足目未来的研究方向。     

New mayflies (Insecta: Ephemerida = Ephemeroptera) from the Upper Permian locality of Isady, Northern European Russia

Two new species of the family Protereismatidae, Alexandrinia vitta sp. nov. and A. ipsa sp. nov., are described from the Upper Permian locality of Isady (Severodvinian Stage, Poldarsa Formation). The third new species Misthodotes dubius sp. nov., belongs to the family Misthodotidae.     

Oldest webspinners from the Middle Jurassic of Inner Mongolia,China (Insecta: Embiodea)

The oldest webspinners, Sinembia rossi gen. et sp. nov. and Juraembia ningchengensis gen. et sp. nov. , are described in the new family Sinembiidae fam. nov. from the Middle Jurassic of Inner Mongolia, China. They differ from the Cretaceous and more recent Embiodea in several plesiomorphic characters, namely they have a long ovipositor, three‐segmented cerci, eyes situated on the posterolateral angles of the head, and the prothoracic prescutum is absent: these characters suggest habits that strongly differ from those of the recent taxa. The loss of the ovipositor and the reduction in the number of cerci can no longer be considered as synapomorphies of the ((Embiodea + Zoraptera) + Plecoptera) and (Embiodea + Zoraptera) clades, respectively.     

The first record of Kaltanidae (Insecta: Mecoptera: Kaltanidae) from the Permian of European Russia

Two new species and a new genus of scorpionflies of the family Kaltanidae, Pseudochorista occidentalis sp. nov. and Kamochorista novokshonovi gen. et sp. nov. (Mecoptera: Kaltanidae), are described from the Urzhumian of Udmurtia (Chepanikha locality). This is the first record of kaltanids on the Russian Platform.     

A new family of the order Grylloblattida (Insecta) from the Middle Permian of Russia

A new family, Ivapteridae fam. nov. (Insecta; Grylloblattida), is described from the Middle Permian locality of Soyana (Arkhangelsk Region; Kazanian Stage). It is most similar to Sojanoraphidiidae O. Martynova, 1952, differing from it in the subcostal field being traversed in the basal half of the wing by long, curved, and strongly oblique crossveins that form a double row of cells, the base of CuA being free, and CuA₁ thin compared to CuA₂. The new family is represented by a single species, Ivaptera sharovi, gen. et sp. nov. An overview of the modern system of the order Grylloblattida is included.