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1.
In the present study, 307 ovaries of eastern Baltic cod Gadus morhua callarias sampled during the prespawning and spawning season 2000 were analysed histologically to estimate the seasonal prevalence and intensity of atresia. The number of atretic oocytes per ovary was estimated using a combination of the physical disector method and volume fraction (Delesse principle). Atretic oocytes were observed in 32% of the ovaries. Prevalence of atresia was independent of female size, but increased significantly with declining female condition from prespawning and through the spawning stages. The relative intensity of atresia, i.e. number of atretic oocytes in relation to normally developed vitellogenic oocytes, was low amounting to 1·4% on average. Similar to prevalence, relative intensity of atresia differed significantly between maturity stages and increased with decreasing female condition. The population egg loss due to atresia amounted to 4·6% indicating that Baltic cod was performing close to maximum productivity, i.e. potential egg production. 相似文献
2.
利用遥感信息研究西藏地区主要植被年内和年际变化规律 总被引:6,自引:3,他引:6
利用1982~2000年NOAA-AVHRR月合成NDVI遥感资料和相关气象台站数据对我国西藏地区的稀疏草地、浓密草地和Tebit森林等主要植被的变化进行了初探。利用月合成NDVI的多年平均值分析了植被指数年内季节性变化规律及其与气候因子的关系,利用多年月合成NDVI的标准差描述了NDVI年际间波动情况。结果表明,在西藏地区,浓密草地和Tebit森林的NDVI植被指数年内变化规律呈明显的季节性,而稀疏草地则不明显;在年际变化方面,浓密草地月合成NDVI值波动幅度最大,Tebit森林次之,稀疏草地最小,且波动幅度较大的月份集中在NDVI值较高的植被生长季节6~10月份。 相似文献
3.
To build a better data foundation for recruitment models of north-east Arctic cod Gadus morhua the construction of fecundity models reflecting variation in the nutritional status of the fish was attempted. The models were based on fecundity time series covering 9 years within the period 1986–2004 and included both general and year-specific approaches. Initial data analysis revealed that the potential fecundity ( F P ) (standing stock of vitellogenic oocytes) was significantly reduced as the vitellogenic oocytes increased in size towards the start of spawning. Histological examination strongly indicated that this seasonal reduction was caused by atresia. Regression analysis showed that the F P was positively correlated to fish total length ( L T ) and the Fulton's condition factor ( K ). A multiple regression including data for all years using fish L T , K and mean oocyte diameter ( D O ) as independent predictors described the F P with an r 2 = 0·94. This was considerably higher than comparable univariate L T or mass-based regressions. These univariate regressions had fairly high r 2 values when split by years, but not as high as found for year-specific multiple regressions. An important application for individual-based fecundity models may be to generate outputs that can be fed into stock level fecundity and recruitment models. Overall, the multivariate models seemed to be the most accurate. The multivariate model including mean D O , however, also had the potential to correct for maturity and thus provide unbiased fecundity comparisons between years, stocks and locations. 相似文献
4.
Limburg KE Walther Y Hong B Olson C Storå J 《Proceedings. Biological sciences / The Royal Society》2008,275(1652):2659-2665
Combining Stone Age and modern data provides unique insights for management, extending beyond contemporary problems and shifting baselines. Using fish chronometric parts, we compared demographic characteristics of exploited cod populations from the Neolithic Period (4500 BP) to the modern highly exploited fishery in the central Baltic Sea. We found that Neolithic cod were larger (mean 56.4 cm, 95% confidence interval (CI)+/-0.9) than modern fish (weighted mean length in catch =49.5+/-0.2 cm in 1995, 48.2+/-0.2 cm in 2003), and older (mean ages=4.7+/-0.11, 3.1+/-0.02 and 3.6+/-0.02 years for Neolithic, 1995, and 2003 fisheries, respectively). Fishery-independent surveys in 1995 and 2003 show that mean sizes in the stock are 16-17 cm smaller than reflected in the fishery, and mean ages approximately 1-1.5 years younger. Modelled von Bertalanffy growth and back-calculated lengths indicated that Neolithic cod grew to smaller asymptotic lengths, but were larger at younger ages, implying rapid early growth. Very small Neolithic cod were absent and large individuals were rare as in modern times. This could be owing to selective harvests, the absence of small and large fish in the area or a combination. Comparing modern and prehistoric times, fishery selection is evident, but apparently not as great as in the North Atlantic proper. 相似文献
5.
Ernst M. von Rudloff 《Phytochemistry》1978,17(1):127-130
The composition of the leaf oil of Sitka spruce was determined and the quantitative variation within trees, within populations and amongst 3 southern, 5 central and 2 northern populations was recorded. No regional or geographic differences were found. In addition to the relatively large amounts of myrcene, piperitone, 1:8-cineole, and relatively low camphene and bornyl acetate percentages, the apparently unique isoamyl and isopentenyl isovalerate may serve to distinguish hybrids in areas of introgression with white spruce. 相似文献
6.
Steffen Funk Romain Frelat Christian Möllmann Axel Temming Uwe Krumme 《Journal of fish biology》2021,98(3):707-722
This study presents the diet composition of western Baltic cod Gadus morhua based on 3150 stomachs sampled year-round between 2016 and 2017 using angling, gillnetting and bottom trawling, which enhanced the spatio-temporal coverage of cod habitats. Cod diet composition in shallow areas (<20 m depth) was dominated by benthic invertebrate species, mainly the common shore crab Carcinus maneas. Compared to historic diet data from the 1960s and 1980s (limited to depth >20 m), the contribution of herring Clupea harengus decreased and round goby Neogobius melanostomus occurred as a new prey species. Statistical modelling revealed significant relationships between diet composition, catch depth, fish length and season. Generalized additive modelling identified a negative relationship between catch depth and stomach content weight, suggesting reduced food intake in winter when cod use deeper areas for spawning and during peak summer when cod tend to avoid high water temperatures. The results of this study highlight the importance of shallow coastal areas as major feeding habitats of adult cod in the western Baltic Sea, which were previously unknown because samples were restricted to deeper trawlable areas. The results strongly suggest that historic stomach analyses overestimated the role of forage fish and underestimated the role of invertebrate prey. Eventually, this study shows the importance of a comprehensive habitat coverage for unbiased stomach sampling programmes to provide a more reliable estimation of top predator diet, a key information for food web analyses and multispecies models. 相似文献
7.
A. N. Stroganov M. Bleil R. Oeberst A. V. Semenova H. Winkler 《Zeitschrift fur angewandte Ichthyologie》2018,34(3):527-534
Two cod stocks (western Baltic cod, WBC, and eastern Baltic cod, EBC) are managed in the Baltic Sea which is characterized by different main spawning areas and different main spawning periods. In this study we analyse the spatial and temporal occurrence of spawning individuals of both cod stocks in the main spawning grounds of the Baltic Sea based on eight microsatellite loci. Our results suggest that EBC (Gadus morhua callarias) has formed currently temporally stable, substantially homogeneous population not only in the Bornholm Sea (ICES SD: 25) but also in the Arkona Sea (ICES SD: 24). The presented analyses proved that EBC (G. m. callarias) can temporarily also spawn in the Belt Sea. 相似文献
8.
Recent environmental changes have influenced the ecology and biology of eastern Baltic cod. Declining somatic condition, maturation at smaller size and restricted size distribution of the population suggest that growth rates have decreased between the early 2000s and the 2010s. Extensive age estimation problems have until now precluded testing of this hypothesis. This study presents evidence for a decrease in somatic growth rate of Baltic cod. Temporal patterns of growth, condition and maturation were analysed based on two complementary analyses: length frequency mode progression derived from DATRAS bottom trawl survey data and known-age samples, where size at age was back-calculated from daily otolith growth patterns. In the known-age samples, growth was positively related to somatic condition at capture with maturity dependent differences. Immature individuals had experienced significantly lower growth and were in lower condition at capture than mature individuals. Growth rates in the known-age samples were estimated at 9.5, 7.8 and 5.7?cm per year for age classes 1, 2 and 3 respectively. Growth between age 2 and 3 decreased significantly from 8.8?cm in the 1997 year class to 7.6?cm in the 2010 year class. While the 2001 and 2004 known-age samples were representative for the population, the 2013 sample was biased towards individuals with a higher condition and growth. Complementary length frequency analysis following the length mode of fish from age 2 to age 3 confirmed growth estimates from the early 2000s, while suggesting a 37.5% lower growth in 2013 compared with 2005. 相似文献
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10.
In the distributional overlap volume of Baltic cod Gadus morhua and its prey, studied in the Bornholm Basin in the southern Baltic Sea, only a fraction of the sprat Sprattus sprattus population vertically overlapped with the Baltic cod population. Sprat occurred in the intermediate water, in the halocline and in the bottom water, while herring Clupea harengus and Baltic cod occurred exclusively in the halocline and in the bottom water. Only parts of the sprat population were hence accessible for Baltic cod, and only a fraction of the sprat had access to the Baltic cod eggs below the halocline. Baltic cod–clupeid overlap volumes appeared to be determined by salinity stratification and oxygenation of the bottom water. Hydrography time series were used to estimate average habitat volumes and overlap from July to September in 1958–1999. In the 1999 survey spawning Baltic cod had greater ratios of empty stomachs and lower average rations than non-spawning Baltic cod. The average ration for Baltic cod caught within 11· 4 m from the bottom (demersal) did not differ from the average ration of Baltic cod caught in shallower waters (pelagic), because spawning and non-spawning Baltic cod in both strata were caught at equal rates. The diet of the Baltic cod caught demersally contained more benthic invertebrates, especially Saduria entomon, but Baltic cod caught pelagically also had fresh benthic food in their stomachs, indicating vertical migration of individual fish. 相似文献
11.
Kate McQueen Monica Mion Annelie Hilvarsson Michele Casini Hans J. Olesen Karin Hüssy Krzysztof Radtke Uwe Krumme 《Journal of fish biology》2019,95(6):1486-1495
An aggregated sample of 925 Atlantic cod Gadus morhua collected by four countries in different regions of the Baltic Sea during different seasons were measured (total length, LT = 161–890 mm and weighed (mass, M = 45–6900 g) both before freezing and after defrosting. The cod were found to decrease significantly in both LT and M following death and frozen storage. There was an average (±SD) change in LT of −2.91% (±0.05%) following freezing, independent of starting LT. Total M changed by −2.65% (±0.14%), independent of starting mass. Shrinkage of LT and M did not differ significantly between 1 and 4 months frozen storage, though LT shrinkage was significantly greater after 1 or 4 months in the freezer compared with after 5 days. There was significant variation in LT and M shrinkage between regions of capture. A significant negative relationship between condition of cod and LT or M change was also observed. Equations to back-calculate fresh LT and M from thawed LT, M and standard length (LS), gutted LT, gutted LS and gutted M are provided. 相似文献
12.
采用水培试验方法,研究了不同浓度镉(Cd)(0、5、10、25和50 mg·L-1) 胁迫条件下藤本植物金银花的生长和生理特性.结果表明: 与对照相比,Cd胁迫对金银花的生长未造成明显影响,在5~50 mg·L-1 Cd处理下,其生物量无明显差异(P>0.05),在低浓度Cd(5 mg·L-1)处理下生物量有所增加,叶、根生物量和总生物量分别增加了2.88%、2.33%和1.25%,说明金银花对Cd具有较强的抗性.在低浓度Cd胁迫下,植物各器官的含水量和可溶性蛋白含量均有所降低,而根系和叶片的丙二醛含量分别增加51.90%和23.07%,叶绿素和类胡萝卜素含量则增加15.87%和24.89%,超氧化物歧化酶活性也显著增强.随着Cd浓度的增高,金银花体内的叶绿素和类胡萝卜素含量,以及超氧化物歧化酶活性均有所降低. 相似文献
13.
Cod Gadus morhua population in the Barents Sea was found to be spatially structured with regard to length-at-age. Results were based on data collected during research surveys in the Barents Sea between 1982 and 1997. The identified spatial structure was most pronounced for age groups 2–4 years and decreased for the older age groups with higher potential for migration. A positive linear correlation between mean length-at-age and mean geographical temperature was established for age groups 2–4 years. This correlation was shown to be strongest when based on mean temperatures during 3 year periods ending with the year of capture. The spatial structure in length-at-age was shown to follow the temperature gradient of the Barents Sea. A large part of the observed area effects could be explained by temperature variation between areas. Evidence is also presented which indicates that the predictability and sensitivity of the dependence of length-at-age on temperature increases under extreme environmental conditions, i.e. in the northern and eastern areas of the Barents Sea. 相似文献
14.
BACKGROUND AND AIMS: Stem respiration of trees is a major, but poorly assessed component of the carbon balance of forests, and important for geo-chemistry. Measurements are required under naturally changing seasonal conditions in different years. Therefore, intra- and inter-annual carbon fluxes of stems in forests were measured continuously from April to November in three consecutive years. METHODS: Stem respiratory CO2 fluxes of 50-year-old Scots pine (Pinus sylvestris) trees were continuously measured with a CO2 analyser, and, concomitantly, stem circumference, stem and air temperature and other environmental factors and photosynthesis, were also measured automatically. KEY RESULTS: There were diurnal, seasonal and inter-annual changes in stem respiration, which peaked at 1600 h during the day and was highest in July. The temperature coefficient of stem respiration (Q10) was greater during the growing season than when growth was slow or had stopped, and more sensitive to temperature in the growing season. The annual Q10 remained relatively constant at about 2 over the three years, while respiration at a reference temperature of 15 degrees C (R15) was higher in the growing than in the non-growing season (1.09 compared with 0.78 micromol m(-2) stem surface s(-1)), but was similar between the years. Maintenance respiration was 76 %, 82 % and 80 % of the total respiration of 17.46, 17.26 and 19.35 mol m2 stem surface in 2001, 2002 and 2003, respectively. The annual total stem respiration of the stand per unit ground area was 75.97 gC m(-2) in 2001 and 74.28 gC m(-2) in 2002. CONCLUSIONS: Stem respiration is an important component in the annual carbon balance of a Scots pine stand, contributing 9 % to total carbon loss from the ecosystem and consuming about 8 % of the carbon of the ecosystem gross primary production. Stem (or air) temperature was the most important predictor of stem carbon flux. The magnitude of stem respiration is modified by photosynthesis and tree growth. Solar radiation indirectly affects stem respiration through its effect on photosynthesis. 相似文献
15.
Thorsten Blenckner Marcos Llope Christian M?llmann Rudi Voss Martin F. Quaas Michele Casini Martin Lindegren Carl Folke Nils Chr. Stenseth 《Proceedings. Biological sciences / The Royal Society》2015,282(1803)
Overfishing of large predatory fish populations has resulted in lasting restructurings of entire marine food webs worldwide, with serious socio-economic consequences. Fortunately, some degraded ecosystems show signs of recovery. A key challenge for ecosystem management is to anticipate the degree to which recovery is possible. By applying a statistical food-web model, using the Baltic Sea as a case study, we show that under current temperature and salinity conditions, complete recovery of this heavily altered ecosystem will be impossible. Instead, the ecosystem regenerates towards a new ecological baseline. This new baseline is characterized by lower and more variable biomass of cod, the commercially most important fish stock in the Baltic Sea, even under very low exploitation pressure. Furthermore, a socio-economic assessment shows that this signal is amplified at the level of societal costs, owing to increased uncertainty in biomass and reduced consumer surplus. Specifically, the combined economic losses amount to approximately 120 million € per year, which equals half of today''s maximum economic yield for the Baltic cod fishery. Our analyses suggest that shifts in ecological and economic baselines can lead to higher economic uncertainty and costs for exploited ecosystems, in particular, under climate change. 相似文献
16.
Martin Lindegren Christian M?llmann Anders Nielsen Keith Brander Brian R. MacKenzie Nils Chr. Stenseth 《Proceedings. Biological sciences / The Royal Society》2010,277(1691):2121-2130
Good decision making for fisheries and marine ecosystems requires a capacity to anticipate the consequences of management under different scenarios of climate change. The necessary ecological forecasting calls for ecosystem-based models capable of integrating multiple drivers across trophic levels and properly including uncertainty. The methodology presented here assesses the combined impacts of climate and fishing on marine food-web dynamics and provides estimates of the confidence envelope of the forecasts. It is applied to cod (Gadus morhua) in the Baltic Sea, which is vulnerable to climate-related decline in salinity owing to both direct and indirect effects (i.e. through species interactions) on early-life survival. A stochastic food web-model driven by regional climate scenarios is used to produce quantitative forecasts of cod dynamics in the twenty-first century. The forecasts show how exploitation would have to be adjusted in order to achieve sustainable management under different climate scenarios. 相似文献
17.
The study of hybrid zones is central to our understanding of the genetic basis of reproductive isolation and speciation, yet very little is known about the extent and significance of hybrid zones in marine fishes. We examined the population structure of cod in the transition area between the North Sea and the Baltic Sea employing nine microsatellite loci. Genetic differentiation between the North Sea sample and the rest increased along a transect to the Baltic proper, with a large increase in level of differentiation occurring in the Western Baltic area. Our objective was to determine whether this pattern was caused purely by varying degrees of mechanical mixing of North Sea and Baltic Sea cod or by interbreeding and formation of a hybrid swarm. Simulation studies revealed that traditional Hardy-Weinberg analysis did not have sufficient power for detection of a Wahlund effect. However, using a model-based clustering method for individual admixture analysis, we were able to demonstrate the existence of intermediate genotypes in all samples from the transition area. Accordingly, our data were explained best by a model of a hybrid swarm flanked by pure nonadmixed populations in the North Sea and the Baltic Sea proper. Significant correlation of gene identities across loci (gametic phase disequilibrium) was found only in a sample from the Western Baltic, suggesting this area as the centre of the apparent hybrid zone. A hybrid zone for cod in the ecotone between the high-saline North Sea and the low-saline Baltic Sea is discussed in relation to its possible origin and maintenance, and in relation to a classical study of haemoglobin variation in cod from the Baltic Sea/Danish Belt Sea, suggesting mixing of two divergent populations without interbreeding. 相似文献
18.
19.
Kc-cells from Drosophila melanogaster, grown under serum-free conditions, produce two beta-hexosaminidases and secrete these enzymes into the medium. The two enzymes were separated by DEAE-exchange chromatography. According to their substrate specificities one enzyme is a beta-N-acetyl-D-glucosaminidase (E.C.3.2.1.30), the other one a beta-N-acetyl-D-hexosaminidase (E.C.3.2.1.52). The beta-N-acetyl-D-glucosaminidase is predominant in the medium, the beta-N-acetyl-D-hexosaminidase within the cells. The Km values for the substrates pNP-GlcNAc, pNP-GalNAc, and (GlcNAc)2 are 0.8, 16.73, and 1.67 mM for the beta-N-acetyl-D-glucosaminidase and 0.24, 0.44, and 0.2 mM for the beta-N-acetyl-D-hexosaminidase. Both enzymes are inhibited by the products and the beta-N-acetyl-D-glucosaminidase is also inhibited stereospecifically by the substrates pNP-GlcNAc and (GlcNAc)2. Both enzymes are inhibited in a partial competitive way by acetamidolactones, the Kis being as low as 0.1 microM. 相似文献
20.
Frederique Olivier Jan A. van Franeker Jeroen C. S. Creuwels Eric J. Woehler 《Polar Biology》2005,28(9):687-699
Demographic parameters were estimated for snow petrels Pagodroma nivea nesting at the study colony of Reeve Hill near Casey station, Antarctica between 1984 and 2003. Average breeding success for the colony varied from 18.2% to 76.5%. Breeding effort, hatching and fledging success were subject to a high interannual variability. We examined the influence of regional sea-ice extent on the breeding performance of snow petrels at Reeve Hill. Fewer birds were breeding when sea-ice had been extensive during April–May. Overall breeding success and fledging success were improved during years with extensive sea-ice cover in winter. Successful breeding effort and breeding success were depressed when there was extensive sea-ice cover during January–February. Sea surface temperatures also correlated to snow petrel breeding performance parameters. Previous work showed that large-scale climatic events (ENSO, Antarctic circumpolar wave) and the related sea-ice cover around the Antarctic might affect the lower trophic levels of the marine environment and consequently food availability for snow petrels. A comparison with the long-term study conducted at Ile des Pétrels (Terre Adélie) suggests that despite similarities in the underlying biological processes that control snow petrel breeding performance, the nature of the correlation of large-scale environmental factors with breeding performance differs substantially between the two colonies, probably because of the confounding effects of other environmental factors acting at a local scale (local weather, nest quality), which also affect bird body condition. 相似文献