Index to Vegetatio,volumes 51–73, and some notes on the position of the journal

An index is presented of 375 articles which appeared in Vol. 51–73 of Vegetatio. A Source index is followed by a Keyword index, a Discipline index, a Vegetation type index, a Geographical index, and a Latin name index for taxa and syntaxa. The codes used for discipline, vegetation type and geographic area are the same as those used in the Index Vol. 1–50. After a short introduction a history of the journal is presented, followed by some statistics on the origin of the authors and the type of article, for seven periods covering the entire history of the journal since 1948. Finally the codes are elucidated.     

Vegetation mapping in Italy

A brief history of the vegetation mapping in Italy is described. The principal types of maps are physiognomical, of vegetation belts, of vegetation series, phytosociological, of dynamical tendencies in vegetation, ecological, of potential vegetation and environmental. The first botanical map of whole Italy was made by Fiori in 1908 and it is of physiognomical type. The last one was realized by Pedrotti in 1989 and represents the natural potential vegetation (in 1:1 500 000 scale).This report was oral presented at the V^th International Congress of Ecology (August 23–30, 1990 - Yokohama, Japan).     

Organic carbon and nitrogen losses influenced by vegetation removal in a semiarid mediterranean soil

A reduction in plant cover can lead to an increase in the erosionprocesses that diminish soil quality. Any rise in temperature resulting frompredicted climate changes may aggravate this effect, particularly in semiaridMediterranean areas. Bearing this in mind, the capacity of a soil to preserveorganic matter becomes very important if the soil is to maintain its physicaland chemical properties. Soil organic carbon and nitrogen changes wereevaluatedin a non-disturbed (with natural vegetation) and a disturbed (all vegetationmanually clipped to ground level) pine system. Nine years after vegetationremoval significant differences (p < 0.01) were found in the soil organiccarbon content between plots (top 20 cm), but not in totalnitrogen. In the disturbed plot 0.0232 Mg ha^–1y^–1 of soil organic carbon were lost through erosionand4.30 Mg ha^–1 y^–1 throughmineralization. In the first 48 months after vegetation removal the soilorganiccarbon content fell from 40.3 to 28.0 g kg^–1. Inthe last 60 months of the experiment the amount of organic carbon in the soilfell from 28.0 to 27.7 g kg^–1. This result wasmainly attributable to the intense oxidization, which took place during thefirst 60 months, of organic matter linked to the coarse soil mineral fraction.Up to the 72^nd month the losses by erosion were a total of 532.7g, which rose to 1284.4 g by the end of theexperiment(108 months). The effect of vegetation removal in a Mediterranean semiarid arealeads to a rapid decline in the amount of organic carbon stored in the soil.Such perturbation is irreversible if left to nature.     

Studies on the vegetation history of Lake Varna region,northern Black Sea coastal area of Bulgaria

On the basis of pollen analytical investigations of two cores from Lake Varna and Lake Beloslav, the vegetation history of the Lake Varna region is traced back to the beginning of the 6th millennium B.C. A two-fold zoning system is used whereby the pollen diagrams are divided into pollen zones, based on tree migration patterns, and settlement periods. Pollen zone 1 is characterised by the absence of Carpinus betulus and Fagus sylvatica. The spread of hornbeam started at ca. 6500 B.C. (pollen zone 2) and beech at ca. 6200 B.C. (pollen zone 3), the latter being the last tree to spread into the region and considerably enrich the forests of the study area. Of the many pollen taxa representing plants that are favoured by open habitats and hence potentially indicative of human impact, only a few taxa are regarded as reliable indicators of human activity. These include above all Triticum-type, and also Secale and to a certain extent Plantago lanceolata, Rumex and Polygonum aviculare. The spatial pattern of settlements is somewhat different in the areas represented by the two profiles. At both sites the first period of settlement occurred during the 6th millennium B.C. (early Neolithic). After the Neolithic period, the main settlement periods of the Eneolithic and the Early and Middle Bronze Age are recorded. On the other hand, land-use history during the Greek and Roman periods is poorly recorded. Studies on the stratigraphy, diatoms and molluscs indicate that the sixth Black Sea transgression (6500–5800 B.C.), which reached –10 m, had considerable influence on the limnological environment.     

Lake Ladoga Region: human impacts and recent environmental changes

This paper presents a landscape-ecological analysis of Ladoga Region, a 30 000 km² territory around Lake Ladoga, which is the largest lake in Europe. The Region includes the contact zone between the crystalline bedrock of Baltic Shield and the sedimentary rocks of the East European plain, and the transition from Boreonemoral to Southern Boreal (Taiga) forest vegetation; it is characterized by abundance of lakes. The Region is divided into 17 discrete landscape areas that are classified in 6 groups.The author uses the landscape-dynamcal approach to the analysis of recent (from 2–3 to 40–50 yr) environmental changes in the region, caused by human activities: forest felling, fires, peat excavation, agriculture, recreational development etc. The following processes are observed: vegetation successions after clearcutting and selective logging; effects of forest fires on the spreading and conservation of pine forests; consequences of peatland drainage; landscape effects of collective farming and gardening; forest succession from pine and deciduous tree dominance towards spruce dominance (observable in all landscape regions); natural vegetation succession on former Finnish arable lands (two types of scenarios: forest succession or development of sedge meadows).     

Vegetation and climatic history of southwest Africa: A marine palynological record of the last 300,000 years

A continuous palynological record from the marine core GeoB1016-3 from the Angola Basin reveals the regional vegetation and climate history of the last 300 ka. Pollen and spores found at the studied site have their source areas in the different vegetation zones of the adjacent part of the West African continent. Those vegetation zones comprise tropical rain forest, coastal mangrove swamp, Miombo woodland, dry forest, Afromontane forest, desert and semi-desert. The main pollen transport agent is the southeast trade wind system. Ocean currents also partly play a role in transporting pollen and spores. During the interglacial periods, ocean currents also transported palynomorphs southward. During the glacial periods, increased trade winds are indicated by high influx of pollen and spores and high pollen percentages of Poaceae and taxa from desert and semidesert vegetations. Reconstruction of the geographical position of palaeo-vegetation zones shows that the northern boundary of the Namib Desert did not move north of 12°S during the last 300 ka. This implies that northward shifts of the Angola-Benguela Front did not pass the latitude of 12°S.     

Accelerated vegetation succession but no hydrological change in a boreal fen during 20 years of recent climate change

Northern mires (fens and bogs) have significant climate feedbacks and contribute to biodiversity, providing habitats to specialized biota. Many studies have found drying and degradation of bogs in response to climate change, while northern fens have received less attention. Rich fens are particularly important to biodiversity, but subject to global climate change, fen ecosystems may change via direct response of vegetation or indirectly by hydrological changes. With repeated sampling over the past 20 years, we aim to reveal trends in hydrology and vegetation in a pristine boreal fen with gradient from rich to poor fen and bog vegetation. We resampled 203 semi‐permanent plots and compared water‐table depth (WTD), pH, concentrations of mineral elements, and dissolved organic carbon (DOC), plant species occurrences, community structure, and vegetation types between 1998 and 2018. In the study area, the annual mean temperature rose by 1.0°C and precipitation by 46 mm, in 20‐year periods prior to sampling occasions. We found that wet fen vegetation decreased, while bog and poor fen vegetation increased significantly. This reflected a trend of increasing abundance of common, generalist hummock species at the expense of fen specialist species. Changes were the most pronounced in high pH plots, where Sphagnum mosses had significantly increased in plot frequency, cover, and species richness. Changes of water chemistry were mainly insignificant in concentration levels and spatial patterns. Although indications toward drier conditions were found in vegetation, WTD had not consistently increased, instead, our results revealed complex dynamics of WTD as depending on vegetation changes. Overall, we found significant trend in vegetation, conforming to common succession pattern from rich to poor fen and bog vegetation. Our results suggest that responses intrinsic to vegetation, such as increased productivity or altered species interactions, may be more significant than indirect effects via local hydrology to the ecosystem response to climate warming.     

Effect of Bursaphelenchus xylophilus on the Assimilation and Translocation of 14 C in Pinus sylvestris

The effect of wound, wound + water, wound + Bursaphelenchus xylophilus culture filtrate, or wound + lethal B. xylophilus doses on the assimilation and translocation of ¹⁴C by 8-month-old Pinus sylvestris seedlings was tested. In two separate experiments, pine seedlings were exposed to 28.35 μCi of ¹⁴CO₂ for 20 minutes below or above (to the pine shoot leader) the point of nematode inoculation. After 2 and 4 hours of dark adaptation, 80% ethanol soluble ¹⁴C tissue extracts were determined by liquid scintillation counting. Nematode infection significantly (P = 0.05) decreased ¹⁴C assimilation. Treatments translocated less than 6% of the total amount of the fixed ¹⁴C and translocation generally decreased with increasing size of nematode inoculum. However, infected pines translocated a greater proportion of the amount of ¹⁴C fixed per gram of exposed nematode-plant tissue than did the control pines. The lower levels of photoassimilate entering the plant system probably resulted in a reduced metabolic capacity in B. xylophilus-infected pine seedlings. The effect on photosynthesis could be one of the key factors leading to death of pines through starvation, and it is possible that it was preceded by an effect on related physiological processes such as water uptake.     

A study on relations of vegetation,climate and soils in Shanxi province,China

Relationships of vegetation, climate and soils in Shanxi plateau wereanalyzed by use of Canonical Correspondence Analysis (CCA). Shanxi province,located at 34°35–40°43 N, 110°15–114°33 E, was divided into a series of rectangular districts of30 latitude by 20 longitude. Areas of vegetation formations and soil types ineach district were measured carefully using fine grain on the vegetation andsoil maps of Shanxi. Climatic data were mean values of 25 years records in eachdistrict. Three data matrices of climate, vegetation and soil were combined byCCA. The results showed that the distribution of vegetation is closely relatedto the variety of climates and to soils distribution.     

Action of Phytochrome During Prechilling of Amaranthus retroflexus L. Seeds

Dark germination of Amaranthus retroflexus L. seeds at 35° increased after several days of prechilling at 20° or lower. Irradiation with far-red light for short periods during the early hours of a prechilling period at 10° inhibited subsequent dark germination at 35°. The inhibition was completely reversible with red light. Far-red irradiation in the latter part of the prechilling period was less effective. Increased dark germination of A. retroflexus seeds following a prechilling period at 20° or less is attributed to action of preexistent P_FR, the far-red absorbing form of phytochrome, within the seeds. Inactivation of P_FR was found to proceed ca. 4 times more rapidly at 25° than at 20°. Failure of imbibition temperatures above 20° to increase dark germination of A. retroflexus seeds is attributed to the rapid thermal reversion of pre-existent P_FR. We suggest that the action of prechilling (layering) on many other seed kinds arises in a similar way.     

Effects of Forest Age on Soil Autotrophic and Heterotrophic Respiration Differ between Evergreen and Deciduous Forests

We examined the effects of forest stand age on soil respiration (SR) including the heterotrophic respiration (HR) and autotrophic respiration (AR) of two forest types. We measured soil respiration and partitioned the HR and AR components across three age classes ∼15, ∼25, and ∼35-year-old Pinus sylvestris var. mongolica (Mongolia pine) and Larix principis-rupprechtii (larch) in a forest-steppe ecotone, northern China (June 2006 to October 2009). We analyzed the relationship between seasonal dynamics of SR, HR, AR and soil temperature (ST), soil water content (SWC) and normalized difference vegetation index (NDVI, a plant greenness and net primary productivity indicator). Our results showed that ST and SWC were driving factors for the seasonal dynamics of SR rather than plant greenness, irrespective of stand age and forest type. For ∼15-year-old stands, the seasonal dynamics of both AR and HR were dependent on ST. Higher Q₁₀ of HR compared with AR occurred in larch. However, in Mongolia pine a similar Q₁₀ occurred between HR and AR. With stand age, Q₁₀ of both HR and AR increased in larch. For Mongolia pine, Q₁₀ of HR increased with stand age, but AR showed no significant relationship with ST. As stand age increased, HR was correlated with SWC in Mongolia pine, but for larch AR correlated with SWC. The dependence of AR on NDVI occurred in ∼35-year-old Mongolia pine. Our study demonstrated the importance of separating autotrophic and heterotrophic respiration components of SR when stimulating the response of soil carbon efflux to environmental changes. When estimating the response of autotrophic and heterotrophic respiration to environmental changes, the effect of forest type on age-related trends is required.     

A vegetation map of The Netherlands,based on the relationship between ecotopes and types of potential natural vegetation

Summary The method of mapping the vegetation on scale 1: 200,000 and the starting points in relation to the potential natural vegetation and ecotopes, are discussed.In view of the planological background of this study, some restrictions have been added to the concept of potential natural vegetation, concerning the period of development and the human influence.The relationship between soil, ground water and vegetation was studied, which resulted in the map of the potential natural vegetation.Each type of potential natural vegetation stands for a series of vegetation types on the same site. Seven main series, with a number of sub-series are distinguished. Within each vegetation series the plant communities have been spread over five groups, according to their structure and naturalness.Ecotopes and ecotope complexes are considered as landscape ecological units. A list of ecotopes was obtained by interpreting topographical maps and by inventory data.The actual vegetation was mapped by estimating the size of the ecotopes within the separate areas. It was expressed in a five figure code for the five groups from the vegetation and ecotopes is combined into the vegetation map of The Netherlands.Interpretation problems, some of them specific for The Netherlands, are discussed and some remarks are made on the necessity of further research.Contribution to the Symposium on Plant Species and Plant Communities, held at Nijmegen, 11–12 November 1976, on the occasion of the 60th birthday of Professor Victor Westhoff.Nomenclature follows Heukels-van Ooststroom, Flora van Nederland, 18e druk, 1975, Wolters-Noordhoff, Groningen; nomenclature of syntaxa follows Westhoff & den Held (1969)     

Altitudinal changes in composition and structure of mountain-temperate vegetation: a case study from the Western Carpathians

Changes in composition and structure of plant communities in relation tothe soil and snow cover variation were analyzed along an altitudinal transect(1150–1750 m) from the mountain-temperate forests to a woodyshrub community and alpine meadows on Mt Velký Gápel', Slovakia.The soils below the treeline (1510 m) had a more developedorganic layer above the mineral substratum. Generally, soil depth decreased asthe altitude increased, although the maximum values were recognized at a middlealtitude in a beech stand. Snow was redistributed by westerly winds from theridgeline down to the upper forest margin. Mean snow depth decreased withaltitude up to almost snow-free sites around the summit. In the 48 plots at 16sites we recorded 118 taxa including 6 tree, 7 shrub, 18 grass, 42 herb, 5fern,25 moss and 15 lichen species. The species diversity showed no distinctrelationship to altitude but declined with canopy consolidation. The TWINSPANfloristic classification distinguished five groups of community typescharacterised by different dominants, and a further three clusters of samplesfrom transition zones. Horizontal compositional heterogeneity increased inareaswhere trees were aggregated and tree basal area was smaller. Vegetationcomposition became more patchy at open-canopy Acerpseudoplatanus–Abies alba mixed forest at 1150 m,in Picea abies forest limit 1470 m, andin Pinus mugo krummholz at 1590 m. Speciesturnover of the entire transect was 6.1 half-changes as estimated by DCA.Despite this heterogeneity, none of the 15 elevational bands had significantaggregation of species' limits. Vegetation varied continuously, with individualspecies overlapping in transition zones delimited by dominant taxa. Thecoincident aggregation of up-slope and down-slope boundaries was found at abelt1430–1510 m. This discrete ecotone corresponds to a shiftfrom the closed coniferous forest to P. mugo krummholz.Thesecond inherent up-slope boundary aggregation indicated the P.mugo krummholz – alpine meadow vegetation transition at1700 m. Spatial analysis (K-function) of eight forest plots(0.12 ha each) showed that at lower elevation, adult trees of thebroad-leaf forest were closer to a random arrangement while at higherelevation,trees of evergreen coniferous stands became aggregated toward the forest limitwith the highest intensity from 2 to 4 m. Altitudinal gradient andrelated factors explained 35% of the variance in vegetation data.Canonical correspondence analysis also showed that main vegetation changesabovethe treeline area were associated with the topographic pattern of pine shrubsand snow cover.     

Scots pine growing on forested mires in Abernethy Forest,Strathspey, Scotland

Summary

A substantial proportion of the Abernethy Forest Reserve has Scots pine (Pinus sylvestris) growing on the surfaces of a variety of mires. The hydrology of the mires has been affected by drainage and peat cutting but this area is unusual in having had a long period of protection from grazing by domestic stock. There are three main types of pine populations found on these mires. Woodland bog comprises predominantly bog vegetation with abundant pine seedlings due to the heavy seed rain from the surrounding woodland. Only a few very small trees survive, which are stunted, heavily diseased and have very low seed production. Wooded bog also comprises predominately bog vegetation but there are scattered mature trees of a moderate height with an open canopy. The trees are fertile and can form uneven aged stands with regeneration. Bog woodland is a predominantly woodland vegetation with tall, dense tree cover on deep peat. The trees are well grown with a dense canopy. A few remnants of bog vegetation remain in the ground flora although most have been replaced by woodland bryophytes and shrubs. Each of these three types is described and their development is discussed.     

A rare 300 kilometer dispersal by an adult male white‐tailed deer

Despite the key roles that dispersal plays in individual animal fitness and meta‐population gene flow, it remains one of the least understood behaviors in many species. In large mammalian herbivores, dispersals might span long distances and thereby influence landscape‐level ecological processes, such as infectious disease spread. Here, we describe and analyze an exceptional long‐distance dispersal by an adult white‐tailed deer (Odocoileus virginianus) in the central United States. We also conducted a literature survey to compare the dispersal to previous studies. This dispersal was remarkable for its length, duration, and the life history stage of the dispersing individual. Dispersal is typical of juvenile deer seeking to establish postnatal home ranges, but this dispersal was undertaken by an adult male (age = 3.5). This individual dispersed ~300 km over a 22‐day period by moving, on average, 13.6 km/day and achieving a straight‐line distance of ~215 km, which was ~174 km longer than any other distance recorded for an adult male deer in our literature survey. During the dispersal, which occurred during the hunting season, the individual crossed a major river seven times, an interstate highway, a railroad, and eight state highways. Movements during the dispersal were faster (mean = 568.1 m/h) and more directional than those during stationary home range periods before and after the dispersal (mean = 56.9 m/h). Likewise, movements during the dispersal were faster (mean = 847.8 m/h) and more directional at night than during the day (mean = 166.4 m/h), when the individual frequently sheltered in forest cover. This natural history event highlights the unpredictable nature of dispersal and has important implications for landscape‐level processes such as chronic wasting disease transmission in cervids. More broadly, our study underscores how integrating natural history observations with modern technology holds promise for understanding potentially high impact but rarely recorded ecological events.     

Accumulation,Allocation, and Risk Assessment of Polycyclic Aromatic Hydrocarbons (PAHs) in Soil-Brassica chinensis System

Farmland soil and leafy vegetables accumulate more polycyclic aromatic hydrocarbons (PAHs) in suburban sites. In this study, 13 sampling areas were selected from vegetable fields in the outskirts of Xi’an, the largest city in northwestern China. The similarity of PAH composition in soil and vegetation was investigated through principal components analysis and redundancy analysis (RDA), rather than discrimination of PAH congeners from various sources. The toxic equivalent quantity of PAHs in soil ranged from 7 to 202 μg/kg d.w., with an average of 41 μg/kg d.w., which exceeded the agricultural/horticultural soil acceptance criteria for New Zealand. However, the cancer risk level posed by combined direct ingestion, dermal contact, inhalation of soil particles, and inhalation of surface soil vapor met the rigorous international criteria (1×10⁻⁶). The concentration of total PAHs was (1052±73) μg/kg d.w. in vegetation (mean±standard error). The cancer risks posed by ingestion of vegetation ranged from 2×10⁻⁵ to 2×10⁻⁴ with an average of 1.66×10⁻⁴, which was higher than international excess lifetime risk limits for carcinogens (1×10⁻⁴). The geochemical indices indicated that the PAHs in soil and vegetables were mainly from vehicle and crude oil combustion. Both the total PAHs in vegetation and bioconcentration factor for total PAHs (the ratio of total PAHs in vegetation to total PAHs in soil) increased with increasing pH as well as decreasing sand in soil. The total variation in distribution of PAHs in vegetation explained by those in soil reached 98% in RDA, which was statistically significant based on Monte Carlo permutation. Common pollution source and notable effects of soil contamination on vegetation would result in highly similar distribution of PAHs in soil and vegetation.     

Simulating the impacts of southern pine beetle and fire on the dynamics of xerophytic pine landscapes in the southern Appalachians

Question: Can fire be used to maintain Yellow pine (Pinus subgenus Diploxylon) stands disturbed by periodic outbreaks of southern pine beetle? Location: Southern Appalachian Mountains, USA. Methods: We used LANDIS to model vegetation disturbance and succession on four grids representative of xeric landscapes in the southern Appalachians. Forest dynamics of each landscape were simulated under three disturbance scenarios: southern pine beetle, fire, and southern pine beetle and fire, as well as a no disturbance scenario. We compared trends in the abundance of pine and hardwood functional types as well as individual species. Results: Yellow pine abundance and open woodland conditions were best maintained by a combination of fire and southern pine beetle disturbance on both low elevation sites as well as mid‐elevation ridges & peaks. On mid‐elevation SE‐W facing slopes, pine woodlands were best maintained by fire alone. Conclusions: Our simulations suggest that fire can help maintain open pine woodlands in stands affected by southern pine beetle outbreaks.     

The formation of “mega‐flocks” depends on vegetation structure in montane coniferous forests of Taiwan

A mixed‐species bird flock is a social assemblage where two or more bird species are moving together while foraging and might benefit from increased foraging efficiency and antipredator vigilance. A “mega‐flock,” which includes flocking species from different vegetation strata, often exhibits high species diversity. Mechanisms for the formation of mega‐flocks have not yet been explored. In this study, we evaluated the influence of vegetation structure and bird species diversity in driving the occurrence of mega‐flocks. We investigated the composition of mixed‐species flocks, local bird communities, and vegetation structure in five vegetation types of two high‐elevation sites in central Taiwan. Mega‐flocks occurred more frequently in pine woodland than later successional stages of coniferous forests. However, species richness/diversity of local bird communities increased along successional stages. Therefore, vegetation variables exhibit more influence on the occurrence of mega‐flocks than local bird communities. Besides foliage height diversity, understory coverage also showed positive effects on flock size of mixed‐species flocks. Our results indicated that pine woodlands with more evenly distributed vegetation layers could facilitate the interactions of canopy and understory flocks and increase the formation of mega‐flocks and thus the complexity of mixed‐species flocks.     

Kinetics of the Daily Rate of Photosynthesis at Low Temperatures for two Conifers

The daily course of photosynthesis at low temperatures in 2 coniferous species, Pinus ponderosa Laws., and Pseudotsuga menziesii (Mirb.) Franco, were studied using controlled environment facilities. After having been grown at a 23° day, and 19° night for a year, seedlings were acclimatized for 4 months to either a 3°, 7° or 11° day all under 1200 ft-c of light and followed by a 16-hour night at 3°. Measurement of photosynthesis at 1200 ft-c revealed 3 separate responses. First, the rapidity at which the plants attained their maximum photosynthesis when the lights were turned on depended upon the species, the current temperature, and the previous temperature condition to which the plants had become acclimatized. The warmer the day temperature the sooner the daily maximum was reached. Second, fluctuations in the rate of photosynthesis during the day varied with the species and the day temperature. Photosynthesis in both fir and pine kept at an 11° day and pines kept at a 7° day attained a daily peak rate followed by a decline. This decline occurred even though temperature and light were kept constant, the CO₂ level was returned to 320 ppm from 290 ppm, and the plants were kept well watered. At a 3° day neither species showed this decline. Third, a plant transferred to another temperature acquired a new stable daily photosynthetic pattern. The number of days required for stabilization depended upon the previous temperature history of the plant. The adjustment rate was faster when the temperature was raised than when it was lowered.     

Palaeoecological evidence for Holocene vegetation, climate and land-use change in the low Don basin and Kalmuk area, southern Russia

Two¹⁴C-dated pollen profiles from mires in the steppe belt of southern Russia are presented. On the basis of these and data from earlier investigations, the Holocene forest history of the southern part of Russia and Ukraine is reconstructed. The steppe belt is very sensitive to climatic oscillations and, in particular, to changes in evapotranspiration. The most favourable climate occurred between 6000 and 4500 B.P. (6800–5200 cal. B.P.), when forest attained its maximum extent in the steppe belt. The period 4500–3500 B.P. (5200–3800 cal. B.P.) was characterised by drier climate with the most arid phase occurring between 4200–3700 B.P. (4700–4000 cal. B.P.). During arid phases, the area under forest and also peat accumulation rates declined. Subsequently, a number of less pronounced climatic oscillations occurred such as in the period 3400/3300–2800 B.P. (3600/3500–2900 cal. B.P.) when there was a return to more humid conditions. During the last 2500 years, the vegetation cover of the steppe belt in southern Russia and Ukraine took on its present-day aspect. On the one hand, there is close correlation between the Holocene vegetation history of southern Russia and Ukraine and, on the other hand, the steppe belt of Kazakhstan and transgressions in the Caspian sea. Human impact on the natural vegetation became important from the Bronze Age onwards (after 4500 B.P.; 5200 cal. B.P.). Particular attention is given to the history of Scots pine (Pinus sylvestris), which had a much wider distribution in the southern part of eastern Europe in the early Holocene. The reduction in range during recent millennia has come about as a result of the combined effects of both climatic deterioration and increased human impact.