Biostratigraphy and paleoecology of the Oligo-Miocene succession in Fars and Khuzestan areas (Zagros Basin,SW Iran)

Paleontological and biostratigraphical studies on carbonate platform succession from southwest Iran documented a great diversity of shallow-water benthic foraminifera during the Oligocene–Miocene. Larger foraminifera are the main means for the stratigraphic zonation of carbonate sediments. The distributions of larger benthic foraminifera in two outcrop sections (Abolhayat and Lali) in the Zagros Basin, Iran, are used to determine the age of the Asmari Formation. Four assemblage zones have been recognized by distribution of the larger benthic foraminifera in the study areas. Assemblage 3 (Aquitanian age) and 4 (Burdigalian age) have not been recognized in the Abolhayat section (Fars area), due to sea-level fall. The end Chattian sea-level fall restricted marine deposition in the Abolhayat section and Asmari Formation replaced laterally by the Gachsaran Formation. This suggests that the Miocene part of the formation as recognized in the Lali section (Khuzestan area) of the Zagros foreland basin is not present in the Abolhayat outcrop. The distribution of the Oligocene larger benthic foraminifera indicates that shallow marine carbonate sediments of the Asmari Formation at the study areas have been deposited in the photic zone of tropical to subtropical oceans. Based on analysis of larger benthic foraminiferal assemblages and microfacies features, three major depositional environments are identified. These include inner shelf, middle shelf and outer shelf. The inner shelf facies is characterized by wackestone–packstone, dominated by various taxa of imperforate foraminifera. The middle shelf is represented by packstone–grainstone to floatstone with a diverse assemblage of larger foraminifera with perforate wall. Basinwards is dominated by argillaceous wackestone characterized by planktonic foraminifera and large and flat nummulitidae and lepidocyclinidae. Planktonic foraminifera wackestone is the dominant facies in the outer shelf.     

Upper Maastrichtian depositional environments and sea-level history of the Kopet-Dagh Intracontinental Basin, Kalat Formation, NE Iran

The Kopet-Dagh Basin is located in northeastern Iran and southern Turkmenistan. The Late Maastrichtian Kalat Formation caps the Cretaceous interval in this Basin. Based on eight measured stratigraphic sections, the depositional environments and the sea-level history of the Kalat Formation have been interpreted. Petrographic and field observations led to identification of four major carbonates (A–D) and two siliciclastic lithofacies types. Carbonate rocks were deposited on a ramp setting within three zones including restricted and semi-restricted lagoons, bars, and open marine environments, while the siliciclastics were deposited at the shoreline. Sequence stratigraphic analysis identified two depositional sequences in the western and eastern parts and three depositional sequences in the central parts of the study area. Comparing the sea-level curve of Late Maastrichtian time in the Kopet-Dagh Basin with the global sea-level curve for the same time interval, there are some geometrical similarities and differences. The variations in the Basin are related to regional tectonic settings and sediment loading of the study area. Reconstructions of depositional environment during eight time slices of the Late Maastrichtian are presented. These results could be used for comparison with other localities worldwide and provide additional data for Late Cretaceous paleogeographic reconstructions.     

Relationships between foraminiferal assemblages and depositional sequences in Jahrum Formation,Ardal area (Zagros Basin,SW Iran)

The Jahrum Formation was deposited in the foreland basin in southwest Iran (Zagros Basin). The Zagros mountain belt of Iran, a part of the Alpine–Himalayan system, extends from the NW Iranian border through to SW Iran, up to the strait of Hormuz. The various facies of the Jahrum Formation were deposited in four main genetically related depositional environments, including: tidal flat, lagoon, shoal and open marine. These are represented by 14 microfacies. The Jahrum Formation represents sedimentation on a carbonate ramp. Tidal flat facies are represented by fenestral fabric, stromatolitic boundstone and thin-bedded planes. Carbonate deposition in a shallow marine lagoon was characterised by wacke–packstone, dominated by various taxa of imperforate foraminifer. The shoals are made up of medium- to coarse-grained skeletal and peloidal grainstone. This facies was deposited predominantly in an active high energy wave and current regime, and grades basinward into middle ramps facies are represented by wackestones–packstones with a diverse assemblage of echinoderm and large benthic foraminifers with perforate wall. Outer ramp facies consist of alternating marl and limestones rich in pelagic foraminifera. There is no evidence for resedimentation processes in this facies belt. The sequence stratigraphy study has led to recognition of three third-order depositional sequences.     

Facies,depositional sequences,and biostratigraphy of the Oligo-Miocene Asmari Formation in Marun oilfield,North Dezful Embayment,Zagros Basin,SW Iran

The Asmari Formation in Marun oilfield (south-west Iran), is about 440 m-thick marine carbonate succession with subordinate siliciclastic rocks, characterized by abundant benthic foraminifera (perforate and imperforate). Foraminiferal biostratigraphy indicates that this unit is Oligocene–Miocene in age. The distribution of benthic foraminifera and other components have led to the recognition of three siliciclastic and ten carbonate facies that were deposited in inner ramp (shoreline, tidal flat, restricted and open lagoon and shoal), middle and outer ramp sub-environments. Based on vertical facies trends, three third-order sequences in the Oligocene and three third-order sequences in the Miocene sediments have been identified. These depositional sequences are bounded by both type 1 and type 2 sequence boundaries. The transgressive systems tracts (TST) of sequences show deepening-upward facies trend with a gradual upward increase in perforate foraminifera, whereas the highstand systems tracts (HST) have a shallowing-upward facies trend and contain predominantly imperforate foraminifera. Deposition of these depositional sequences (DS) were controlled by both eustasy and tectonic subsidence.     

Applications of nummulitids and other larger benthic foraminifera in depositional environment and sequence stratigraphy: an example from the Eocene deposits in Zagros Basin,SW Iran

The Tale-Zang Formation in Zagros Mountains (south-west Iran) is a Lower to Middle Eocene carbonate sequence. Carbonate sequences of the Tale-Zang Formation consist mainly of large benthic foraminifera (e.g. Nummulites and Alveolina), along with other skeletal and non-skeletal components. Water depth during deposition of the formation was determined based on the variation and types of benthic foraminifera, and other components in different facies. Microfacies analysis led to the recognition of ten microfacies that are related to four facies belts such as tidal flat, lagoon, shoal and open marine. An absence of turbidite deposits, reefal facies, gradual facies changes and widespread tidal flat deposits indicate that the Tale-Zang Formation was deposited in a carbonate ramp environment. Due to the great diversity and abundance of larger benthic foraminifera, this carbonate ramp is referred to as a “foraminifera-dominated carbonate ramp system”. Based on the field observations, microfacies analysis and sequence stratigraphic studies, three third-order sequences in the Langar type section and one third-order sequence in the Kialo section were identified. These depositional sequences have been separated by both type-1 and type-2 sequence boundaries. The transgressive systems tracts of sequences show a gradual upward increase in perforate foraminifera, whereas the highstand systems tracts of sequences contain predominantly imperforate foraminifera.     

Size-related isotopic trends in some Maastrichtian planktic foraminifera: methodological comparisons, intraspecific variability, and evidence for photosymbiosis

The serial test dissection and sieve fraction methods for determining the pattern of size-related change in oxygen and carbon isotopic ratios are compared using four Late Cretaceous planktic foraminifer species (Racemiguembelina fructicosa, Planoglobulina acervulinoides, Planoglobulina multicamerata, and Pseudoguembelina palpebra) from a subtropical site in the North Atlantic (DSDP Hole 390A). Despite the extra labor required, we identify several clear advantages of the dissection method, including: (1) it provides a means of obtaining size-dependent changes in isotopic signatures that are unequivocally ontogenetic, whereas isotopic variation observed from sieve-separated size fractions could be ontogenetic or ecotypic; (2) the taxonomic identity of smaller sized specimens using the dissection method is unequivocal, whereas species identification is increasingly ambiguous in smaller size fractions using the sieve method; (3) it reveals a greater total range and a greater complexity in the pattern of ontogenetic change in stable isotopic values, whereas the sieve method averages the isotopic signal across the entire ontogenetic range preserved within the whole tests that are used. Our results from serial dissections demonstrate that among the species analyzed, R. fructicosa and P. acervulinoides yield relatively negative adult δ¹⁸O values, a large size-related change in δ¹³C values (1.32 and 2.05‰, respectively), and virtually no correlation between size-related δ¹³C and δ¹⁸O values. On this basis we suggest that these were photosymbiotic species that inhabited relatively shallow surface waters. Evidence for photosymbiosis is not as compelling for P. palpebra, as this species yields a 1.06‰ shift in δ¹³C and relatively negative δ¹⁸O values in adult chambers, but much stronger correlation between size-related δ¹³C and δ¹⁸O values (r²=0.40) than in R. fructicosa and P. acervulinoides. Planoglobulina multicamerata yields the most positive adult δ¹⁸O values of the species studied, a strong covariance between size-related δ¹³C and δ¹⁸O values (r²=0.77), and a 0.97‰ shift in δ¹³C composition during ontogeny. We conclude that this species lacked photosymbionts and migrated to a deeper surface water paleohabitat as it increased in size. Single specimen analyses of tightly constrained size fractions reveal a high degree of intraspecific variation. δ¹³C and δ¹⁸O values vary by up to 0.70 and 0.28‰ in R. fructicosa, 1.41 and 0.80‰ in P. acervulinoides, 0.66 and 0.82‰ in P. palpebra, and 0.18 and 0.33‰ in P. multicamerata, respectively. Such a range of isotopic variation has been observed in modern day planktic foraminifer assemblages, and likely results from growth of individuals during different phases of the seasonal cycle and/or the kinetic effect of intraspecific variation in shell calcification rates. As suggested by other investigators, large sample sizes should be analyzed to provide the most reliable correlation of stable isotopic stratigraphic records.     

A Foraminiferal Parasite on the Sea Urchin Echinocorys: Ichnological Evidence from the Late Cretaceous (Lower Maastrichtian,Northern Germany)

Archaeonassa is a poorly known ichnogenus, originally described from the Cambrian of North America, which is a member of the Scolicia ’group’, but can be regarded as distinct from Scolicia. Archaeonassa was originally based on modern material, and therefore falls into a taxonomic grey zone concerning its availability, but as A. fossulata was erected on fossil material the ichnogenus is here retained. Although exhibiting a degree of morphological plasticity in its style of ornament, Archaeonassa is considered to be monospecific, with only the ichnospecies A. fossulata recognised. The ichnogenus appears to be restricted to the Paleozoic, although similar material is well documented from modern environments. Thus the ichnogenus is expected to be geographically and stratigraphically more widespread than it otherwise appears. Archaeonassa is a valuable tool for environmental reconstruction within the Paleozoic as it occurs within intertidal deposits. The ichnogenus is interpreted chiefly as the work of gastropods, although in many cases may have been produced by trilobites or even echinoids, and represents the exogenic expression of either surface or shallow subsurface locomotion.     

Autochthonous facies and allochthonous debris flows compared: Early oligocene carbonate facies patterns of the Lower Inn Valley (Tyrol, Austria)

Summary This study presents a microfacies analysis and palaco-environmental interpretations of Early Oligocene carbon ates from the Lower Inn Valley Tertiary (“Unterinntal-Terti?r”) of Austria. The well preserved biogenic components allow detailed investigations of component relationships and controlling ecological parameters. The carbonates are dominated by coralline algae, corals, small and large benthic foraminifers, bryozoans and lithoclasts. Bivalves, gastropods, echinoderms, brachiopods and serpulids are subordinate. The limestones are present as A) autochthonous carbonates transgressing directly above the Triassic basement and B) allochthonous debris flows within deeper-water marls. These carbonates are found within the Paisslberg Formation. The Werlberg Member within this formation, pertains to the autochthonous carbonates and larger debris flows. Five facies types are separated following fabric analysis and statistical treatment (correlation, cluster analysis, principal components analysis) of semi-quantitative data consisting of component frequencies of thin sections. Facies distribution patterns are principally controlled by variations in substrate characteristics, turbulence and light along a depth gradient. Reconstruction of facies pattern distribution reveal both lateral and proximal-distal facies trends: coral-coralline algal facies, coralline algal facies as well as foraminiferal facies were situated in shallower environments, laterally adjacent to each other. These grade distally into coralline algal-bryozoan facies, bryozoan facies and finally into mollusc rich marls. Debris flows consisting of reworked material from all of the known facies (bioclastic packstone facies) is restricted to the debris flow and possible represents transport induced differentiation of components and grain size within distal debris flows.     

Benthic foraminiferal ecomarker species of the terminal Cretaceous (late Maastrichtian) deep-sea Tethys

Benthic foraminiferal distribution patterns throughout the late Maastrichtian Tethyan deep sea are analyzed. Many species are ubiquitously distributed throughout this region and therefore it is hard to assess their ecological preferences. However, five species show distribution patterns, which suggest that they may have distinctive paleoenvironmental preferences. These preferences are interpreted from hypothesized surface circulation and upwelling patterns. Additional information comes from Recent benthic foraminiferal ecology and from responses to the Cretaceous/Paleogene (K/Pg) boundary event. This enables us to assess the ecological preferences of these late Maastrichtian taxa, and establish them as ecological-marker (ecomarker) species for paleoenvironmental interpretation of the late Maastrichtian bathyal-abyssal Tethyan realm.(1) Eouvigerina subsculptura is suggested to be indicative of reasonably oxygenated upper-middle bathyal environments, though with high abundance of utilizable organic matter. (2) Sliteria varsoviensis is linked to areas of late Maastrichtian upwelling and seems to have been an epibenthic species with an opportunistic life mode. (3) Gavelinella beccariiformis and (4) Nuttallides truempyi are considered to be indicative of oligotrophic conditions unless they occur with a large proportion of endobenthic morphotypes. (5) Gavelinella pertusa is proposed to indicate neritic-middle bathyal environments of the ‘boreal’ realm, which might be influenced by more seasonal food-fluxes and by higher oxygen levels than similar settings in the (sub)tropics.Finally, the anomalous high abundances of the buliminid species Sitella cf. plana in deep open ocean environments is discussed in terms of possible mechanisms permitting such a (morphologically) opportunistic species to thrive in such an assumedly oligotrophic environment.     

Depositional environment and sequence stratigraphy of the Oligo-Miocene Asmari Formation in SW Iran

The Asmari Formation, a thick carbonate succession of the Oligo-Miocene in Zagros Mountains (southwest Iran), has been studied to determine its microfacies, paleoenvironments and sedimentary sequences. Detailed petrographic analysis of the deposits led to the recognition of 10 microfacies types. In addition, five major depositional environments were identified in the Asmari Formation. These include tidal flat, shelf lagoon, shoal, slope and basin environmental settings and are interpreted as a carbonate platform developed in an open shelf situation but without effective barriers separating the platform from the open ocean. The Asmari carbonate succession consists of four, thick shallowing-upward sequences (third-order cycles). No major hiatuses were recognized between these cycles. Therefore, the contacts are interpreted as SB2 sequence boundary types. The Pabdeh Formation, the deeper marine facies equivalent of the Asmari Limestone is interpreted to be deposited in an outer slope-basin environment. The microfacies of the Pabdeh Formation shows similarities to the Asmari Formation.     

Rediscovery of the Tagragra Tracksite (Maastrichtian,Agadir, Morocco): Agadirichnus elegans Ambroggi and Lapparent 1954 is Pterosaurian Ichnotaxon

ABSTRACT

Discovered in 1954 by Ambroggi and Lapparent, the Tagragra tracksite (Maastrichtian, Agadir, Morocco) yielded tracks of theropod dinosaurs, birds and enigmatic forms tentatively attributed to Lacertilia under the name Agadirichnus elegans. The original specimens are today considered to be lost, and the status of A. elegans has been questioned, even though these footprints can retrospectively be referred to pterodactyloid pterosaur tracks. In order to discuss the status of these foot prints, the historical site has been actively sought and recently rediscovered by one of us (MdD). New material confirms the presence of pterodactyloid footprints, with two morphotypes, in two different facies, respectively associated with bird and theropod tracks. Morphotype I is tentatively conferred to Pteraichnus Stokes 1957 Stokes, W. L. 1957. Pterodactyl tracks from the Morrison Formation. Journal of Palaeontology, 31:952–4.[Web of Science ®] , [Google Scholar], which extends the stratigraphical distribution of this ichnotaxon to the Maastrichtian. Morphotype II corresponds to the original ichnospecies A. elegans Ambroggi and Lapparent 1954 Ambroggi, R., and A. F. de Lapparent 1954. Les empreintes de pas fossiles du Maestrichtien d'Agadir. Notes du Service Géologique du Maroc, 10:43–6. [Google Scholar], which is here reassessed as a valid ichnotaxon, with the designation of a neotype. Thus, A. elegans Ambroggi and Lapparent 1954 Ambroggi, R., and A. F. de Lapparent 1954. Les empreintes de pas fossiles du Maestrichtien d'Agadir. Notes du Service Géologique du Maroc, 10:43–6. [Google Scholar] is historically the first pterosaurian ichnotaxon ever described.     

The Pleistocene lake deposits of the NE Baza Basin (Spain): salinity variations and ostracod succession

The Guadix-Baza Basin in Spain covers an area of approximately 3000 km² and yields a sedimentary sequence ranging from Lower Miocene to Pleistocene. Twenty five meters of Lower Pleistocene lacustrine sediments have been located in the NE part of the Basin at about 1000 meters in altitude. This sequence which overlies dolomitic mud flat deposits consists of limestones, calcareous and dolomitic mudstones, dolostones, silty clays, sands and gravels. Salinity fluctuations and short dry episodes, related to lake level oscillations, have been recorded by textural, mineralogical and faunal changes throughout the sequence. Ostracods, which are the most commonly encountered fossils, permit to detect recurrent changes in water salinity and regime, and solute composition. The faunal changes indicate an alternation of slightly saline and bicarbonate-rich water (when ostracods and gastropods occur) with a saline NaCl-dominant water (in which ostracods, Cerastoderma bivalves and non-marine foraminifers are found).The frequent and recurrent hydrochemical changes in the Baza Basin in the Early Pleistocene point to a climate of high contrast like in the Mediterranean region today but with a greater availability of water within the system compared to the present situation in the area.     

Population dynamics of Marginopora kudakajimensis Gudmundsson (Foraminifera: Soritidae) in the Ryukyu Islands, the subtropical northwest Pacific

The population dynamics of Marginopora kudakajimensis Gudmundsson, a dinoflagellate endosymbiont-bearing soritid foraminifer, was studied in the Ryukyu Islands, the subtropical northwest Pacific. Macroalgal samples were collected monthly between November 1995 and November 1996 at a 1-m-deep lagoonal site colonized by this species. Monthly variations in the size–frequency distributions and population density indicate that this foraminiferal population replaces itself in one year. Asexual reproduction occurs twice a year, in late spring and winter; in late spring, some of the adult individuals reproduce by multiple fission simultaneously. As a result, two cohorts (a late spring and a winter cohort) are found in the population during a year. Life span of the former cohort is up to one year, while that of the latter cohort is up to six months. Megalospheric specimens comprise up to 99% of the population with a few microspheric individuals throughout the year. The life tables and survivorship curves revealed that size-specific mortality rates were very low during the first several size classes and increased thereafter, indicating low juvenile mortality with high mortality later in life. The carbonate production rate by this Marginopora population is approximately 5 kg CaCO₃ m⁻² yr⁻¹, which is extremely higher than those reported for other larger foraminiferal species.     

Contributions to the Life Cycle of Peneroplis planatus (Fichtel and Moll) Montfort

Direct observations of schizonts and agamonts releasing megalospheres clarified the asexual phase of the life cycle of Peneroplis planatus and made it most probable that this species has a paratrimorphic life cycle. Specimens with maximum lengths between 837 and 3,503 μm released about 500 to 1,500 megalospheric juveniles, which possessed two chambers (proloculi and flexostyles) prior to emergence from the parental shell. The presence of gamonts was not shown and was only implied by the occurrence of the agamonts. Since agamonts and schizonts have been found from December to May and since asexual reproduction occurs in spring in Elat, sexual reproduction probably occurs at another time of year (June to December). More detailed studies of this species need to be conducted throughout the year to improve our knowledge of the life cycle of this species.     

Palaeoenvironments of the Buia Homo site: High-resolution facies analysis and non-marine sequence stratigraphy in the Alat formation (Pleistocene Dandiero Basin, Danakil depression, Eritrea)

The Early to Middle Pleistocene Dandiero rift basin is located near the Buia village, 110 km south of Massawa (Eritrea), and is filled by about 1000 m of continental deposits bearing 1 My erectus-like human remains. The basin fill consists of six formations (from bottom up): Bukra sand and gravels (fluvial), Alat formation (fluvio-lacustrine), Wara sand and gravel (fluvial), Goreya formation (lacustrine), Aro sand (fluvio-deltaic) and Addai fanglomerate (alluvial fan). This paper is focused on the Homo-bearing deposits of the Alat formation. The Alat formation records several episodes of changing accommodation space, with repeated shifts from fluvial to lacustrine conditions. The first episode points to rapid lake formation followed by a progressive shallowing. The second episode records fluvial aggradation above a sequence boundary, followed by the development and progressive infill of a second lake. The third episode points to a rapid lake formation followed by deposition of deltaic and fluvial sediments. The last episode is characterised by fluvial deposition under low-accommodation conditions. The development of the lacustrine environments was probably controlled by tectonics, in accordance with its formation during regionally dry climatic conditions, although minor shifts toward wetter conditions cannot be excluded. Human remains occur in the upper Alat formation, where the transition from a deltaic to alluvial setting is characterised by high frequency, potentially millennial-scale, lake-level oscillations. The palaeoanthropological record in the Buia area seems to be correlated with water availability (lacustrine coastal plains and floodplains) in a relatively open and grassy environment (grassland- and savannah-dominated). The increase in fluvial discharge that occurred at the top of the Alat formation led to erosion and winnowing of the underlying fossil-bearing, fluvio-deltaic deposits. A large number of bones and artefacts were consequently accumulated at the base of channel fills, which represent a preferential horizon for fossil searching.     

Dasycladales from the Upper Maastrichtian of Salento Peninsula (Puglia, southern Italy)

Summary A rich dasycladalean assemblage, mainly consisting of new taxa, has been discovered in upper Maastrichtian coarse bioclastic limestones of shelf margin facies cropping out along the southeastern coast of Salento peninsula (Puglia, Southern Italy). It consists of 8 species grouped into 6 genera:Cymopolia decastroi Parente 1994;C. barattoloi Parente, 1994;Zittelina fluegeli n.sp;Jodotella koradae (Dieni, Massari & Radoicic, 1983) nov. comb.;Barattoloporella salentina n. gen. n. sp.;Morelletpora dienii n. sp.;Neomeris spp. (two different species). Zittelina fluegeli n. sp. is characterized by an ovoid thallus with calcification made by a calcareous wall enveloping only the proximal part of the branches (except in basal whorls) and by closely packed, and partly coalescent, calcified ampullae arranged all around the median and distal portion of branches. The transferral ofNeomeris (Larvaria) koradae Dieni, Massari & Radoicic, 1983 into the genusJodotella, with the new combinationJodotella koradae, is proposed on the basis of new observations on the number and arrangement of fertile ampullae. The new genusBarattoloporella, type-speciesBarattoloporella salentina n. gen. n. sp., is erected for dasycladalean algae characterized by a segmented thallus. Each segment consists of: a) basal and apical sterile whorls made by primary branches only, b) central fertile whorls made by primary branches bearing in terminal position one fertile ampulla and one or two secondary branches. morelletpora dienii n. sp. is characterized by a segmented thallus with barrel shaped to pear shaped repetitive elements, consisting of simple whorls of first order branches only. Shape of the branches varies from regularly phloiophorous to more or less differentiated in a stalk and a swollen portion, with or without a subterminal constriction. This is by far the most diverse dasycladalean assemblage ever found in the Maastrichtian. Its diversity supports the conclusion that, within the Late Cretaceous, the Maastrichtian represents a maximum in dasycladalean diversity. Its taxonomic composition strenghtens the hypothesis that dasycladaleans were hardly affected by K/T mass extinction.     

Lithostratigraphy and facies development of upper cretaceous carbonates in east central Sinai,Egypt

Summary The Upper Cretaceous exposures in east central Sinai are represented by carbonate-dominated successions interbedding few sandstone, chert, shale and marl horizons. The recognised rock units are correlated with their counterparts commonly used in the Gulf of Suez region and central Sinai including from base to top: the Raha Formation, Abu Qada Formation, Wata Formation, Matulla Formation and the Sudr Chalk. Twelve limestone microfacies are encountered and are categorised as mudstones (pelmicrite and ostracod micrite), wackestones (pelagic biomicrite and foraminiferal biomicrite), grainstones (foraminiferal biopelsparite and oosparite), boundstones (bindstone and framestone), floatstones (coated-grained biomicrudite, rudist biomicrudite and shelly biomicrudite) and rudstones (shelly biosparudite). The dolostone microfacies include fine-medium crystalline ostracod dolostones and shelly dolostones. These microfacies have been compared with the Standard Microfacies Types and their depositional environments are discussed. The encountered litho- and biofacies suggest that the Cenomanian shallow transgressive sea had covered east central Sinai as far south as the Dahab region. By the advent of the Turonian, open marine subtidal conditions prevailed. This was followed by transitional conditions with shoals and tidal bars in the Late Turonian pointing to a regressive phase more pronounced at the southern localities. The rocks of the Matulla Formation were deposited in an oscillating environment of shallow subtidal to intertidal conditions during Coniacian-Santonian. In the Late Santonian and during most of the Campanian-Maastrichtian, sedimentation was influenced by open marine conditions with low sedimentation rates; local shallow subtidal regressive events occurred.     

Stratigraphic modelling of carbonate platform-to-basin sediments (Maastrichtian to Paleocene) in the Eastern Desert, Egypt

In the Galala Mountains of the Eastern Desert, Egypt, carbonate platform and basin deposits have excellent exposure. These exposures show a late Campanian–early Paleocene rimmed platform evolving into a late Paleocene distally steepened ramp. We modelled the evolution of the platform–basin transition from the Maastrichtian to Selandian (68.7–59 Ma) with the 2-D stratigraphic simulation program PHIL and compared the modelled results with outcrop sections. Stratigraphic, facies, and environmental data are summarized and operate as input and control parameters for the computer simulation. The most important parameters that control the depositional geometries of the late Cretaceous mixed carbonate siliciclastic platform and the Paleogene carbonate platform are changes in relative sea level, sediment flux and initial topography. The simulation provides an understanding of platform growth and slope to basin deposition, particularly in areas of the platform that are poorly exposed or have been eroded. Moreover, the simulated geologic parameters like lithology, overall thickness and palaeowater depth closely resemble field and laboratory measurements of the individual sections. In an earlier study, the Maastrichtian slope angle was calculated to be 5–8° and this was confirmed in this study. In this earlier study also the timing of the transition from a rimmed platform to a distally steepened ramp was established to be during latest Maastrichtian–early Paleocene. The present study shows that the rimmed platform persisted at least until the late Paleocene (59 Ma), as indicated by the relatively high slope angle of 6°.     

Oichnus Bromley Borings in the Irregular Echinoid Hemipneustes Agassiz from the Type Maastrichtian (Upper Cretaceous,The Netherlands and Belgium)

Three ichnospecies of Oichnus Bromley occur in tests of the large holasteroid echinoid Hemipneustes striatoradiatus (Leske) in the type area of the Maastrichtian (Upper Cretaceous) in The Netherlands and Belgium; Oichnus simplex Bromley (penetrative), Oichnus paraboloides Bromley (nonpenetrative and showing two distinct morphologies), and Oichnus excavatus isp. nov. (nonpenetrative). The two distinct morphologies of O. paraboloides (both shallow, one with a central boss) are gregarious, but do not occur together on the same specimens, suggesting they were generated by different taxa. Oichnus paraboloides with a central boss occurs on H. striatoradiatus from the upper Nekum Member, Maastricht Formation (Maastrichtian, Upper Cretaceous). Tests of the host echinoid are smaller in the overlying Meerssen Member, Maastricht Formation, where they are infested by O. excavatus, the largest borings considered herein, which have concave walls and a large central boss. Blisters inside tests from the Meerssen Member show that this infestation occurred when the echinoid was alive. It is postulated that producers of these borings in H. striatoradiatus may have been genetically related and increased in size during the Maastrichtian even as the host echinoids showed a size decrease. This size increase in H. striatoradiatus was genetic and cannot be related to increase in size of borings.