陕西子洲中侏罗世早期恐龙足迹初步研究

陕西子洲县中侏罗统延安组砂岩中,同一地点共厚约1.7 m的5个层面发现4种食肉类恐龙足迹,自上而下分别为:1)第5层面大型三趾型足迹,实雷龙足迹科王氏子洲足迹(新遗迹属、新遗迹种)Zizhoupus wangi ichnogen.et ichnosp,nov.;2)第4和第3层面中型三趾型足迹,实雷龙足迹科龙尾峁张北足迹(新遗迹种)Changpeipus longweimaoensis inchnosp.nov.;3)第2层面小型三趾或四趾型足迹,虚骨龙类小理河陕西足迹(新遗迹种)Shensipus xiaoliheensis inchnosp.nov.;4)第1层面小型三趾型足迹,虚骨龙类的铜川陕西足迹Shensipus tungchuanensis.     

云南早侏罗世一新具嵴冠兽脚类恐龙

报道了具嵴冠的兽脚类恐龙一新属种,安龙堡双柏龙(Shuangbaisaurus anlongbaoensis gen.et sp.nov.).双柏龙发现于云南省楚雄彝族自治州双柏县下侏罗统冯家河组中,保存了部分头骨带下颌.双柏龙沿两侧眼眶背缘向上有嵴冠发育,这在其他兽脚类中未曾报道过.与其他早侏罗世体型较大且具一对矢状嵴冠的兽脚类(双嵴龙属和中国龙属)相比,双柏龙还独具一些特征组合,如相对较高的前颌骨体、抬高的前颌骨腹缘、后腹向延伸的轭骨后突及较小的上颞孔.比较研究表明,尽管中国“双嵴龙”(“Dilophosaurus”sinensis)可能应归入中国龙属(Sinosaurus),但未必属于模式种.双柏龙的发现将有助于研究基干兽脚类的演化,尤其是各类头骨骨饰在其中的作用.     

New dinosaur tracksites from the Middle Jurassic of Madagascar: ichnotaxonomical,behavioural and palaeoenvironmental implications

Abstract: New dinosaur tracksites are described from the Bajocian–Bathonian Bemaraha Formation of western Madagascar. Two track‐bearing surfaces can be followed over a distance of at least 4 km, suggesting the existence of a hitherto unrecognized megatracksite. The track assemblage is theropod dominated, but sauropod tracks also occur at one site. Qualitative and quantitative analysis of the abundant theropod track material suggests that most, if not all, theropod footprints are attributable to a single trackmaker and are referred to Kayentapus isp. Although this ichnogenus, originally described from the Lower Jurassic of North America, has never been recorded from Gondwana nor from the Middle Jurassic, track morphology strongly suggests this attribution. Palaeogeographical, sedimentological and ichnological data suggest that the dinosaur tracks formed in an intertidal to supratidal setting where the coastline influenced the preferred walking direction of the animals.     

四川威远恐龙足迹一新属

1985年,自贡恐龙博物馆的朱时达、高人彦、彭光照根据四川威远县荣胜乡沙坝村村民李元吉、李鸣章的报告和地矿部第二地质大队提供的资料,在威远县荣胜乡老鸦坡下侏罗统珍珠冲组下部黄色砂岩上发掘出6个连续恐龙足迹化石,并把这批标本运回自贡恐龙博物馆保存。     

The dinosaur tracksite of Loulle (early Kimmeridgian; Jura,France)

More than 1500 sauropod and theropod tracks have been discovered and excavated on the floor of a disused 3800 m² quarry, near the village of Loulle (French Jura). These levels correspond to a tidal-flat environment, from intertidal to supratidal zone, dated from the earliest Kimmeridgian (∼157 M.a.). Eighteen sauropod trackways have been recognized, ranging from 3.8 m to 51.5 m in length. These trackways do not correspond to a unique herd passage, because trackways are diversely oriented, most of them are crossing others and prints are more or less deeply marked, depending of different stages of substratum competence. According to the trackway gauges, the three types (narrow-, medium and wide-gauge) are represented. Some trackways could be referred to the ichnogenus Brontopodus Farlow et al., 1989, and others to Parabrontopodus Lockley et al., 1994a. Based on footprint size, the body-sizes of these 18 sauropod trackways range from tiny (Pes Length < 25 cm) to large individuals (PL > 75 cm), showing a regular continuum of sizes. Several biometrical ratios classically used in sauropod ichnology appear to be closely related to size (e.g., the smallest individuals apparently moved faster than the largest; large individuals are all narrow-gauge while small individuals are all wide-gauge; large specimens are characterized by a high heteropody ratio when the smallest show a low heteropody ratio). Associated with these sauropod trackways, four theropod trackways have also been recorded. Three small to medium-sized theropod are referred to the ichnogenus Carmelopodus Lockley et al., 1998a, while a large individual (PL = 77 cm) is tentatively referred to Megalosauripus Lessertisseur, 1955.     

A sauropod dinosaur megatracksite from the Late Jurassic of northern Switzerland

The largest known sauropod trackway site from the Upper Jurassic in Europe has been found in the northern Jura Mountains of Switzerland. Since the initial discovery of the site in 1988, detailed ichno‐facies mapping of the sites has been undertaken and completed. Six separate sites (from 5 to 45km apart) have been located to date; the largest one, displaying 345 single imprints, extends over a surface of 7000 m². As all of the reported sites occur within the same bed, these vertebrate prints form a megatracksite covering an area of more than 360 km². All localities are stratigraphically within the Reuchenette Formation. Two biostratigraphically diagnostic ammonites have been found within the sequence (Aulacostephanus; Gravesia), indicating an Upper Kimmeridgian age (sensu gallico). In the easternmost sites, imprints occur on mud‐cracked tidal pond deposits, whereas the western tracksites are found in supratidal algal marsh deposits. The track‐bearing horizon is immediately below a trans‐gressive event at all the sites, suggesting that the ichnofaunas represent dinosaur activity during a relative sea‐level low‐stand. Footprint size and morphology, as well as trackway dimensions, suggest that the track producers belong to the largest sauropods yet recorded in the European Jurassic, closely resembling the Breviparopus tracks from the Middle Jurassic of Morocco.     

内蒙古鄂托克早白垩世恐龙足迹化石8号点风化机理研究

内蒙古鄂托克旗查布地区是罕见的恐龙足迹等遗迹化石分布区,分布着大量白垩纪恐龙足迹化石。这些足迹化石在野外遭受着不同程度的风化破坏,为此在足迹密集的核心区8号点建立了野外地质遗迹博物馆保护原址。跟踪观察后发现室内足迹化石的风化程度相较于露天保存更加严重,尤其在馆内靠近四周围墙的区域,足痕表面酥粉破碎,有的足迹甚至完全消失。为研究8号点足迹化石的风化原因,本文对8号点馆内外赋存于同一层位的岩石进行取样,使用偏光显微镜(PM)、X射线衍射(XRD)、压汞(MIP)、离子色谱(IC)、拉曼光谱(Raman)、傅里叶变换红外光谱(FTIR)以及扫描电子显微镜能谱(SEM-EDS)等方法进行分析,结果发现室内岩石的孔隙率更高,孔径更大,可溶盐含量是室外上层的2倍,室外下层的约11倍,种类以NaCl和Na₂SO₄为主。模拟实验显示,相比冻融作用,Na₂SO₄更具有破坏性。由此认为导致鄂托克遗迹博物馆内恐龙足迹化石风化的机理是由地下水和降雨形成的地表水共同作用的结果,尤以Na₂SO_4...     

First Early Jurassic Ornithischian and theropod footprint assemblage and a new ichnotaxon Shenmuichnus wangi ichnosp. nov. from Yunnan Province,southwestern China

The new ichnospecies, Shenmuichnus wangi ichnosp. nov., is the first evidence for the presence of large ornithischians in the Early Jurassic of Yunnan Province, whereas the known skeletal record documents small species only. Until now Shenmuichnus was known from a single locality in Shaanxi Province by the ichnospecies Shenmuichnus youngteilhardorum. Compared with the latter, Shenmuichnus wangi is larger and shows a different trackway configuration, particularly in the relative position of manus and pes imprints. Palecologically, the occurrence of Shenmuichnus wangi in a red bed facies indicates the preference of distinctive environments of trackmakers of both ichnospecies, questioning former hypotheses of exclusivity of ornithischians in more humid climates. By abundance both skeletons and footprints of ornithischians suggest their role as a minor component in Early Jurassic saurischian dominated dinosaur faunas in this region.     

山西天镇晚白垩世一新鸭嘴龙超科恐龙（英文）

记述了山西天镇晚白垩世灰泉堡组发现的一鸭嘴龙超科恐龙新属种:天镇大同龙Datonglong tianzhenensis gen.et sp.nov.。标本为一保存较完好的右侧齿骨并带牙齿。与其他鸭嘴龙超科恐龙不同,大同龙每个齿槽有两个功能齿,而且舌面齿冠上嵴的发育也具有独特的特征组合(主嵴向远中方向偏移;次嵴发育;无其他附嵴;主嵴上半段略后弯)。对比研究发现,在较进步的非鸭嘴龙科鸭嘴龙超科类群中齿骨和牙齿的演化过程复杂,许多鸭嘴龙科的特征在这些类群中时而出现,如:巴克龙Bactrosaurus齿冠具有较大的高/宽比;吉尔摩龙Gilmoreosaurus齿冠上只有一个主嵴和纤弱的附嵴;张衡龙Zhanghenglong齿冠上主嵴中置;南宁龙Nanningosaurus下颌冠状突前倾;大同龙Datonglong每个齿槽有两个功能齿。这表明为了获得更有效的牙齿咀嚼方式,在鸭嘴龙科起源前它的姐妹群们曾进行过多种尝试;这也告诫我们,受大量趋同演化的影响,要厘清这些类群间的关系并非易事。     

甘肃早白垩世恐龙蛋化石新类型（英文）

甘肃省早白垩世地层中,出土了大量恐龙骨骼以及恐龙足迹化石,但是至今未有蛋化石的报道。根据发现于兰州-民和盆地下白垩统河口组的蛋壳化石,建立一恐龙蛋新蛋属、蛋种,并将其归于一新蛋科:Polyclonoolithidae(多小枝蛋科)。新发现的蛋化石标本不同于所有已知的恐龙蛋类型,具有独特的显微特征组合:分叉的蛋壳单元向外延伸至蛋壳外表面,并未在靠近蛋壳外表面处融合成层;弦切面上具相互链接或独立的多角形的蛋壳单元;以及不规则的气孔道。中国的恐龙蛋化石大多出自晚白垩世地层,仅在辽宁有早白垩世恐龙蛋的报道。新发现扩展了中国恐龙蛋化石的地质和地理分布,也有可能为圆形蛋科蛋壳结构的起源提供新的认识。     

Dinoturbation in Upper Jurassic siliciclastic levels at Cabo Mondego (Lusitanian Basin,Portugal): evidences in a fluvial-dominated deltaic succession

At Cabo Mondego (western central Portugal), the Upper Jurassic marine to coastal succession contains several stratigraphic levels preserving dinosaur footprints on the surface bedding plane, as well as convolute bedding and soft sediment injection structures interpreted as dinoturbation structures. At least nineteen new three-dimensional structures observed in cross-sections are interpreted as produced by dinosaur trampling. The identification of three-dimensional structures of dinosaur footprints provides an important complement to the information obtained from footprints preserved on single bedding surfaces, such as the substrate consistency, potential trackmaker identification, and the possibility to enhance the distinction of sauropods and tridactyl dinosaurs, and paleoenvironmental interpretations. In the lower part of the Arenitos da Boa Viagem Formation, eight levels of probable lowermost Kimmeridgian age (ca. 157–156 Ma), displaying the above-mentioned deformational structures, were analyzed in detail. They support interpretations concerning the relationship between the footprints and the substrate consistency at the time of their formation. Three distinct cohesiveness patterns, defined by the penetration of the feet from the paleosurface, are the result of different degrees of substrate cohesiveness. Identifying the trackmakers of levels belonging to the middle Oxfordian–lower Kimmeridgian has important implications for Late Jurassic ecosystem reconstructions, as the footprints observed in Cabo Mondego indicate a change in the morphotypes throughout the Upper Jurassic succession.     

Two new dinosaur tracksites from the Lower Cretaceous Jiaguan Formation of Sichuan Basin,China: specific preservation and ichnotaxonomy

Two new dinosaur tracksites are reported from the Lower Cretaceous Jiaguan Formation in the Sichuan Basin, Qijiang District of Chongqing. These are the Gaoqing-Yongsheng and the Huibu tracksites, which represent the 13th and 14th reports from this formation. The Gaoqing-Yongsheng tracksite reveals the trackway of a large biped (ornithopod) in association with isolated sauropod tracks and large indeterminate undertracks with radial cracks. These features are preserved as natural casts with pebble infillings in a coarse, cross bedded and very thick bedded sandstone sequence. The Huibu tracksite reveals isolated theropod tracks and ornithopod tracks, the latter having a quadripartite, Caririchnium-like morphology, preserved in a thin bedded sandstone sequence with intercalated mudstone.     

First Thyreophoran Type Tracks from the Middle Jurassic Chuanjie Formation of Yunnan Province,China

Two tridactyl footprints from the Chuanjie Formation (Middle Jurassic) of Yunnan Province, China are morphological characteristics of thyreophoran tracks. They show some similarities to Shenmuichnus, known from the Early Jurassic strata of both Shaanxi and Yunnan provinces, but are somewhat larger, thereby resembling the ichnogenus Stegopodus. Based on their general morphology and size being congruent with this ichnogenus, they are tentatively assigned here to cf. Stegopodus. This is the fourth report of large ornithischian (probably thyreophoran) tracks from the Lower-Middle Jurassic of China that indicates relatively large trackmakers that were likely to be taxonomically distinct from much smaller and gracile Anomoepus trackmakers, also of ornithischian affinity. The larger tracks indicate a hitherto unreported abundance, size range and diversity of track types attributed to this group. The parallel orientation of the two best preserved trackways may indicate gregariousness.     

Chemical preservation of tail feathers from Anchiornis huxleyi,a theropod dinosaur from the Tiaojishan Formation (Upper Jurassic,China)

A panel of geochemical techniques is used here to investigate the taphonomy of fossil feathers preserved in association with the skeleton of the Jurassic theropod Anchiornis huxleyi. Extant feathers were analysed in parallel to test whether the soft tissues morphologically preserved in the fossil also exhibit a high degree of chemical preservation. Scanning electron microscopy (SEM) and energy dispersive spectroscopy (EDS) indicate that clays and iron oxide pseudomorphs occur in the surrounding sediment and also reveal the preservation of melanosome-like microbodies in the fossil. Carbon gradient along a depth profile and co-occurrence of carbon and sulphur are shown in the fossil by elastic backscattering (EBS) and particle-induced x-ray emission (PIXE), which are promising techniques for the elemental analysis of fossil soft tissues. The molecular composition of modern and fossil soft tissues was assessed from micro-attenuated total reflectance fourier transform infrared spectroscopy (micro-ATR FTIR), solid-state ¹³C nuclear magnetic resonance (CP-MAS ¹³C NMR) and pyrolysis gas chromatography mass spectrometry in the presence of TMAH (TMAH-Py-GC-MS). Results indicate that the proteinaceous material that comprises the modern feathers is not present in the fossil feathers. The fossil feathers and the embedding sediment exhibit a highly aliphatic character. However, substantial differences exist between these samples, revealing that the organic matter of the fossil feathers is, at least partially, derived from original constituents of the feathers. Our results suggest that, despite the morphological preservation of Anchiornis feathers, original proteins, that is keratin, were probably not preserved in the 160-myr-old feathers.     

Review and Detailed Description of Sauropod-Dominated Trackways from the Upper Cretaceous Jiangdihe Formation of Yunnan,China

Type specimens of the sauropod ichnotaxon Chuxiongpus changlingensis, which was later reassigned to Brontopodus changlingensis, as well as the theropod ichnotaxon Yunnanpus huangcaoensis, both from the Cretaceous Jiangdihe Formation of Yunnan Province, are redescribed in order to document their morphological features. Both, but particularly Y. huangcaoensis, which is considered now a nomen dubium, were originally based on poorly preserved material. Nevertheless, the specimens document a saurischian dominated biota that existed during the deposition of the Jiangdihe Formation from which no skeletal remains are known. B. changlingensis trackways were left by small sauropods that show consistent partial or complete overprint of the manus by the pes. This pattern makes it difficult to calculate manus length, and heteropody can be only estimated.     

Dinosaur tracks in Lower Jurassic coastal plain sediments (Sose Bugt Member,Rønne Formation) on Bornholm,Denmark

Fluvial palaeochannels of coastal plain sediments of the Lower Jurassic Sose Bugt Member of the Rønne Formation exposed in the coastal cliffs at Sose Bugt, Bornholm, contain abundant dinosaur or other large vertebrate tracks in the form of deformation structures exposed in vertical section. The tracks are represented by steep‐walled, flat‐to‐concave‐bottomed depressions, with a raised ridge at each side. The tracks are filled with laminated sediments, draping the contours of the bottom of the depression. Underprints, stacked concave deformations beneath the prints, are present beneath each track. Contemporary Upper Triassic – Lower Jurassic strata from southern Sweden and Poland contain a diverse track fauna, supporting our interpretation. This is the earliest evidence of dinosaur activity in Denmark.     

贵州蕨类植物新记录

报道了贵州省新记录的4种蕨类植物,分属3科,4属,它们是上毛凤丫蕨（Coniogramme suprapilosa）、具腺凸轴蕨（Metathelypteris glangdulosa）、光滑方秆蕨（Glaphyropteridopsis glabrata）和混淆鳞毛蕨（Dryopteris commixta）。     

Hoof-Like Unguals,Skin, and Foot Movements Deduced from Deltapodus Casts of the Galve Basin (Upper Jurassic-Lower Cretaceous,Teruel, Spain)

Galve (Teruel, Spain) is a town in the interior of a synclinal fold with Upper Jurassic marine limestones along its flanks, and, in its core, Upper Jurassic–Lower Cretaceous continental and shoreline sediments crop out. The core sediments cover an area about 8 km², and contain a concentration of sites with footprints, bones, and eggshells of dinosaurs. The footprints are both shafts and natural casts. Some casts are attributed to stegosaurs (Deltapodus). The Deltapodus casts are characterized by features that allow us to make direct observations on the skin formed by polygonal scales, and ellipsoidal “hooves,” as well as deductions on the movement of the limbs during walking. According to the opinion of some authors, dinosaur footprints are indicators of the motion of their limbs and sometimes of the whole body. So far, results have been deduced from theropod, ornithopod, and sauropod footprints. This article shows the results obtained from analis of the aforementioned Deltapodus casts, i.e., forelimb movement similar to that of the forelimbs of sauropods, and the rigid structure of the autopodial part of the hind limb.     

Lavinipes Cheminii Ichnogen., Ichnosp. nov., A Possible Sauropodomorph Track from the Lower Jurassic of the Italian Alps

A new ichnotaxon is described from the Lower Jurassic (Upper Hettangian-Lower Sinemurian) carbonate tidal flats on the central-eastern Italian Alps. The narrow-gauge trackway is that of a large quadrupedal dinosaur. The pes is functionally tetradactyl with three rounded antero-medially directed digits, and the manus is pentadactyl. This quadrupedal form is close to Otozoum and Pseudotetrasauropus jaquesi both traditionally related to sauropodomorph trackmakers. The similarity with Otozoum is so marked that Lavinipes and Otozoum could be cogeneric. But the overall evidence today is that the Otozoum trackmaker was generally bipedal, whereas the trackmaker of L. cheminii is fully quadrupedal. The manual prints of L. cheminii show five short clawless digits and are different from the tetradactyl slender toed manual prints of Otozoum. The possible sauropodomorph affinity of the L. cheminii trackmaker is here discussed with an attempt to a revision of the Late Triassic-Jurassic tracks which have been traditionally related to sauropod and prosauropod.     

Dinosaur Footprints from Northeastern Brazil: Taphonomy and Environmental Setting

The Brazilian Cretaceous basins located at the northeastern region of Brazil present a wide distribution of dinosaur tracks and isolated footprints in many environmental settings: alluvial fans, braided and meandering fluvial floodplains, marginal lake borders and tidal flats. They are mainly theropod and sauropod footprints of Neocomian and Cenomanian ages that are found in Sousa, Uiraúna-Brejo das Freiras, Araripe, Cedro, Malhada Vermelha, Lima Campos and São Luís Basins. The vertebrate ichnofossils are important biogenic sedimentary structures, a picture of the interaction between dinosaur's behavior and the substrate nature, allowing the paleobiological analysis and inferences about the palaeoenvironments.