Platform-basin transect of a middle to late Jurassic large-scale carbonate platform system (Shotori mountains,Tabas area,east-central Iran)

Summary Following a phase of predominantly siliciclastic sedimentation in the Early and Middle Jurassic, a large-scale, low-latitude carbonate depositional system was established in the northern part of the Tabas Block, part of the central-east Iranian microplate, during the Callovian and persisted until the latest Oxfordian/Early Kimmeridgian. Running parallel to the present eastern block margin, a NNW/SSE-trending carbonate platform developed in an area characterized by reduced subsidence rates (Shotori Swell). The growth of this rimmed, flat-topped barrier platform strongly influenced the Upper Jurassic facies pattern and sedimentary history of the Tabas Block. The platform sediments, represented by the predominantly fine-grained carbonates of the Esfandiar Limestone Formation, pass eastward into slope to basin sediments of the Qal'eh Dokhtar Limestone Formation (platform-derived allochthonites, microbialites, and peri-platform muds). Towards the west, they interfinger with bedded limestones and marlstones (Kamar-e-Mehdi Formation), which were deposited in an extensive shelf lagoon. In a N−S direction, the Esfandiar Platform can be traced for about 170 km, in an E-W direction, the platform extended for at least 35–40 km. The width of the eastern slope of the platform is estimated at 10–15 km, the width of the western shelf lagoon varied considerably (>20–80 km). During the Late Callovian to Middle Oxfordian, the Esfandiar Platform flourished under arid climatic conditions and supplied the slope and basinal areas with large amounts of carbonates (suspended peri-platform muds and gravitational sediments). Export pulses of platform material, e.g. ooids and aggregate grains, into the slope and basinal system are interpreted as highstand shedding related to relative sealevel variations. The high-productivity phase was terminated in the Late Oxfordian when the eastern platform areas drowned and homogeneous deep water marls of the Upper Oxfordian to Kimmeridgian Korond Formation onlapped both the Qal'eh Dokhtar Limestone Formation and the drowned Esfandiar Limestone Formation. Tectonic instability, probably caused by faulting at the margins of the Tabas Block in connection with rotational movements of the east-central Iranian block assemblage, was responsible for the partial drowning of the eastern platform areas. In some areas, relicts of the platform persisted to produce shallow-water sediments into the Kimmeridgian.     

Evidence of synsedimentary tectonics in the Northern Tabas Block,East-Central Iran: The Callovian (Middle Jurassic) Sikhor Formation

Summary The Sikhor Formation (new) is a predominantly siliciclastic sediment package intercalated between the marly-silty Baghamshah Formation (below) and the calcareous Esfandiar Limestone and Qal'eh Dokhtar Limestone formations (above). All stratigraphic evidence points to an Early Callovian age of the formation. The Sikhor Formation is restricted to the southern and central Shotori Mountains and consists of two members: The Kuh-e-Neygu Member (new) is composed of fluvialdeltaic conglomerates, sandstones, and siltstones grading into marly silt of the Baghamshah Formation. The overlying Majd Member (new) is characterised by mixed siliciclastic-carbonate sediments that record the interfingering of carbonate ramp sediments with fluvialdeltaic sands and silts. Evidence of erosional truncation of the underlying Baghamshah Formation and confinement of the siliciclastic sediment to a comparatively narrow, NNW-SSE elongated strip suggest that the formation had its origin in the asymmetric uplift of a westdipping tilted fault block in the southern Shotori Mountains that shed its sediment predominantly in a northern and eastern direction. After erosional levelling, the former uplifted areas were overgrown by the highly productive Esfandiar Carbonate Platform. The Sikhor Formation thus is evidence of an extensional tectonic pulse in the early Callovian and underlines that this area of the Tabas Block was a tectonically highly unstable area during most of the Jurassic.     

Toarcian and Aalenian (Jurassic) ammonites from the Shemshak Formation of the Jajarm area (eastern Alborz, Iran)

In the Jajarm area (eastern Alborz Mountains, NE Iran) the ?Upper Triassic — Lower Bajocian siliciclastic Shemshak Formation is up to 2000 m thick. Whereas the lower third of the formation is nearly exclusively non-marine, the upper two-thirds are fully marine. The middle part is characterized by several levels containing a relatively diverse and well preserved assemblage of ammonites of the Toarcian to Aalenian stages. Two sections of the ammonite-bearing strata, spaced 20 km, are presented and correlated by means of ammonite biostratigraphy. The ammonite fauna consists of 27 taxa, some of which are recorded for the first time from the Alborz Mountains. The ammonites are briefly described and their palaeobiogeographic context is reviewed. The ammonite fauna of the Shemshak Formation at Jajarm, as elsewhere in North and Central Iran, is exclusively Northwestern Tethyan in character and is closely related to the faunas of Northwestern and Central Europe.     

Early Jurassic (latest Toarcian) brachiopods from the northeastern margin of the Western Tethys (Central Iran) and their paleobiogeographical significance

Brachiopod fauna from central Iran, recorded in the upper part of the Shemshak Group and attributed to the upper Toarcian (Pseudoradiosa-Aalensis zones), are reported for the first time in Iran. The assemblage recognized includes six different taxa: Homoeorhynchia sepahanensis nov. sp., formally described in this paper, Globirhynchia subobsoleta, Pseudogibbirhynchia sp., Tetrarhynchiidae sp. indet., Monsardithyris? aff. haresfieldensis, and Zeilleria cf. leckenbyi. Analysis of faunal affinities with other paleobiogeographical regions shows a free connection of the central Iranian brachiopod fauna and wide areas of the northern shelf margin of the Tethys Ocean. This is due to an apparent disruption of bioprovinciality inferred for the late Toarcian-earliest Aalenian, congruent with a connection through the northern seaway across the peri-Laurasian epicontinental platforms.     

Taxonomy and palaeobiology of the Bathonian (Middle Jurassic) tulitid ammonite Morrisiceras

Taxonomic revision of a number of species belonging to the Middle Bathonian tulitid genus Morrisiceras (Morrisiceras Buckman - macroconchs and Holzbergia Torrens - microconchs) from the Polish Jura (south-central Poland) and England, shows that the genus is in fact represented by one, albeit morphologically very plastic (especially macroconchs), species Morrisiceras morrisi (Oppel). In the Polish Jura area, all of the previously described morphotypes of the macroconch, along with its microconch Holzbergia, are characterized by their identical stratigraphic ranges, both being confined to the Middle Bathonian Morrisi Zone. Much smaller microconchs (males) may have attained maturity earlier than their macroconchs (females). Morphologically very plastic macroconchs seem to have attained full maturity at various shell diameters. Such phenomenon is known in recent coleoids and similarly may be the result of changing environmental conditions. Palaeobiogeographically, the main area occupied by the species was confined to the epicratonic seas, which spread across what is now present-day central and North-West Europe, with some occurrences in western Asia (Turkmenistan/Uzbekistan) and sporadic occurrence in a deep-water facies of Apulia (Italy). Taphonomical observations suggest that Morrisiceras shells have not undergone significant post-mortem drifting or transport after deposition. The preservation of some of the Morrisiceras morphotypes, occurring as dorso-ventrally flattened specimens in the host clays, may indicate that they were quickly buried in life position after they have landed on the sea-floor.     

Probable crocodilian tracks and traces from the Morrison Formation (Upper Jurassic) of eastern Utah

A tetradactyl pes impression and tridactyl manus impression are described as the type specimen of Hatcherichnus sanjuanensis ichnogen. et ichnosp. nov., a probable large crocodilian ichnite from the Salt Wash Member of the Upper Jurassic Morrison Formation in eastern Utah. A similar pes track from the Morrison Formation at Garden Park, Colorado, may also belong to this ichnogenus. The type specimen from Utah consists of plaster replicas of natural casts of a left pes impression and a left manus impression. Associated with the type specimen were possible tail and body drag impressions. The tracks do not appear to be part of a walking trackway and may be swim tracks associated with an animal in shallow water. The tracks occur at a visible contact between slightly fining‐upward channel sandstone units.     

Benthic foraminiferal assemblages of the Middle and Upper Jurassic sediments from the northeastern Alborz and western Koppeh Dagh,Iran: Systematic palaeontology and palaeoecology

This research is focused on a poorly studied Jurassic sequence in the eastern part of Alborz and Western Koppeh Dagh basins. The foraminifera are reported from Callovian (Middle Jurassic) to Tithonian (Upper Jurassic) sediments of the Farsian and Chaman Bid formations. The benthic foraminiferal associations belong to the families Ammodiscidae, Epistominidae, Hauerinidae, Hormosinidae, Ichthyolariidae, Lagenidae, Nodosariidae, Nubeculariidae, Ophthalmidiidae, Polymorphinidae, Rhabdamminidae, Saccamminidae, Spirillinidae, Spirocyclinidae, Textulariidae, Trocholinidae, and Vaginulinidae. In the studied area, the foraminifera are dominated by the families Nodosariidae, Vaginulinidae and Epistominidae. Palaeoecological analysis of the foraminiferal assemblages from the Farsian Formation indicates that these sediments were deposited on the inner to mid shelf environment, with normal salinity (euhaline), and relatively well oxygenated waters, with eutrophic conditions in the Goznawwi area and oligotrophic conditions in the Chaman Bid area. The foraminiferal assemblages in this study are the first record from Jurassic sediments of the eastern part of the Alborz and Koppeh Dagh basins in northern Iran.     

New data on the Rectogordius (foraminifera) abundance zone (Latest Carboniferous: Gzhelian) of the Zaladou Formation (east-central Iran,Tabas block,Shishtu section)

The Rectogordius (Foraminifera) abundance zone is described in the east of the Shishtu village in the Ozbak Kuh Mountain. The samples were collected in the Zaladou Formation., which is 60 m thick and composed of shales, sandstones, sandy limestones, microconglomerate, bioclastic limestones, coral limestone and fusulinid limestones. The Rectogordius abundance zone was found in sandy bioclastic limestone. It displays two species and three subspecies of this foraminifer, including Rectogordius iranicus, R. iranicus gadukensis, R. minimus and R. minimus shishtuensis n. subsp., R. iranicus ozbakensis n. subsp. The age of this abundance zone is considered to be Gzhelian, due to the distribution of Rectogordius in Central Iran (Ozbak Kuh; Zaladou Formation), central and eastern Alborz (Emarat Fm.), Sanandaj-Sirjan zone (Vazhnan Formation), as well as in the Donets, Arctic Canada, Afghanistan, and the Carnic Alps. The genus Rectogordius is possibly restricted to the northern Paleotethys margin, northern Cimmerian margin, shelf of the Uralian Ocean as far as the northernmost part of North America. Two new subspecies Rectogordius minimus shishtuensis n. subsp. and Rectogordius iranicus ozbakensis n. subsp. are described.     

Microtaphonomy of bioclasts and paleoecology of microencrusters from Upper Jurassic spongiolithic limestones (External Prebetic,Southern Spain)

Microencrusters and microtaphonomic features of the Oxfordian spongiolithic limestones of the External Prebetic were studied using thin-section analysis. The spongiolithic limestone is a bioclastic-rich packstone with common echinoderm, mollusc and brachiopod remains. The bioclasts show a high fragmentation index and frequent microborings. The encrustation index (E _i) is higher for fragments of serpulids, ammonoids and bivalves, and increases with the initial grain-size of bioclasts. The main microencrusters consist of benthic microbial communities (BMC) and nubeculariids, as well as subordinate calcareous and siliceous agglutinated foraminifera, serpulids and bryozoans. BMC are usually the first colonizers, and encrusting foraminifera mainly appear on bioclasts larger than 2 mm. BMC dominate in well-developed encrustations on upward facing surfaces of larger bioclasts that are also colonized by foraminifera (nubeculariids and Subdelloidina). Bullopora, serpulids and bryozoans are more common on lower surfaces. The fact that the values of encrustation index, encrustation thickness and diversity of the microencrusters increase with the size of bioclasts is related to a higher stability and exposure time of the available bioclastic substrate. The microencruster distribution on upper and lower surfaces of large bioclasts may be related to photic control, space competition and/or predation avoidance.     

An ammonite–fish association from the Kimmeridgian (Upper Jurassic) of La Rochelle, western France

A partially disarticulated actinopterygian fish preserved in a large three-dimensional ammonite body chamber is described from the Kimmeridgian of western France. Taphonomic observations on the degree of preservation of the fish and the development of epibiont organisms on the inner wall of the shell indicate a rather long time interval before sediment totally filled the body chamber. The fish, referred to an indeterminate Macrosemiidae, probably used this empty ammonite ( Rasenioides , Aulacostephanidae) shell as a refuge, or possibly for spawning and/or brooding. It can be assumed that ammonite shells may have constituted common shelters for demersal fishes living in an open-marine shelf environment, near to a muddy bottom devoid of rocks.     

A refined ammonite biostratigraphy for the Middleand Upper Carixian (Ibex and Davoei Zones,Lower Jurassic) in North-West Europe and stratigraphical details of the Carixian-Domerian boundary

A refined biostratigraphical scheme is described forthe Ibex and Davoei Zones (Carixian Substage, Lower Jurassic) in NW Europe, and the biostratigraphical details of the Carixian — Domerian boundary are outlined. It is based on detailed stratigraphical collections of ammonites from the major Pliensbachian localities in Britain, France and Germany, with supplementary data from Portugal. The zones and subzones, which are largely unchanged from the well-established scheme described by Dean & alii (1961), are further divided into zonules which can be recognised throughout NW Europe. These zonules, nine for the Ibex Zone and seven for the Davoei Zone, probably represent the greatest biostratigraphical resolution readily obtainable using ammonite faunas. The problems inherent in the precise recognition of the chronostratigraphical Carixian-Domerian boundary, owing to slightly discordant species ranges, are discussed.     

Ammonite diversity and its palaeobiogeographical structure during the early Pliensbachian (Jurassic) in the western Tethys and adjacent areas

The early Pliensbachian (Early Jurassic) is known as a time of marked provincialism in the marine realm, notably between the Mediterranean Tethys and North–West Europe. In order to test this observation quantitatively, we compiled 104 locality-level species lists from those areas based on a comprehensive revision of early Pliensbachian ammonites. With this dataset, we also explore the relationship between ammonite richness and biogeography at the scale of the sub-chronozone during the early Pliensbachian. Using various multivariate statistics and rarefaction techniques, we show that: (i) there is a sharp contrast between the NW European (NWE) and the Mediterranean (MED) provinces, although there is some mixing in Austroalpine and Pontic ammonite faunas; (ii) species richness in the MED province is about twice that in the NWE province for each chronozone; (iii) ammonite species richness tends to decrease during the early Pliensbachian, especially at the Ibex–Davoei transition; and (iv) the NWE and MED sensu stricto provinces both record the same pattern of variations in richness despite the fact that their taxonomic compositions have virtually nothing in common at the species level. We suggest that the low ammonite richness of the Davoei chronozone may be related to a coeval warming of seawaters, but that this was insufficient to affect the sharp palaeobiogeographic contrast between the two provinces. This persistent compartmentalisation probably reflects a major palaeogeographical structure, such as an emerged or near-emerged barrier running from the Betic range to the Briançonnais ridge. Overall, it seems that the diversity and distribution of early Pliensbachian ammonite species were simultaneously controlled by climate, palaeogeography and eustasy.     

Middle and Upper Jurassic bivalve associations from the Iberian Range (Spain)

The four sections richest in bivalves from the Middle and Upper Jurassic of the Iberian Range (Spain) were selected for a quantitative palaeoecological analysis of the bivalve fraction of the macrobenthos. Five bivalve associations and two assemblages were recognized with the help of a Q-mode hierarchical cluster analysis (Ward method). The main environmental factors controlling bivalve associations are thought to be substrate, water energy and distribution of organic matter. The bivalves exhibit a distinct spatial and temporal distribution pattern within the Aragonian Branch of the Iberian Range. Four of the bivalve associations occur in the Upper Oxfordian (Sot de Chera Fm) and one association in the Lower Callovian (Chelva Fm). In the Sot de Chera and Loriguilla formations, the abundance of bivalves decreases from NW to SE i.e., from relatively close to the shore line towards the distal-most part of the carbonate platform. In the Chelva Fm, bivalves are abundant in the Ariño region, interpreted as a palaeogeographic high. The spatial distribution of bivalves might have been largely controlled by the availability of nutrients.     

Taxonomic and biostratigraphic re-evaluation ofPerisphinctes internispinosus Krantz, 1926 (Upper Jurassic, Ammonoidea)

The lectotype ofPerisphinctes internispinosus Krantz, 1926, type species ofWindhauseniceras Leanza, 1945, from the upper Middle Tithonian of Cerro Lotena (Neuquén-Mendoza Basin, Argentina) is designated. The type series, housed in the Paläontologisches Institut der Universität Bonn, was originally illustrated by hand-drawings of two of the eleven specimens. The lectotype is illustrated by a photograph for the first time. The specimen is probably a large microconch; the most likely corresponding macroconch appears to beH. aff.steinmanni (Steuer, 1897) from Cerro Lotena.Windhauseniceras internispinosum is the index and guide species of theInternispinosum Biozone, upper Middle Tithonian, most probablyPonti Zone in age. The occurrence in Arroyo del Yeso (Mendoza) below a succession ofMicracanthoceras species includingM. mirum (Leanza, 1945) which is very close toM. microcanthum, strongly supports this correlation. The species occurs throughout the Neuquén-Mendoza Basin and probably also in the Tarapacá Basin, Northern Chile.     

The occurrence and preservation of ammonites in the Blue Lias Formation (lower Jurassic) of Devon and Dorset, England and their palaeoecological, sedimentological and diagenetic significance

More than two thirds of beds in the lowest Jurassic, Blue Lias Formation lack ammonites, which are commonly preserved in irregular or planar-bedded, bioturbated limestones, very rarely in laminated limestones and almost never in laminated black shales. Ammonites are preserved in 3D in nodular and planar-bedded limestones and at any orientation to bedding. Co-occurrence with macrobenthos and absence from beds without benthos suggest that Blue Lias ammonites were nektobenthonic. Scour structures and imbrication of ammonites in the Best Bed imply presence of traction currents. Lack of epifauna on large cephalopod shells (and other fossils) implies rapid deposition in event beds. Blue Lias deposition was episodic, not slow and continuous as the fine grain size implies. Undistorted trace fossils, uncrushed ammonites and stable isotope values all suggest early cementation of limestone beds from pore waters of a similar composition to contemporary Jurassic sea water. A clear diagenetic trend exists, with limestones having least, and laminated black shales most, modified stable isotope values. Contrast between trace fossil fills and host sediment demonstrates that Blue Lias rhythms are primary, but limestone beds have been diagenetically cemented.     

Facies and palaeoecology of Upper Jurassic (Middle Oxfordian) coral reefs in England

Summary This study documents the facies and fauna of Late Jurassic (Middle Oxfordian) coral reefs in England. Sedimentological and palaeoecological analysis of these reefs distinguishes three generic reef types: (1) small reef patches and thickets associated with siliciclastic deposits; (2) small reef patches and thickets associated with siliciclastic-free bioclastic grainstones and packstones; and (3) biostromal units associated with deep water facies. The depositional environments of these reef types are discussed. Two coral assemblages are identified: (1) the microsolenid assemblage; and (2) theThamnasteria, Isastraea, Fungiastraea andThecosmilia assemblage (Thamnasteria assemblage). TheThamnasteria assemblage developed in all shallow water environments in the study area, regardless of local environmental conditions. The fauna is very eurytopic,r-selected and can tolerate significant environmental fluctuations on short temporal scales (sub-seasonal). The main control on the development of the microsolenid assemblage was low light intensity, low background sedimentation rates and low hydrodynamic energy levels.     

Stratigraphy,facies and synsedimentary tectonics in the Jurassic Rosso Ammonitico Veronese (Altopiano di Asiago,NE Italy)

Summary The red, pelagic limestones of the Rosso Ammonitico Veronese (Upper Bajocian-Tithonian) of the Altopiano di Asiago area can be subdivided into eight facies. They differ from each other in structure (bedding style, presence and type of nodularity) and texture (nature of components, grain-vs mud-support, compactional features). Several discontinuities could be recognized, based on sedimentological or biostratigraphic evidence. In the context of a drowned platform, where sediments essentially consist of skeletal remains of both planktonic and benthic organism, the different facies are interpreted as reflecting a varying influence of currents on the winnowing of micrite and on triggering early cementation. Early cementation in turn, controlled the patterns of bioturbation and the response of sediments to later compaction and pressure-dissolution. At times, microbial mats, of unidentified nature, were important in trapping and binding sediment, giving rise to early lithified nodules and layers of stromatolitic aspect. The Rosso Ammonitico Veronese can be subdivided into three units: lower Rosso Ammonitico (RAI: Upper Bajocian-Lower Callovian), middle Rosso Ammonitico (RAM: Upper Callovian-Middle Oxfordian), and upper Rosso Ammonitico (RAS: Lower Kimmeridgian-Tithonian). Frequent ammonite moulds allow the precise dating of the base and top of each unit, and the documentation of facies heteropies and hiatusses in the more fossiliferous RAS. The lower unit (RAM) is massive and essentially nodular; the middle unit (RAM) is well bedded, non-nodular, and cherty; the upper unit (RAS) is richer in clay and typically nodular. The RAI and the RAS are present everywhere, though significant facies and thickness changes affect particularly the RAS; the RAM is much more variable, ranging from 0 to 10 metres. These variations, that may be gradual or abrupt, are inter-preted as the result of Middle-Late Callovian block-faulting which generated an irregular sea floor topography where the swells were more exposed to currents that continuously removed sediments, inducing long-lasting periods of nondeposition. Sediments preferentially accumulated in the adjacent lows. A confirmation of this hypothesis is provided by evidence of synsedimentary tectonics, described for the first time in the Rosso Ammonitico Veronese. Neptunian dykes, both vertical and horizontal, are developed at the top of the RAI and are filled with laminated sediments or collapse breccias. Glides of metre-size blocks and slumps are present at the top of the RAI and at the base of the RAM, respectively. Cm-thick layers of mud supported breccias are intercalated in the upper part of the RAI and within the RAM: they are interpreted as seismites. All these features document a tensional regime that generated fractures in more or less lithified sediments and failure along steep fault scarps or gently dipping slopes of tilted fault blocks. Recognition of this Callovian-Oxfordian tectonic activity shows that, after the Bajocian drowning, the Trento Plateau did not simply experience a uniform and general foundering: a small-scale block-faulting was still active and affected the pattern of facies distribution.     

Jurassic biostratigraphy and paleoenvironmental evolution of the Malaguide complex from Sierra Espuña (Internal Betic Zone, SE Spain)

Jurassic studies in the Internal Zones of the Betic Cordillera are scarce since this zone is composed mainly of pre-Jurassic metamorphic rocks. Only the “Dorsal” and the Malaguide domains include fossiliferous Jurassic successions, as in Sierra Espuña (SE Spain), which is one of the bigger and well-exposed Jurassic outcrops of the Internal Zones. Collected Ammonite assemblages update and improve the precision of previous biostratigraphic data by the recognition of: the Domerian (= Upper Pliensbachian, in the Mediterranean Domain) Lavinianum (Cornacaldense Subzone), Algovianum (Ragazzoni, Bertrandi, Accuratum and Levidorsatum Subzones) and Emaciatum (Solare and Elisa Subzones) Zones; the Lower Toarcian Polymorphum and Serpentinum Zones; the Middle Toarcian, Bifrons and Gradata Zone; the Upper Toarcian Reynesi Zone; the Lower/Upper Bajocian, the Lower Callovian Bullatus and Gracilis Zones; the Middle/Upper Oxfordian Transversarium, Bifurcatus, Bimammatum and Planula Zones; and the Lower and Upper Kimmeridgian Platynota, Strombecki, Divisum and Beckeri Zones.The paleoenvironmental evolution of the Malaguide Jurassic at Sierra Espuña shows similarities with other Mediterranean Tethyan paleomargins. The biostratigraphic precision along with the litho- and biofacies analyses has enabled the interpretation that the Malaguide paleomargin evolved as a passive margin, developing shallow carbonate platforms, until the Domerian (Lavinianum Zone). Then, the platform broke up (Domerian, Lavinianum Zone-Upper Toarcian, Reynesi Zone) with the beginning of the rifting stage, beginning the development of horst-graben systems and the coeval drowning of the area. This stage ended in the upper Lower Callovian (Gracilis Zone) to the Middle Oxfordian (Transversarium Zone) interval, starting the drifting stage, which accentuated the horst-graben systems, leading to the deposition of condensed nodular limestones in the raised sea bottom.     

Ammonite eggs and ammonitellae from the Kimmeridge Clay Formation (Upper Jurassic) of Dorset, England

Eight clusters of small spherical and subspherical objects, some isolated and some associated with shells of perisphinctid ammonites, have been recovered from the Lower and Upper Kimmeridge Clay (Upper Jurassic) of the Dorset coast, England. They have been interpreted as ammonite egg sacs and represent the freshest and best-preserved examples known so far. Their structures and the ecological framework in which they occur are discussed. The parents are thought to be members of the two eudemic genera Aulacostephanus and Pectinatites that dominate the biostratigraphy of the ammonites in the range of the Kimmeridge Clay in which they occur. Isolated nuclei of ammonitellae have also been recovered.