Marine influence and incursion in the Gondwana basins of Orissa, India: A review

The Permian–Triassic succession of the Indian Gondwana Sequence was previously considered to have been deposited in a fluviatile-lacustrine environment. Similarly, earlier Lower Gondwanas of Orissa State (a major part of the Mahanadi Master basin) were considered entirely fresh water deposits. Faunal evidence is still scanty in this master basin. Ichnology and palynology along with a few sedimentary records are reviewed and analysed for inferring marine signature. The marine nature of the Talchir, Karharbari, Barakar, Barren Measures and Kamthi sediments of three major basins (Talcher, Ib River and Athgarh) in Orissa State was predicted on the basis of typical marine ichnofossils. Most of these sediments also contain acritarchs reflecting marine marginal environment throughout the Permian. Moreover, evidence of wave activity, salinity raise and discovery of phosphorite in Permian sediments also strengthen this view.Hence, the previous model of continental facies for the Lower Gondwanas is found to be incorrect. The ichnofossils (Skolithos and Cruziana ichno-facies), acritarchs (Foveofusa, Leiosphaeridia, Greinervillites, etc.) and other palynofossils of marine origin can be utilized as a tool for palaeoenvironmental reconstruction. In the Gondwana basins of Orissa (Mahanadi Master basin), consistent occurrence of marine acritarchs and trace fossils with some typical sedimentary structures such as wave ripples has been studied and reviewed from the Talchir (Early Permian) to Upper Kamthi (Triassic) formations at various time intervals. Here marine incursion could have occurred due to the well known global transgressions during Permian and Triassic.     

山东滕县煤田石炭纪、二叠纪孢粉植物群及古气候

根据前人在原位孢子方面已积累的大量资料,并参考现代孢子花粉形态研究成果,大致推测了本区石炭-二叠纪分散孢子花粉的大多数属的植物亲缘关系。通过已建立的含煤地层孢粉组合带推测相应时期的古植被类型,并与前人有关地质时期及地层的大植物化石带进行了对比,不仅验证了各地质时期植被的兴起、发展、繁盛以及衰退绝灭的历史过程,更大意义上将微体形态的孢子花粉与宏观上的植被进行了对比归类,从而使得对研究区晚古生代含煤地层的古植被的形态面貌有了更直观的了解。最后,利用孢粉分析恢复相应地质时期的古气候。     

Permian, Jurassic and Early Cretaceous palynofloral assemblages from subsurface sedimentary rocks in Chuperbhita Coalfield, Rajmahal Basin, India

The results of a palynological analysis of the sedimentary sequence of Borehole RCH-151, Chuperbhita Coalfield, Rajmahal Basin, Bihar are presented here. The borehole penetrated the Rajmahal Formation (comprising two traps sandwiching an intertrappean bed), the thinly represented Dubrajpur Formation and in its lower part, the Coal Measures. The coal-bearing interval is associated with Scheuringipollenites barakarensis, Faunipollenites varius, Densipollenites indicus, Gondisporites raniganjensis and Densipollenites magnicorpus Assemblage Zones. The presence of these biostratigraphic units indicates correlation with the Barakar Formation (Early Permian) and the Barren Measures and Raniganj Formations (both Late Permian). This is the first record, in the Chuperbhita Coalfield, of Late Permian strata, which appear to represent a condensed sequence. Prior to the present study, the Permian succession was thought to have been associated entirely with the Barakar Formation. The overlying Dubrajpur Formation yielded a distinct spore-pollen assemblage (in association with the first report of dinoflagellate, Phallocysta), which is assigned to the newly identified Callialasporites turbatus palynozone of latest Early to early Middle Jurassic age. The diverse spore-pollen flora of the intertrappean bed (Rajmahal Formation) incorporates several age marker taxa, viz. Undulatisporites, Leptolepidites, Klukisporites, Ruffordiaspora, and Coptospora. The assemblages from intertrappean beds are correlated with the Ruffordiaspora australiensis palynozone of Australia. Thus the palynodating indicates Permian, latest Early to early Mid-Jurassic and Early Cretaceous age for the strata studied. This is the first record of definite Jurassic microfossils from the non-marine sequence of Rajmahal Basin, India.     

印度Mahanadi盆地井下二叠纪下冈瓦纳岩层的微体植物化石组合

Mahanadi河盆地的冈瓦纳型诸盆地位于印度的近中心地区,呈北西—东南走向分布。Mahanadi主要盆地内的几个重要煤田为Korda煤田,Mand—Raigarh煤田和Ib河煤田。Korda煤田的冈瓦纳系沉积向南东向延伸,经Mand—Raigarh煤田延续至Ib河煤田。Korda和Ib河两个煤田储量巨大,且长期来有小构造活动;而Mand—Raigarh煤田至今很少受到重视,勘探开发晚得多。文中首次对Mand-Raigarh煤田5个钻孔样品作了孢粉研究,以初现面、相对丰度等为特征,划分出7个孢粉组合带,时代从二叠纪最早期(Talchir组)至二叠纪末期(Raniga由组)。填补了东西两个煤田之间的生物地层研究空白。7个孢粉组合带为：i)Parasaccites-Plicatipollenites assemblage zone(相当于Talchir组）,ii)Sulcatisporites-Brevitriletes assemblage zone(相当于下Barakar组）,iii)Sulcatisporites-Rhizomaspora assemblage zone(少量Densipollenites存在说明其相当于上Barakar组）,iv)Densipollenites-Striatopodocarpites assemblage zone(相当于Barren Measures组）,v)Striatopodocarpites-Densipollenites assemblage zone(相当于下Raniganj组）,vi)Striatopodocarpites-Densipollenites-Sulcatisporites assemblage zone(相当于上Raniganj组）vii)Sriatopodocarpites-Alisporites assemblage zone(相当于Raniganj最上部到Kamthi组最下部）,Raniganj组沉积中见到小的皱缩裂隙、植物垂直根和根的印痕,似代表古土壤。     

The oldest record of a boring foraminifer: Early Permian of New Mexico,USA

The bioeroding foraminifer Troglotella incrustans Wernli and Fookes (Bolletino della Societa Paleontologica Italiana 31, 1992, 95), is widely reported from Bajocian?, and Oxfordian to Lower Cenomanian (with a Late Jurassic acme) shallow‐water limestones of the Tethyan realm. A single specimen of a boring foraminifer, assigned to T. incrustans, has now been observed from the Lower Permian (Sakmarian) Community Pit Formation of the Doña Ana Mountains, New Mexico, USA. Surviving the end‐Permian mass extinction, T. incrustans might be a Lazarus taxon that persisted in refuges. This finding represents the oldest record of a foraminifer exhibiting an euendolithic way of life. Boring foraminifera have not been previously recorded from strata older than the Jurassic. Boring traces of potentially foraminiferan origin, however, have been already reported from the Lower Carboniferous (?Ordovician).     

Lower Permian Bryozoa of the Jamal Formation from Bagh-e Vang (Shotori Mountains,northeast Iran)

Lower Permian Bryozoan fauna of Jamal Formation, exposed in Bagh-e Vang (Shotori Mountains, northeast Iran) includes six species. Three species – Streblotrypa (Streblascopora) marmionensis (Etheridge 1926), Rhabdomeson bispinosum (Crockford 1944) and Alternifenestella kungurensis (Stuckenberg 1898) – indicate the Lower Permian (Artinskian to Kungurian) age of the formation. Three additional taxa – Fistulipora sp. 1, Fistulipora sp. 2 and undetermined timanodictyid bryozoan ?Timanodictya sp. could not be identified at the species level. The investigated fauna refers to Uralian and Australian palaeobiogeographic provinces.     

A GLOSSOPTERIS FLORA FROM THE OHIO RANGE,ANTARCTICA

Cridland , Arthur A. (Ohio State U., Columbus.) A Glossopteris flora from the Ohio Range, Antarctica. Amer. Jour. Bot. 50(2): 186–195. Illus. 1963.—Leaves of Glossopteris indica are the commonest fossils in the Mount Glossopteris Formation on Mount Schopf. Other fossils present are: Glossopteris ampla, G. anguslifolia, G. damudica, G. browniana, detached scales, sporangia of Arberiella containing bisaccate pollen grains, seeds of Samaropsis (principally S. longii), and some obscure fossils. These remains, and associated animal fossils, suggest that the Mount Glossopteris Formation is Permian. The plants evidently grew in a seasonal but favorable climate. These growth conditions indirectly support paleomagnetic calculations which indicate that in Late Permian time, Mount Schopf lay near latitude 50°S.     

New species of Dzhulfian (Late Permian) ammonoids from the Hambast Formation of Central Iran

New ammonoid species of the genus Araxoceras (A. abarquense sp. nov. and A. iranense sp. nov.) are described from the Early Dzhulfian (Early Wuchiapingian) beds of the Hambast Formation of Central Iran (Abade). The data on the distribution of Paraceltites (Paraceltites sp.) and Vedioceras (V. umbonovarum Ruzhencev), for the first time discovered in the Dzhulfian beds of Abade, and of Eoaraxoceras are provided. The correlation of the Late Permian assemblages of ammonoids of Iran and Transcaucasia is amended.     

Scorpionid Resting Trace from The Lower Permian of Southern New Mexico,USA

We describe an arthropod body impression associated with arthropod trackways of the ichnogenus Stiaria from the Lower Permian (upper Wolfcampian) Robledo Mountains Formation (Hueco Group) in the Prehistoric Trackways National Monument of southern New Mexico. The probable producer of these traces was a scorpion, and we name the likely scorpionid resting trace Alacranichnus braddyi, new ichnogenus and ichnospecies. There are no prior reports of scorpionid body impressions from terrestrial settings in the fossil record.     

Permian algae and algal microfacies from unayzah,quassim district,saudi arabia

Summary Late Permian bioclastic calcarenite beds of the middle Khuff Formation were sampled for their algal constituents near the city of Unayzah, Quassim district in central Saudia Arabia. The algal flora includes two species of udoteacean algae (Succodium difficile andSuccodium sp.), the dasycladacean algaMizzia velebitana and two species of red algae (Gymnocodium bellerophontis, Permocalculus plumosus). Other algal floral remains found forming isolated single layers which have generally been named algal microfacies. These include a phylloid microfacies, an oncoid microfacies and algal stromatolites. The Khuff Formation is well-known for its accumulation of non-associated gas and particularly for its oil accumulation in eastern Saudi Arabia and the Arabian Gulf states.     

Insect traces on early Permian plants of India

The study of fossil leaves from the Glossopteris flora of the Lower Permian Gondwana sequence of the Barakar Formation of the Raniganj Coalfield, India, has revealed the presence of different types of insect traces. The structures are compatible with a variety of insect activities on fossil leaves. Nibbled and cuspate margins, trench marks, obliterated surfaces, blotch marks, and holes of various shape constitute feeding traces, whereas egg pouches along the midrib and irregularly distributed oviposition marks are traces of egglaying. Although fossil records of both insects and their activities in the Glossopteris flora of India remain scarce, the different types of insect traces identified in the present study demonstrate the existence of a diverse insect fauna during the Permian Period of India.     

FIRST REPORT OF THE FERTILE PLANT GENUS UMKOMASIA FROM LATE PERMIAN BEDS IN INDIA AND ITS BIOSTRATIGRAPHIC SIGNIFICANCE

Abstract:  The genus Umkomasia Thomas, a female fructification of Dicroidium , is reported from Late Permian beds exposed in the Behra Rivulet near the village of Karaonda in the Tatapani-Ramkola Coalfield, Chhattisgarh, India. This is the first record of this genus from the Late Permian, and from Indian Gondwana. Two species have been recognized: Umkomasia polycarpa Holmes and U. uniramia Axsmith, Taylor, Taylor and Cuneo. The find suggests that the genus Dicroidium appeared in the Late Permian before reaching its acme in the Middle–Upper Triassic.     

Lower Permian Bryozoa from Ellesmere Island (Canada)

A collection of fossils sampled during the 1898–1902 expedition of theFram to the Canadian Arctic Islands includes abundant bryozoans from the Lower Permian (Artinskian) Great Bear Cape Formation of Ellesmere Island. From this material a new genus with one new species —Nansenopora peculiaris n. gen., n. sp. — as well as three new species —Streblotrypella arctica n. sp.,Phragmophera patricki n. sp. andKallodictyon spinatum n. sp. — are described. Furthermore, the speciesUlrichotrypa ramulosa Bassler, 1929 is reported for the first time from the Lower Permian of the Arctic region.     

Large Pelycosaur Footprints from the Lower Pennsylvanian of Alabama,USA

Relatively large tetrapod footprints from Lower Pennsylvanian strata of the Pottsville Formation in the Warrior basin coal field of Alabama were previously assigned to the ichnogenus Attenosaurus. However, three morphotypes are present, and we assign these tracks to Attenosaurus, a new ichnogenus (Alabamasauripus) and Dimetropus. Most of these tracks apparently were made by large pelycosaurs, for which there is no bone record older than Late Pennsylvanian.     

Evolutionary trend of Zoophycos morphotypes from the Upper Cretaceous–Lower Miocene in the type pelagic sections of Gubbio,Italy

About 200 Zoophycos specimens, including 90 specimens studied in detail, have been analysed in the continuous Upper Cretaceous–Lower Miocene pelagic sedimentary type sections of the Gubbio area (the Contessa Highway, Contessa Quarry and Bottaccione sections, Northern Apennines). The sediments are reddish to grey limestones and marls of the Scaglia Group and marls with volcaniclastic deposits of the Bisciaro Formation. The aim was to examine the evolutionary trend of what is probably the most debated trace fossil of all time, from the Upper Cretaceous to Lower Miocene. Despite having been found in beds ranging from the Cambrian to the present, no consensus has been reached regarding mode of construction, tracemaker or ethological explanation for Zoophycos. Four Zoophycos morphotypes are recognized at Gubbio showing variations of major and minor lamellae, apex, lobes and whorls: the Cretaceous–Eocene cone‐shaped type 1, the Upper Eocene–Middle Oligocene helicoidal type 2, the Oligocene lobate type 3 and the Upper Oligocene–Lower Miocene flat type 4. The very high ichnodensity in some beds (hundreds of specimens in discrete levels of the Bisciaro Formation, now destroyed by quarrying) seems to find explanation in abnormal concentrations of phytodetritus and organic matter on the seafloor in some periods. This very high abundance in discrete levels reflects a change in sedimentation and seafloor conditions at pre‐flysch deposition. Due to such high ichnodensity, many adjacent specimens display deformed outer margins. Taphonomic analysis shows a variation of whorls, laminae and U‐shaped lobes, reflecting ontogenetic development of the tracemaker(s) (?sipunculid worms).     

Palynostratigraphy of subsurface Upper Permian and Mesozoic Succession, Rakshitpur area, Raniganj Coalfield, West Bengal

In borehole RRK-1, Rakshitpur, Raniganj Coalfield in Damodar Basin, the lithounits identified in the interval of 115.00–610.00 m are in unconformable contact with each other. Tertiary sediments (115.00–179.70 m) that contain Callialasporites, Araucariacites, and Podocarpidites show continuity with the underlying supra-trappean strata (179.70–185.70 m). The two palynozones identified within the Rajmahal Formation (Foraminisporis asymmetricus 179.70–185.70 m and Foraminisporis wonthaggiensis 198.92–216.63 m) suggest Valanginian to Barremian age. The chocolate facies of the Panchet Formation (253.70–471.70 m) is devoid of palyniferous matter except at 260.55 m depth where the Foraminisporis wonthaggiensis assemblage of the younger horizon (inter-trappean) continues to occur. Downwards, at 261.20–262.40 m depth, Permian taxa Striatopodocarpites, Faunipollenites, and Densipollenites are abundant, but Arcuatipollenites pellucidus, Playfordiaspora cancellosa, and Goubinispora indica are rare. Besides, Callialasporites turbatus/C. microvelatus, first occurred at 262.40 m depth, are still present in the Panchet Formation. This kind of mixed population of palynomorphs is suggestive of disturbed zone in the deposite, and had happened at the on-set of volcanic activity in the study area, and re-defines the status of this part (253.70–262.40 m) to be infra-trappean. Occurrence of pebble bed at 262.40–262.80 m further supplements this hiatus. In the coal horizon (471.70–610.00 m), two identified palynoassemblages, Densipollenites magnicorpus and Gondisporites raniganjensis, indicate Late Permian age for this part of the Raniganj Formation. Abundant wood shreads, plant tissues in the chocolate facies of the Panchet Formation and at intervals in the Coal horizon suggest high-energy flow during sedimentation in the study area.     

Tetrapod footprints of the lower permian choza formation at Castle Peak, Texas

The tetrapod track ichnofauna of the Lower Permian Choza Formation at Castle Peak, Texas, consists after augmentation of sample size and revision of three distinet ichnotaxa:Erpetopus willistoni Moodie, 1929,Varanopus curvidaetylus Moodie, 1929 andDromopus palmatus (Moodie, 1929). These are the tracks of protorothyridis, captorhinomorphs and araeoscelids, respectively. Castle Peak is the type locality of the ichnogeneraErpetopus andVaranopus. With the tetrapod ichnofauna of the Robledo Mountains Formation in New Mexico, the Castle Peak ichnofauna is the second Lower Permian track oecurrence that can be clearly correlated to the marine timescale. Both tetrapod ichnofaunas support assigning an age up to Artinskian to most Lower Permian red-bed tetrapod ichnofaunas of the North American Wolfcampian-Leonardian and the European Rotliegendes. The ichnogeneraBatrachichnus, Limnopus, Amphisauropus, Erpetopus, Varanopus, Dromopus andDimetropus characterize an Artinskian red-bed ichnofauna of Euramerican distribution.      

Annalepis, a pioneering lycopsid genus in the recovery of the Triassic land flora in South China

Fossil plants are scarce in the Earliest Triassic marine deposits of western Guizhou and eastern Yunnan. Only Annularia shirakii, Lobatannularia sp., Paracalamites stenocostatus, Gigantopteris sp., Pecopteris sp. were reported from the base of the Kayitou Formation dated as Early Induan by marine fauna. Recently, we discovered numerous representatives of the genus Annalepis in the same Lowermost Triassic beds: A. latiloba, A. brevicystis, A. angusta, Annalepis spp. occur associated with a basal Triassic marine fauna. This discovery fills the biostratigraphic gap between the Late Permian “Gigantonoclea guizhouensis-Ullmannia cf. bronnii-Annularia pingloensis” and the late Lower Triassic “Neuropteridium–Albertia–Voltzia” assemblages reported from South China. It represents an important datum dealing with the very beginning of a new terrestrial flora installation after the Permian flora disappearance following the Permian–Triassic boundary mass extinction. This “starting point” of a new vegetal cover in South China is to be taken into account in reconstructing through space and time the settlement process of the Mesozoic floristic provinces.     

烟叶大羽羊齿Gigantopteris nicotianaef olia Schenk的时空分布和相关地层的对比

烟叶大羽羊齿Gigantopteris nicotianae f olia Schenk是地质和古生物工作者熟知的植物化石。由于Halle（1927）曾将单网羊齿的种误归并于这一种内,致使它一度成为广见于华夏古植物区二叠纪陆相地层的植物化石。后来的研究表明（Guand Zhi,1974;姚兆奇,1983;Glasspool et al．,2004）,除了河南平顶山的可疑标本外（Sze,1954）,此种可作为华南冷坞期的标志性植物。最新资料证实（杨关秀等,2006）,产自河南禹州和临汝上石盒子组（云盖山组）的标本完全显示了G．nicotianaefolia的特征,说明了这个种在华北的存在,从而扩大了这种植物的分布区。根据烟叶大羽羊齿的时空分布规律,文章认为河南的上石盒子组完全可与江苏龙潭组和江西乐平组老山段下亚段的含煤层位对比,时代应属瓜德鲁普世冷坞期（Capitanian）。文中较详细阐述将河南上石盒子组的时代定为冷坞期而不是晚二叠世早期（吴家坪期）的观点,并引证磁性地层方面证据,支持这一结论。     

A new scanilepiform from the Lower Triassic of northern Gansu Province,China, and phylogenetic relationships of non‐teleostean Actinopterygii

A new scanilepiform, Beishanichthys brevicaudalis gen. et sp. nov. , is named and described based on fossils from the Lower Triassic lake deposits exposed in Beishan area, Gansu Province, China. The discovery documents a new record of this group, which is significantly older than other known scanilepiforms from China, and is slightly younger than Evenkia from the Lowest Triassic of Central Siberia. Although the Beishan beds were previously interpreted as Late Permian in age, based on megaplant fossils, this new discovery supports the reinterpretation of the deposits as Early Triassic in age, based on vertebrate fossils from the same locality and horizon. Phylogenetic analysis was conducted to resolve the relationships of Scanilepiformes with other actinopterygian clades, and the inter‐relationships within Scanilepiformes. Contrary to previous thought that scanilepiforms are closely related to the Amiidae, the phylogenetic results of this study recognize the Scanilepiformes as stem‐group neopterygians. Relationships of the Scanilepiformes and Australosomus with other neopterygians remain unresolved. With a characteristic long‐based dorsal fin, scanilepiforms represent a small group that emerged in Early Triassic freshwater environments, inhabited Eurasia and North America during the Middle–Late Triassic, briefly invaded the marine environment by the Late Triassic in Europe, and became extinct at the end of Triassic. © 2011 The Linnean Society of London, Zoological Journal of the Linnean Society, 2011, 161 , 595–612.