Paleoenvironmental conditions preceding the Messinian Salinity Crisis in the Central Mediterranean: Integrated data from the Upper Miocene Trave section (Italy)

Integrated data of calcareous plankton and benthic foraminifers from the pre-evaporitic interval of Trave section (Central Italy) allowed the reconstruction of surface and bottom-water conditions in the Central Mediterranean during the interval from 7.61 to 6.33 Ma, preceding the Messinian Salinity Crisis.Our data point out a three-step paleoenvironmental evolution. During the first stage (7.61-7.02 Ma) benthic foraminiferal assemblages depict stable, well-oxygenated and ventilated bottom-water conditions, while the surface water records variable temperature and high nutrient conditions, probably associated with strong seasonality. The second stage (7.02-6.70 Ma) points to unfavourable bottom-water condition, triggered by deep-sea stagnation. This is witnessed by a significant decrease in oxygen concentration and biotic diversity, and by the presence of stress-tolerant taxa. A general warming of the surface water and a strongly stratified water column, characterized by an expanded mixed layer, are also recorded.From 6.70 Ma onwards (third stage), a prominent change to more restricted, low-oxygenated, hypersaline conditions at the sea floor is testified by the total disappearance of deep-dwelling planktonic foraminifers and the increasing abundance of stress-tolerant species. Calcareous plankton reflects high instability of the surface water in terms of nutrients, temperature and salinity. During this stage the environmental deterioration reaches intermediate depths in the water column.The initial change toward a step-wise isolation of the Central Mediterranean bottom-waters is probably related to a general warming, responsible for a first slowing-down of the vertical circulation, favouring stratification of surface and intermediate waters and stagnation of bottom-waters. This warming is related to the restricted connection between the Mediterranean Sea and the Atlantic Ocean, which occurred since 7.146 Ma.In the Trave section, the isolation of bottom-waters most likely occurred at the same time as in other Mediterranean sections. However, due to the presence of a hiatus it cannot be excluded that it occurred with a delay of ~ 100 kyr, probably related to the shallower paleodepth of the basin.     

Nannoplankton biostratigraphic calibration of the evaporitic events in the Neogene Fortuna Basin (SE Spain)

The Fortuna Basin is an example of a marginal Mediterranean basin with evaporitic sedimentation during the Late Tortonian and Messinian. This basin shows an early restriction event before the main Messinian Salinity Crisis (MSC) that allows the Tortonian Salinity Crisis (TSC) to be proposed as a tectonic uplift event isolating the eastern Betic basins. Four evaporitic events are present in the central part of the Fortuna Basin, from bottom to top: Los Baños Marls Formation (composed by Fenazar Conglomerate Bed, Lower Gypsum Member [Mb] and Sanel Mb), Tale Gypsum Formation (Fm), Chicamo Diatomites and Gypsum Cycles Fm, and Rambla Salada Gypsum Fm. The present work documents the first biostratigraphic dating based on calcareous nannoplankton of these events. The lowest occurrence (LO) of Amaurolithus primus is registered at the upper part of the Sanel Mb, below the Tale Gypsum Fm. The LOs of Amaurolithus delicatus and Reticulofenestra rotaria, which mark the base of the Messinian, occur in the lower part of the Chicamo Cycles Fm, above the Tale Gypsum Fm, the Triquetrorhabdulus rugosus-Nicklithus amplificus integrate form and the LO of Nicklithus cf. amplificus in the upper part of the Chicamo Cycles Fm. Taking into account these results, a new calibration of the available magnetostratigraphic data is presented: the Chicamo Cycles Fm were formed during the reverse chron C3Ar and the Tortonian-Messinian boundary should be found within the Tale Gypsum Fm or near the top of the Sanel Mb. The onset of the TSC, the first restriction phase of the Fortuna Basin, is represented by the Fenazar Conglomerate Bed, bottom of the Los Baños Fm, and not by the Tale Gypsum Fm, as previously considered.     

Calcareous nannofossils and foraminifers herald the Messinian Salinity Crisis: The Pollenzo section (Alba, Cuneo; NW Italy)

During the Messinian, the Mediterranean area experienced fast and prominent paleoenvironmental changes, culminating in the so-called Messinian Salinity Crisis, with the deposition of the evaporitic series. This work investigates the micropaleontological assemblages in the pre-evaporitic sediments of the Sant’Agata Fossili Marls (SAF) of the Pollenzo section (Cuneo area, North Western Italy). A semiquantitative analysis is carried out on the upper part of the marly and pelitic sediments of the SAF underlying the first gypsum bed, ascribed to the Vena del Gesso Fm. (VDF). The studied interval belongs to the planktonic foraminifer Globorotalia conomiozea Zone and “non distinctive Zone” of Iaccarino and to the calcareous nannofossil MNN11b/c Zone of Raffi et al. (1998, 2003) ( [Raffi et al., 1998] and [Raffi et al., 2003]). Decrease of diversity and abundance of the foraminifer and calcareous nannofossil assemblages is recorded 12 m below the VDG and clearly reflects environmental stress. From bottom to top, six paleoecological events are recorded: (1) the first peak abundance of “small” Reticulofenestra and the last recovery (LR) of planktonic foraminifers; (2) the peak abundance of Pontosphaera japonica and the last recovery of warm water taxa Discoaster spp.; (3) the last recovery of benthic foraminifers; (4) the co-occurring peak abundances of Helicosphaera carteri and Sphenolithus abies, and the last recovery of warm water taxa Amaurolithus spp.; (5) the second peak of “small” Reticulofenestra; (6) the definitive disappearance of calcareous nannofossils. These paleoecological events describe a progressive isolation of the basin from the world ocean and increasingly stressed environment (LR planktonic foraminifers; LR Discoaster spp.), increasing dysoxic to anoxic conditions at the sea floor (LR benthic foraminifers), shallowing of the water column (peak of H. carteri), increasing salinity in surface waters (peak of S. abies), and enhanced nutrient concentration in surface waters (peak of “small” Reticulofenestra); these are related to paleoenvironmental changes predating gypsum deposition at Pollenzo and affecting the whole Mediterranean basin.     

Sierra del Colmenar 1A,a new late Messinian (Late Miocene) locality in the Bajo Segura Basin (SE Spain): biostratigraphic and palaeoenvironmental implications

Abstract

In this paper, we have studied a new micromammal site from the Sierra del Colmenar section (Elche, SE Spain), named Sierra del Colmenar 1A (SCO-1A), representing the uppermost levels of Messinian age of the Bajo Segura Basin. The sedimentary context of this locality corresponds to a costal lagoon with marine influence. The fossil site has yielded remains of Apodemus aff. gorafensis, Paraethomys meini, Apocricetus alberti, Occitanomys alcalai, Ruscinomys sp., Eliomys cf. truci, Muscardinus sp., Parasorex ibericus, Prolagus michauxi and Soricidae indet. Based on this assemblage, we propose a Late Miocene age (MN13) for the locality SCO-1A, matching the previously inferred age for the stratigraphic unit in which the fossil site is situated. According to its faunal composition, we infer warm and humid climate conditions in the moment of formation of this locality, agreeing with the environmental conditions of the Messinian age in a parallic sedimentary context.     

Integrated quantitative biostratigraphy of the latest Tortonian-early Messinian Pissouri section (Cyprus): An evaluation of calcareous plankton bioevents

A quantitative study was performed on planktonic foraminifera and calcareous nannofossils of the astronomically dated Late Miocene Pissouri section (Cyprus). Our results confirm the reliability of well-known planktonic foraminiferal events as Catapsydraxparvulus LO (Last Occurrence), sinistral coiling change of Globorotaliascitula, Globorotaliamiotumida group FRO (First Regular Occurrence), Globorotalianicolae FO (First occurrence) and LO, sinistral coiling change of Neogloboquadrinaacostaensis and also of several important calcareous nannofossil events (Amaurolithusprimus FO, Amaurolithusdelicatus FO, Reticulofenestrarotaria FO and FCO). Integrated planktonic foraminifera and calcareous nannoplankton data contribute to an enhanced time resolution of the Tortonian - early Messinian interval in the Levantine basin, and contribute to detailed correlations throughout the Eastern Mediterranean. In addition, we compare methodologies commonly used in calcareous plankton biostratigraphy, and shortly outline sources of bias that can influence the results of stratigraphic studies.     

The Late Tortonian-Early Messinian Foraminiferal Record Of The Abad Member (Turre Formation), Sorbas Basin, Almería, South-East Spain

The Late Tortonian-Early Messinian Abad Member in the Sorbas Basin, south-east Spain contains abundant foraminifera. Planktonic foraminifera have been used to reconstruct the environmental conditions that developed within the basin. Benthonic foraminifera reveal a detailed picture of the palaeoenvironment. After an initial deepening episode, from upper epibathyal ( c . 200 m) depths to lower epibathyal ( c . 1000 m) depths, the basin progressively shallowed to shelf depths ( c . 100 m); this palaeobathymetric reconstruction thus describes an initial rapid transgressive episode, a highstand and a regressive phase; the basin water mass became progressively warmer, and yet there was no significant increase in salinity or oxygen level in the water mass. The lack of any microfossil evidence to support an increase in the salinity levels towards the top of the Abad Member indicates a rapid environmental switch from normal marine conditions during the deposition of the Abad Member to hypersaline conditions during deposition of the overlying Yesares Member and precipitation of great thicknesses of gypsum. This research illustrates the necessity for high resolution sampling, approximate to the standard interval (10 feet: c . 3 metres) used in industrial boreholes, for Mediterranean Late Miocene stratigraphies, and indicates that all previous palaeoenvironmental studies on the Sorbas Basin have had inadequate sampling regimes which do not represent the evolving environmental conditions of the Abad Member.     

Late Miocene Erinaceinae from the Teruel Basin (Spain)

Classifying fossil teeth of Erinaceinae (spiny hedgehogs) is a challenging task, because of their scanty record and systematic treatment that heavily relies on skull characteristics. In this paper we describe the complete set of isolated dental elements of Erinaceinae from the upper Miocene sediments of the Teruel Basin (eastern Central Spain). Four different species were recognized: Postpalerinaceus cf. vireti, Atelerix aff. depereti, Atelerix steensmai nov. sp., and a form classified as Erinaceinae genus and species indet. All four are relatively derived in showing multi-purpose dentitions, not showing only adaptations to insectivory, but also to carnivory, herbivory and possibly durophagy/malacophagy. The temporal occurrence of spiny hedgehogs during the middle to late Miocene in the Teruel Basin and neighboring Calatayud-Montalbán Basin peaks within periods of relative aridity, a correlation consistent with modern geographic distribution. Messinian cooling is the best candidate for explaining a remarkable demise of Erinaceinae at 7 Ma.     

Reproducibility and reliability of the Pliensbachian calcareous nannofossil biohorizons from the Basque-Cantabrian Basin (Northern Spain)

Based on calcareous nannofossil assemblages identified in four expanded and well-dated sections from the Basque-Cantabrian Basin, the main objective of this paper is to improve the knowledge of the Pliensbachian calcareous nannofossil events, and to calibrate these events to the ammonite zones established for this area. The semiquantitative analysis of more than 140 smear slides from the Tudanca and Santotis sections, and the re-analysis of more than 200 smear slides from the Camino and San Andrés sections, have been carried out in order to describe the succession of calcareous nannofossil assemblages. Related to their composition changes, we have recognized and calibrated to the ammonite zones two main events: the first occurrences (FOs) of Similiscutum cruciulus and Lotharingius hauffii, and six secondary events: the FOs of Biscutum novum, Biscutum grande, Biscutum finchii, Lotharingius barozii and the FCOs (first common occurrences) of Calcivascularis jansae and L. hauffii. In the Camino and San Andrés sections, we also identify the FOs of Biscutum dubium, Bussonius prinsii and Lotharingius sigillatus. The obtained data allow us to assess the degree of reproducibility of the Pliensbachian calcareous nannofossil events in the studied area. The biohorizon succession recognized in the Basque-Cantabrian Basin are compared to those proposed for NW Europe, Lusitanian Basin, Italy and Southern France.     

Spatial distribution of Late Pliensbachian (Early Jurassic) calcareous nannofossils within the Lusitanian basin (Portugal)

Quantitative analyses of Pliensbachian calcareous nannofossils have been carried out on a proximal-distal transect in the Lusitanian Basin (Portugal). The studied sections Vale Venteiro near Tomar and Peniche represent proximal and distal environments with respect to the emerged land of the Iberian Meseta (to the East). The upper portion of the Vale das Fontes Fm (Davoei and Margaritatus ammonite Zones) is studied in both sections and correlated by means of ammonite and nannofossil integrated biostratigraphy. A careful analysis of the preservation state of nannofossils is performed. Preservation state is moderate to good in the two settings; changes in nannofossil assemblages are therefore considered as primary. Samples were analysed for nannofossil absolute and relative abundances, species diversity and wt%CaCO₃. The analysis of calcareous nannofossils (absolute abundance, percentage, average abundance) shows changes in the community structure in space (i.e., from proximal to distal), and vertically within the water column. This pattern suggests a partitioning of habitats within the photic zone, and with respect to emerged lands. Proximal environments within the Lusitanian Basin were probably more eutrophic, because of the proximity of emerged lands to the East (Iberian Meseta). This was the source area for nutrients delivered into the basin by river transport. Distal environments were likely characterized by a greater water depth and by a deeper light penetration leading to a relatively more expanded photic zone. The assemblages recorded in the proximal setting exhibit high mean relative abundance of placolith-bearing coccolithophorids (Lotharingius spp., Biscutum spp. and Similiscutum spp.) and of Schizosphaerella spp., while distal assemblages are dominated by Crepidolithus crassus and Schizospharella spp. Crepidolithus crassus is interpreted as a deep-dweller coccolithophorid, inhabiting preferentially distal and deeper settings in the Lusitanian Basin. Placolith-bearing coccolithophorids were more abundant in proximal settings with respect to emerged lands, under relatively elevated trophic conditions. The probable calcareous dinocyst Schizospharella spp. proliferated in surface waters of both proximal and distal environments.     

New charophyte assemblage from middle Miocene lacustrine deposits of Moneva (Ebro Basin,Spain)

Two new fossil charophyte species from middle Miocene lacustrine deposits of Moneva (Ebro Basin, NE Spain) are described and illustrated. Sphaerochara miocenica nov. sp. is represented by medium sized gyrogonites with a characteristic spheroidal shape and ornamented with numerous small and regularly spaced tubercles arranged along the spiral cells. Several specimens show an unusual carbonate encrusted layer that covers completely or partially the gyrogonite. Psilochara monevaensis nov. sp. is characterized by large gyrogonites, with typical prolate shape and protruding apical poles. A second population of S. miocenica nov. sp. from a distant early upper Miocene lacustrine basin located in the Middle East (Bekaa Valley, Lebanon), identified in a previous study as Sphaerochara sp., is described and illustrated here, allowing the comparison of these distant populations. Ecologically, the associated fauna and facies from both localities suggest that these taxa grew in very shallow athalassic lakes. Interestingly, S. miocenica nov. sp. having a wide geographic distribution in peri-Mediterranean lakes with variable salinity, may become a fossil guide for the Miocene, providing at the same time important palaeoecological information.     

Hipparion material from Gaojiashan locality in the Late Miocene of Linxia Basin,Gansu, China and associated mammalian fossil assemblage

Two Hipparion species, H. forstenae and H. hippidiodus from Gaojiashan locality in the Linxia Basin, Gansu, China are described in this paper, with the first presence of the former in the Linxia Basin. The lower cheek teeth of Hipparion from Gaojiashan bear deep ectoflexids, and after comparisons to other Chinese Hipparion fossils with deep ectoflexids, the mandibles with deep ectoflexids on the lower premolars suggest their attributions to H. forstenae. The presence of a deep ectoflexid on p2 may be interpretable as an intraspecifically variable feature in H. forstenae, and can also be found in H. dermatorhinum. Besides the Hipparion material, at least 8 species of large mammals occur at the Gaojiashan locality, including Adcrocuta eximia, Chilotherium wimani, Shansirhinus ringstroemi, Eucladoceros cf. proboulei, Palaeotragus cf. coelophrys, Giraffidae gen. et sp. indet., and the previously studied Promephitis hootoni and Eostyloceros hezhengensis. Although the sample from Gaojiashan shares five species with the Yangjiashan fauna, the Gaojiashan fossil assemblage may date to the Baodean because of the presence of H. forstenae. The overall taxonomic composition of the Gaojiashan assemblage at least suggests an approximate Baodean age, which would imply survival of C. wimani into the Baodean of the Linxia Basin.     

High-resolution intra- and interbasinal correlation of the Danian-Selandian transition (Early Paleocene): The Bjala section (Bulgaria) and the Selandian GSSP at Zumaia (Spain)

The Danian-Selandian (D-S) boundary has been identified for the first time in the Black Sea coast at Bjala (Bulgaria) based on a new integrated bio-, magneto- and cyclostratigraphic study. Several correlation criteria as established for the basal Selandian GSSP from Zumaia (Basque Basin) are evaluated. Noteworthy, is the almost complete lack of calcareous nannoplankton species Braarudosphaera bigelowi in the Bulgarian sections, a sharp decrease of which was indicated as suitable criteria for defining the D-S boundary as it occurred both at Zumaia and in the classical locations of the North Sea basin. Conversely, the second evolutionary radiation of the calcareous nannofossil genus Fasciculithus together with the occurrence of Fasciculithus tympaniformis that define the NP4/NP5 zonal boundary seem to be reliable criteria to approximate the D-S boundary. In detail, however, the best approach is to integrate biostratigraphic data within a magnetostratigraphic and/or cyclostratigraphic framework. Refinements on the placement of chron C27n at Zumaia and robust bed-by-bed correlation between several Basque sections and Bjala indicates that the D-S boundary is located 30 precession cycles (~ 630 ky) above C27n. In addition to the precession-related marl-limestone couplets and 100-ky eccentricity bundles recognized in the studied sections, expression of the stable 405-ky long eccentricity allows direct tuning to the astronomical solutions. A correlation of the land-based sections with previously tuned data from ODP Site1262 from the Southern Atlantic is challenged. Our choice is consistent with original tuning at Zumaia but shifts one 100-ky cycle older previous tuning from Site 1262 along the interval above C27n. Under the preferred tuning scheme the D-S boundary can be given an age of 61.641 ±0.040 Ma on the La04 orbital solution.     

Crepidolithus cantabriensis nov. sp., a new calcareous nannofossil (Prymnesiophyceae) from the Lower Jurassic of northern Spain

A new calcareous nannofossil, Crepidolithus cantabriensis nov. sp., is described from the Lower Jurassic marls and marly limestones of the Cantabrian Range (Northern Spain). The new species is characterized by medium-sized, normal to narrowly elliptical coccoliths with open central-areas. In the light microscope, under crossed nicols and in distal view, it displays a characteristic bicyclic extinction pattern. Its inner rim edge shows maximum birefringence colours, which vary from white to yellow, whereas the faintly outer rim margin exhibits low birefringence, tending to grey, and they are separated by a dark and sigmoidal isogyre. C. cantabriensis nov. sp. seems closely related to Crepidolithus crassus on the basis of shared morphological characters. Both of them have a distal shield composed of around thirty sub-vertical, dextrally imbricating laths. The central-area width of C. cantabriensis nov. sp. is approximately half of the coccolith width, whilst it is reduced to one or two tiny lenticular slits in C. crassus. Nevertheless, the phyletic relationships between C. cantabriensis nov. sp. and Crepidolithus cavus are not clear, since they show different rim structures, and C. cavus has a delicate bar aligned along the minor axis of its central area. The first specimens of C. cantabriensis nov. sp. have been recognized within the Raricostatum AZ in the Tudanca section. This species seems to be more abundant in the Reinosa area, which occupied a more distal position with respect to the Iberian Massif during the studied time interval. It is hypothesized here that C. cantabriensis nov. sp. could have inhabited the lower-photic zone.     

Paracamelus (Mammalia,Camelidae) remains from the late Messinian of Italy: insights into the last camels of western Europe

In this paper we describe fossil remains of an indeterminate species of the genus Paracamelus (Artiodactyla, Camelidae) from the Messinian post-evaporitic deposits (5.55–5.40 Ma) of Verduno (Piedmont, NW Italy). Camelins dispersed into Eurasia from North America in the late Miocene and almost instantaneously spread in western Europe and Africa. The size and morphology of the fossils found at Verduno are consistent in with those of Paracamelus, the earliest Old World camelin. Up to now, the only fossil camels recovered in the Neogene of Western Europe have been found at Venta del Moro and Librilla in Spain at 6.2 Ma. The remains from Verduno represent the first evidence of a camelin in the Neogene of Italy and they considerably expand the paleobiogeographic range of the Old World early camelins. The presence of a camelid at Verduno reinforces and confirms the importance of the fossiliferous deposits of NW Italy in defining the complex paleobiogeographic patterns of Europe during the Messinian, at the end of the Messinian salinity crisis.     

Sciuridae (Rodentia,Mammalia) from the Late Miocene of Venta del Moro (Cabriel Basin,Spain): biostratigraphical,phylogenetic and palaeoecological inferences

Abstract

Fossil squirrels are relatively abundant in Aragonian and Ramblian assemblages, but very scarce in the Late Turolian and Ruscinian of the Iberian Peninsula. Until now, the locality of Venta del Moro has yielded over 4500 micromammal dental remains, but just nine of them are sciurids. In this assemblage, we have identified the taxa Pliopetaurista pliocaenica, Heteroxerus cf. mariatheresae and Atlantoxerus cf. margaritae. This is the first co-occurrence of the genera Pliopetaurista, Heteroxerus and Atlantoxerus in the same locality, being also one of the oldest records of P. pliocaenica, the oldest record of a form that could be related to A. margaritae, and the youngest record of the genus Heteroxerus. In addition, we hypothesize about the phylogeny of Pliopetaurista in the light of recent discoveries. Finally, the presence of the genera Heteroxerus and Atlantoxerus suggests an open environment and relatively dry environmental conditions, while Pliopetaurista is purportedly a flying squirrel, dweller of forested habitats and wet climates. Based on this, we infer an open environment of grassland type, bordered by forested habitats, agreeing with the conditions supposed for Venta del Moro according to its macromammal and floral assemblages.     

A new coral‐associated decapod assemblage from the Upper Miocene (Messinian) Upper Coralline Limestone of Malta (Central Mediterranean)

Abstract: A rich coral‐associated decapod assemblage is recorded from the ‘Depiru Beds’ of the upper part of the Upper Coralline Limestone (Messinian, Upper Miocene), from the island of Malta. Nineteen species within 17 genera have been discovered, where 14 genera are new for Malta. Four new species are described, namely Micippa annamariae sp. nov., Pilumnus scaber sp. nov., Panopeus muelleri sp. nov. and Herbstia melitense sp. nov. Herbstia melitense sp. nov. constitutes the first record of the genus from the fossil record in the Mediterranean region. This discovery more than doubles the number of known fossil decapod species from Malta. The fossil bivalve Jouannetia (J.) semicaudata Des Moulins, 1830 and the extant decapod Maja goltziana D’Oliveira, 1888, are also recorded for the first time from Malta. Other Neogene coral‐associated decapod assemblages are investigated and correlated with the new assemblage from Malta. The migration of taxa between the Mediterranean region and the Paratethys, particularly during the Lower Badenian (Langhian), is evidenced by the strong affinity of the Maltese decapod assemblage with that of the Middle Miocene Badenian assemblages from Hungary, Poland and Ukraine. Upper Miocene, Messinian assemblages from Spain, Algeria and Morocco are also similar to that from Malta.     

The late Messinian Lago-Mare episode in the Mediterranean Basin: Preliminary report on the occurrence of Paratethyan ostracod fauna from central Crete (Greece)

In the Crete Island, late Messinian Lago-Mare facies are not well known. At present, the occurrence in Crete of the uppermost Messinian post evaporitic deposits is a matter of debate. According to several authors, the well-known late Messinian Lago-Mare facies does not occur in Crete. In this paper the preliminary results obtained from the biostratigraphical analysis of some sections sampled in the Messarà Plain will be shown. Nearby Faneromeni and Ano Akria villages, the Miocene/Pliocene boundary is well exposed. There, gypsum-bearing clay, laminated microcrystalline gypsum and gypsum-rudites characterize the evaporitic deposits of the Messinian stage. In these areas, above the Messinian evaporite, post-evaporitic fine-laminated polychrome clays, with intercalations of sandstones and conglomerates, have been found. In both the Faneromeni and Ano Akria area, the Pliocene grey clays and conglomerates rest unconformably on the uppermost Messinian post-evaporitic deposits. A 20 cm-spaced sampling has been performed in both the sections, for more than 100 samples collected. The results of the micropaleontological analysis performed on the Faneromeni and Ano Akria sections point to the occurrence of ostracod assemblages containing: Loxocauda limata (Schneider in Agalarova et al.), Loxocauda sp., Cytherura pyrama Schneider, Cyprideis anlavauxensis Carbonnel, Cyprideis agrigentina Decima, Amnicythere palimpsesta (Livental), Amnicythere propinqua (Livental), Amnicythere accicularia (Olteanu), Amnicythere costata (Olteanu), Amnicythere multituberculata (Livental), Amnicythere sp. D (Miculan in Bassetti et al.), Amnicythere sp. 2 Gliozzi and Grossi, Amnicythere sp., Euxinocythere (Maeotocythere) praebaquana (Livental in Agalarova et al.), Mediocytherideini indet., Pontoniella pontica (Agalarova), Camptocypria sp. 1 Gliozzi and Grossi, Caspiocypris sp., Zalanyiella venusta (Zalanyi), Tyrrhenocythere sp., Loxoconcha rhombovalis Pokorny, Loxoconcha eichwaldi Livental, Loxoconcha sp. A (Miculan in Bassetti et al.), Loxocorniculina djafarovi (Schneider in Suzin). In the analysed samples, reworked planktonic foraminifers and well-preserved charophyte gyrogonites have been also found. The ostracod assemblages found in the Messarà Plain belong to the Loxocorniculina djafarovi Zone (sensu Carbonnel, 1978), which characterizes the uppermost Messinian deposits of the whole Mediterranean Basin. At that time, the well-known Lago-Mare biofacies was also widespread on the Crete Island. The presence of Paratethyan ostracods in the post-evaporitic Messinian deposits of both Faneromeni and Ano Akria sections suggests that in the Crete Island the latest Messinian sedimentation took place in brackish water palaeoenvironments.     

Microfossil assemblage from the Late of Middle Carboniferous in Western Shaan-Gan-Ning Basin,Northwest China

In this paper, some results of the study on the sporepollen of the Yanghugou Formation in the western Shaan-Gan-Ning (Ordos) Basin of Northwest China axe reported. Plenty of well preserved microfossil, which 136 types belong to 62 microspore genera and one megaspore genus, including 2 new genera , 13 new species, I Acritarch (Chamosphaera pseudozonatus gen. et sp. nov.), from 8 bore-holes and outcrop have been discribed. In generally, the microfiora of the Yanghugou Formation is similar to the Taiyuan Formation (Upper Carboniferous). According to the spore-pollen assemblage of Yanghugou Formation may correlate with those in the Middle Carboniferous of Great Britain, Belgium the north of France, German and North America. Besides, the assemblage are similar to Middle Carboniferous of Nortth Shandong and Hengshanbu of Ningxia in the microfossil assemblage. The geological age of the beds is considered to Late of Middle Carboniferous.