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Abstract: Adequate cover is a critical component of ruffed grouse (Bonasa umbellus) habitat during the brood-rearing period when chick mortality is high. We assessed habitat use by ruffed grouse during the brood-rearing period by comparing characteristics of tree, shrub, and ground layers at ruffed grouse brood and random locations. We captured and radiomarked 29 females with broods in 2 forest settings of the Réserve faunique de Portneuf, Quebec, Canada. We described grouse habitat using ground surveys and forest maps, and we identified the used habitat characteristics using analysis of variance and logistic regression. Females with broods used mixed and regenerated clearcut stands that were 1.5–7 m tall and 11–20 years old. Compared with random locations, grouse locations had higher lateral obstruction (76% vs. 68%), higher small-stem density (29,085 stems/ha vs. 19,340 stems/ha), and were closer to roads and trails. Percentage of coverage by ground vegetation was not higher at grouse locations as often reported in previous studies. Results from this study will help orient ruffed grouse habitat management on Quebec public land and elsewhere in nordic—temperate mixed hardwood—softwood forests to maintain suitable brood habitat after logging operations. Forest management should promote growth of young mixed stands with high horizontal and vertical cover provided by high small-stem density, which offers protection against aerial and terrestrial predation. Edges such as roadsides are also important in brood habitat as they provide food and cover.  相似文献   

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Abstract: Ruffed grouse (Bonasa umbellus) is a popular small game species in northeastern North America. We assessed female ruffed grouse habitat characteristics during winter of 2001-2002 and 2002-2003 in a region dominated by mixed softwood-hardwood forests by comparing used and random locations. We followed 23 radiotagged adult females in 2 forest sites of the Réserve faunique de Portneuf, Québec, Canada, from late November to mid-April. We described grouse habitat using ground surveys and identified selected habitat characteristics using analysis of variance and logistic regression. Females preferred mixed softwood-hardwood stands > 17 m tall and 61-120 years old. Compared with random locations, grouse locations had more well-developed total canopy cover (>4m;75%), canopy (>4m;35%), midstory (1-4 m tall; 35%), and lower-story (<1 m tall; 23%) coniferous cover, and higher coniferous stem density and tree basal area (dbh > 9 cm; 343 stems/ha and 9.0 m2/ha, respectively). Forest management should maintain mature mixed softwood-hardwood stands (50% coniferous), which are adequate winter habitat for ruffed grouse.  相似文献   

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Ruffed grouse (Bonasa umbellus) are a popular game bird and the management indicator species for quaking aspen (Populus tremuloides) in the Black Hills National Forest (BHNF), which requires development of a robust monitoring protocol to evaluate trends in ruffed grouse populations. We used roadside drumming surveys in spring 2007 and 2008 to estimate ruffed grouse occupancy and detection probabilities in the BHNF while simultaneously assessing the influence of sampling and site covariates on these processes. Ruffed grouse occupancy estimates were constant between spring 2007 and 2008 (Ψ = 0.12, SE = 0.03) and were positively influenced by the amount of aspen surrounding the site. Detection probability estimates were constant between spring 2007 and 2008 (p = 0.27, SE = 0.06) and were influenced by survey date in a quadratic form and negatively influenced by wind speed and time of the survey. Collectively, our results demonstrated that ruffed grouse occupancy and detection probabilities in the BHNF were low. Occupancy could be increased by increasing the extent of aspen. To improve monitoring efficiency and maximize probability of detecting ruffed grouse, ruffed grouse monitoring should be conducted during the peak of drumming (mid-May), during favorable weather conditions such as low wind speeds and little precipitation, and during early morning, near sunrise. © 2010 The Wildlife Society.  相似文献   

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Abstract: We surveyed drumming ruffed grouse (Bonasa umbellus) to estimate the probability of detecting an individual, and we used Bayesian model selection to assess the influence of factors that may affect detection probabilities of drumming grouse. We found the average probability of detecting a drumming ruffed grouse during a daily survey was 0.33. The probability of detecting a grouse was most strongly influenced by the temperature change during a survey (temp change = 0.23, 95% probability interval [PI] = 0.13 ≤ ≤ 0.33) and its interaction with temperature at the start of the survey (interaction = 0.01, 95% PI = 1.42 × 10−3 ≤ ≤ 0.03). Although the best model also included a main effect of temperature at the start of surveys, this variable did not strongly correlate with detection probabilities (start temp = −0.03, 95% PI = −0.06 ≤ ≤ 9.80 × 10−5). Model assessment using data collected at other sites indicated that this best model performed adequately (i.e., positive correlation between observed and predicted values) but did not explain much of the variation in detection rates. Our results are useful for understanding the historical drumming index used to assess ruffed grouse populations and for designing auditory surveys for this important game bird.  相似文献   

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Species associated with early successional habitats have experienced dramatic declines in the eastern United States as a result of land use changes and human disruption of natural disturbance regimes. Consequently, active management is required to create early successional habitat and promote plant and animal communities that depend on periodic forest disturbance. Ruffed grouse (Bonasa umbellus) depend on recently disturbed forest habitat, and have experienced dramatic declines over the last half-century. Although ruffed grouse are extensively studied, little effort has been made to link population dynamics with habitat management at landscape scales. We used stochastic, spatially explicit population models that combined landscape conditions derived from a Geographic Information System with demographic data, and applied the model to a declining ruffed grouse population in Rhode Island, USA. We identified vital rates that influence ruffed grouse population dynamics using baseline models constructed with current demographic rates and landscape conditions, and assessed the effect of landscape-scale forest management alternatives on population persistence by running multiple management simulations. Baseline models typically predicted population decline, and we concluded that vital rates (survival and recruitment) had a greater influence on population persistence than did dispersal capability, carrying capacity, or initial population size. Management simulations predicted greater population persistence under a scenario where high-quality habitat was provided in fewer large blocks as opposed to many small blocks, and the rate at which we allowed ruffed grouse to colonize newly created habitat had a substantial impact on management success. Populations of ruffed grouse in the eastern United States are likely to continue to decline given current disturbance regimes, and our work provides a link between ruffed grouse demography and landscape-scale habitat conditions to support management decisions. © 2011 The Wildlife Society.  相似文献   

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1999至2002年,在法国东南部的阿尔卑斯山,通过无线电追踪方法研究了花尾榛鸡的性比问题。首先,我们通过直接观测无线电标记雄性的配对比例,在消除可能的捕捉误差后,发现繁殖种群性比为1.26(即未配对的雄性占21%)。然后,我们估计了标记个体的存活率和繁殖成功率。在假设窝扩散时幼体的性比平衡的前提下,通过两性和两个年龄组(幼体,〈10月龄的个体;成体〉10月龄的个体)的确定性统计模型,得到的理论性比为1.22(即未配对的雄性占18%),这一结果与实际观测的性比十分接近。我们认为,雌性死亡率高导致的两性存活率差异可以解释雄性性比偏高现象。我们进而讨论了花尾榛鸡整个分布区内居间强度的非对称性比的适应意义,以及在衰退的花尾榛鸡种群中雌性数量极端不足的后果[动物学报52(4):655—662,2006]。  相似文献   

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Cyclic dynamics of bird and mammal populations are commonly reported in northern latitudes throughout the world, and recent European observations on rodents and grouse suggest that cycle periods decline towards southern latitudes. To investigate latitudinal patterns of cyclic dynamics in North America, we assembled 27 long‐term data sets collected between 1939 and 2001 for three grouse species. By fitting the data with autoregressive models to measure direct and delayed density dependence, we show that, in contrast to European studies, North American grouse exhibit period increases from north to south, with cycles collapsing via period lengthening. This occurs because delayed density dependence decreases in southern latitudes, whereas direct density dependence increases. These results show that cyclic dynamics can dissipate by period lengthening as well as the period shortening postulated for European grouse and rodents.  相似文献   

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The amount of dispersal that occurs among populations can be limited by landscape heterogeneity, which is often due to both natural processes and anthropogenic activity leading to habitat loss or fragmentation. Understanding how populations are structured and mapping existing dispersal corridors among populations is imperative to both determining contemporary forces mediating population connectivity, and informing proper management of species with fragmented populations. Furthermore, the contemporary processes mediating gene flow across heterogeneous landscapes on a large scale are understudied, particularly with respect to widespread species. This study focuses on a widespread game bird, the Ruffed Grouse (Bonasa umbellus), for which we analyzed samples from the western extent of the range. Using three types of genetic markers, we uncovered multiple factors acting in concert that are responsible for mediating contemporary population connectivity in this species. Multiple genetically distinct groups were detected; microsatellite markers revealed six groups, and a mitochondrial marker revealed four. Many populations of Ruffed Grouse are genetically isolated, likely by macrogeographic barriers. Furthermore, the addition of landscape genetic methods not only corroborated genetic structure results, but also uncovered compelling evidence that dispersal resistance created by areas of unsuitable habitat is the most important factor mediating population connectivity among the sampled populations. This research has important implications for both our study species and other inhabitants of the early successional forest habitat preferred by Ruffed Grouse. Moreover, it adds to a growing body of evidence that isolation by resistance is more prevalent in shaping population structure of widespread species than previously thought.  相似文献   

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  总被引:4,自引:0,他引:4  
Ecologists often assume that dispersing individuals experienceincreased predation risk owing to increased exposure to predatorswhile moving. To test the hypothesis that predation risk isa function of movement distance or rate of movement, we usedradio-telemetry data collected from 193 ruffed grouse (Bonasaumbellus) during 1996–1999 in southeastern Ohio. Cox'sproportional hazards model was used to examine whether the riskof predation was affected by the rate of movement and site familiarity.We found evidence indicating that increased movement rates mayincrease the risk of predation for adult birds but not juveniles.We also found juvenile and adult birds inhabiting unfamiliarspace were consistently at a much higher risk of predation (threeto 7.5 times greater) than those in familiar space. Our resultsindicate that although movement itself may have some effecton the risk of being preyed upon, moving through unfamiliarspace has a much greater effect on risk for ruffed grouse. Thissupports the hypothesis that increased predation risk may bean important cost of dispersal for birds.  相似文献   

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Abstract: Harvest of upland game birds in concert with sampling of age ratios from wings can yield important biological information about populations. Although estimates of productivity are commonly produced, they are often not accompanied by a measure of variance. Thus, we developed standard error estimates for sample productivity ratios, compared 4 methods for creating confidence intervals for population productivity ratios, and developed a test and the corresponding sample size requirements for comparing 2 population productivity ratios. We applied these techniques to greater sage-grouse (Centrocercus urophasianus) wing-data collected in Oregon, USA (1993–2005). Computer simulations indicated that backtransforming the Wilson's score interval on the proportion of immatures in the sample results in the most reliable confidence intervals among the methods considered. We recommend to managers measuring conservation action outcomes with productivity ratios to consider the appropriate sample sizes for the spatial and temporal scale of their monitoring programs.  相似文献   

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1. To examine the effects of forest harvest practices on headwater stream macroinvertebrates, we compiled a 167 site database with macroinvertebrate, fish, physical habitat and catchment land cover data from the three forested ecoregions in western Oregon. For our analysis, headwater streams were defined by catchment areas <10 km2 and perennial water during summer low flows. Almost all sites in the database were selected using a randomised survey design, constituting a representative sample of headwater streams in these ecoregions. 2. Macroinvertebrate taxonomic and functional feeding group composition were very similar among the three ecoregions in the study area (Coast Range, Cascades and Klamath Mountains). On average, 55% of the individuals at each site were in the orders Ephemeroptera, Plecoptera or Trichoptera. Dipteran taxa (mostly chironomids) accounted for another 34%. At almost all sites, non‐insects made up <10% of the macroinvertebrate assemblage. Almost half (49%) of the assemblages were collectors; remaining individuals were about evenly divided among scrapers, shredders and predators. 3. There were 189 different macroinvertebrate taxa at the 167 sites with richness at individual sites ranging from 7 to 71 taxa. Ordination by non‐metric multidimensional scaling revealed a strong association between % Ephemeroptera, especially Baetis, and site scores along the first axis. This axis was also strongly related to % coarse substratum and fast water habitat. The second axis was strongly related to % intolerant individuals, site slope and altitude. No strong relationships were evident between any ordination axis and either logging activity, presence/absence of fish, catchment size or ecoregion. 4. Based on macroinvertebrate index of biotic integrity (IBI) scores, 62% of the sites had no impairment, 31% of the sites had slight impairment and only 6% of the sites had moderate or severe impairment. IBI scores were not strongly related to forest harvest history. All four severely impaired sites and five of the seven sites with moderate impairment were lower altitude, shallower slope stream reaches located in the Coast Range with evidence of agricultural activity in their catchment or riparian zone. % sand + fine substratum was the environmental variable most strongly related to macroinvertebrate IBI.  相似文献   

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  总被引:1,自引:1,他引:0  
Active-acoustic surveys were used to determine the distribution of dusky dolphins and potential prey in two different New Zealand locations. During seven survey days off Kaikoura Canyon, dusky dolphins were found within the DeepScattering Layer (DSL) at 2000 when it rose to within 125 m of the surface. As the DSL rose to 30 m at 0100, the observed depth of dolphins decreased, presumably as the dolphins followed the vertical migration of their prey. Acoustically identified subgroups of coordinated animals ranged from one to five dolphins. Time, depth of layer, and layer variance contributed significantly to predicting foraging dusky dolphin subgroup size. In the much shallower and more enclosed Admiralty Bay, dolphins noted at the surface as foraging were always detected with the sonar, but were never observed in coordinated subgroups during the brief (two-day) study there. In Admiralty Bay dolphin abundance was correlated with mean volume scattering from potential prey in the water column; and when volume scattering, an index of prey density, was low, dolphins were rarely present. Ecological differences between the deep waters of Kaikoura Canyon and the shallow nearshore waters of Admiralty Bay may result in differences in how, when, and in what social groupings dusky dolphins forage.  相似文献   

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Exfoliated skin was collected from bowriding dolphins with the use of a sterilized nylon scrub pad affixed to a wooden dowel. Initial tests of the effectiveness of the technique and dolphin behavioral responses were conducted on dusky dolphins off Kaikoura, New Zealand. During 14 sampling days, 128 contacts using this procedure were made with bowriding dolphins, of which 114 showed behavioral response. Responses during sampling were mild, with 11% of contacted individuals showing no visible response, and 66% of individuals which could be monitored for 30 sec after contact returning to bowride within 30 sec. Mean return time was 10 sec postcontact and did not vary significantly for groups of different sizes. Behavioral controls suggest that a proportion of responses could be explained by typical dolphin behavior in the presence of boats. Seventy-eight percent of contacts resulted in successful collection of tissue samples. Sample time was three minutes on average. Size of group and behavioral state did not appear to influence sample time. Preliminary genetic analyses revealed that tissue collected by this technique was suitable for amplification and sequencing of mitochondrial DNA (mtDNA) via PCR. Comparisons of mtDNA control region sequences with those from known L. obscurus and other delphinid samples verified that this technique was robust against contamination from elements in sea water.  相似文献   

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Deviations from equal sex ratios in mammals can reveal insights into sex-specific growth, survival, movements, and behavior. We assessed blue whale (Balaenoptera musculus) sex ratios based on 21,542 fetal and 311,901 whaling records, finding that males were slightly but significantly more common than females (51.3% fetal, 52.1% postnatal). Antarctic catches shifted from 52.4% male before 1951 to 48.0% male thereafter, even though larger females were preferentially targeted by whalers and should have declined. The southernmost land stations caught more males than those in southern Africa, and at land stations, sex ratios shifted subtly over the course of a year. Pelagic catches demonstrated spatial structure in sex ratios, including more males being caught in the Ross Sea. In utero, the smallest females were often misidentified as males, and there was some evidence for higher prenatal male mortality. Once born, medium-sized blue whales within each region were more often male, while the longest were nearly all female; explained entirely by females growing faster and reaching longer sizes. Overall, though, sex ratios are remarkably close to equality across time, space, and length; with any deviations best explained by faster female growth and size-selective whaling.  相似文献   

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