五福花科花部形态的变异式样及其系统学意义

由于四福花Ｔｅｔｒａｄｏｘａ顶花、侧花均为典型的４数花,并被认为是科内最原始的属,因而长期以来关于五福花Ａｄｏｘａ“总苞一萼”还是“萼一花冠”的争议得到了彻底解决。根据本研究,五福花顶花和侧花的变异痃较五福花科另外３个分类群要高得多。五福花顶花和侧花的变异率分别为２０％和３６１％。通过后位雄蕊的退化和两则相邻花瓣的融合,雄蕊和花瓣从６数至３数的减少系列仅存在于五福花中。本文假设五福花科顶花和侧花的     

五福花科花部维管结构的比较研究

首次报道了华福花属华福花 Sinadoxa corydalifolia和东方五福花 Adoxa orientalis花部维管结构,并对五福花科3属4种植物的花部维管系统进行了比较研究。3属4种的花部维管系统表现出不同的对称性和不同的复化、减化以及不同的融合程度。基于花部维管系统的证据,本工作支持四福花属四福花 Tetradoxa omeiensis为科内原始类群代表的看法。     

五福花科花部维管结构的比较研究

首次报道了华福花属华福花Ｓｉｎａｄｏｘａｃｏｒｙｄａｌｉｆｏｌｉａ和东方五福花Ａｄｏｘａｏｒｉｅｎｔａｌｉｓ花部维管结构,并对五福花科３属４种植物的花部维管系统进行了比较研究。３属４种的花部维管系统表现出不同的对称性和不同的复化、减化以及不同的融合程度。基于花部维管系统的证据,本工作支持四福花属四福花Ｔｅｔｒａｄｏｘａｏｍｅｉｅｎｓｉｓ为科内原始类群代表的看法。     

花锚属一新变型——玉花卵萼花锚

报道了龙胆科卵萼花锚的一个新变型——玉花卵萼花锚。新变型区别于原变型的唯一特征是其花冠为白色,部分花距为黄色。     

四福花染色体核型的分析

四福花[Tetradoxa ometensis (Hara)C.Y.Wu]体细胞具有36个染色体。其核型组成为2n=36=6m+14sm+4st+12t,即具有3对中部着丝点染色体,7对亚中部着丝点染色体,2对亚端部着丝点染色体和6对端部着丝点染色体。 四福花染色体核型分析表明,与传统对五福花科植物染色体具9基数的认识不同,其基数应为X=18。与Noguchi所发现的具18基数的三倍体五福花的核型相比较,二者在核型组成及染色体结构上都有明显差异。     

狭义五福花科的分子系统学和物种分化

狭义五福花科(Adoxaceae.s.s)仅含3属4种,但该科的物种分化、系统发育和分类一直存在争议。本文通过测定东方五福花和血满草的ITS(核糖体DNA内转录间隔区)序列,构建了包括狭义五福花科(4种)、广义忍冬科接骨木属、荚属以及其余4属植物在内的系统发育树。研究结果不支持狭义五福花科内根据形态学证据做出的系统发育假设四福花不是该科中最早分化出来的种;该物种与五福花属的两个物种形成一个单系群,与另一分支华福花属相对应。该科中3个物种,四福花、五福花和华福花之间的分化主要是在二倍体水平上的异域分化,而东方五福花则是通过多倍化形成的。粗略的时间估算表明这些物种之间的分化较晚,可能在第三纪末至第四纪早中期,与青池高原近期强烈隆升以及冰期气候反复变化形成的环境变迁密切相关。     

浅谈花萼及其演化

由若干片萼片组成花的花萼,包被在花的最外层,在花蕾期有保护作用。萼片基本上是小的变态叶或变态苞片。普通萼片多为绿色的叶状体,在结构上类似叶,有大量的绿色薄壁细胞,但无栅栏组织和海绵组织的分化,有一定的光合作用能力。花萼通常一轮,也有两轮的,外轮的花萼叫副萼。如锦葵、棉、草莓等。萼片各自分离的,称离萼,如大多数较原始的植物木兰、莲等。也有彼此连合一起构成为合萼的,大多数植物花的萼片都合生。萼片大小相同或不同,分别称为整齐萼或不整齐萼。萼片比花冠先脱落的称早落萼,如罂粟;有的与花冠一起脱落,称落萼,如油菜、桃;也有的花萼常留在花柄上,随同果实一起发育,称宿萼,如茄、柿、番茄、辣椒等;有的花萼一边引伸成短小的管状突起,称为矩,如凤仙花、旱金莲等植物的花萼;有的花的萼片大而有各种颜色,类似花冠,具有吸引昆虫传粉的作用,如铁线莲和绣球花的花萼。     

中国囊萼花属一新记录种——茎花囊萼花

首次报道了囊萼花属(Cyrtandromoea Zoll.)的1个中国新记录种——茎花囊萼花(Cyrtandromoea grandis Ridl.),该种原记载产于泰国、马来半岛和苏门答腊岛,2019年在中国云南省勐海县布朗山地区发现有分布。茎花囊萼花与囊萼花(Cyrtandromoea grandiflora C.B.Clarke)最为相似,但前者聚伞花序生于老茎上,每一花序3～18朵花,二者容易区别。     

华福花的解剖学特征及其同五福花的比较

本文记载了华福花的地理分布,生长和繁衍的物理条件,详细描述了营养器官的解剖学特征和花粉形态。从四个方面比较了两种植物的共同点和不同点,强调了华福花和五福花相比处于更高的阶段,并在发展和分化中。     

肋柱花属的系统研究

本文对肋柱花属的属下分类、系统发育和地理分布等方面进行了深入研究。文用分支系统学的方法和原理,用计算机PAUP程序处理,获得了几个最简约的支序图。 肋柱花属属于龙胆亚族辐状花冠群,在这群中,论亲缘关系它与辐花属最近,与獐牙菜属次之,而与黄秦艽属关系较远。 獐牙菜属在进化程度上较肋柱花属低,因此它被选为肋柱花属的外类群。 经过支序分析,肋柱花属的18个种根据Hennig的“共近裔性原则”可组合为三个组,其中肉质根茎组为较原始的组,肋柱花组为中级进化水平的组,合萼组是进化程度最高的组。 肋柱花属是北温带分布型的属,分布于亚洲、欧洲及北美洲,直达北极。从种的地理分布型分析,表明秦岭一横断山区是本属的起源与分化中心。 随文报道了一个新组、一个新种和一个新变种。 研究了全部种类的命名模式。     

Molecular Phylogeny and Species Speciation of Adoxaceae. s. s

It remains unclear about the speciation and phylogeny of Adoxaceae s. s ., a small family with 3 genera and 4 species . In this paper , ITS ( nuclear DNA internal transcribed spacer) regions of Adoxa orientalis and Sambucus adnata were firstly sequenced . Phylogenetic trees were constructed for all species of Adoxaceae (four species ) , Sambucus, Viburnum and four genera of Caprifoliaceae . The divergences among four species of this family were further calculated based on the calibration of the fossil records of the Caprifoliaceae and the general evolutionary rate of herbs for ITS . The phylogenetic analyses did not support the previous assumptions on the phylogeny and species divergence of Adoxaceae s. s . based on the morphological evidence : Tetradoxa is not the firstly diverged and it clustered with two species of Adoxa as a monophylogenetic group , paralleling to the other lineage comprising of monotypic Sinadoxa . The allopatric speciation at the diploid level might have contributed to the differentiation among Sinadoxa corydalifolia, Tetradoxa omeiensis and Adoxa moschatellina and the polyploidy to the origin of A. orientalis . The crude timing based on ITS sequence differentiation suggested a recent divergence among all four species probably between the late Miocene and the Tertiary and this speciation process might be closely correlated with habitat fragmentation and change due to the extensive uplifts of the Qinghai-Tibetan Plateau and climatic oscillation during the glacial and interglacial ages occurred at this stage .     

Wuyunanthus gen. nov., a flower of Celastraceae from the Palaeocene of north-east China

A flower assigned to Wuyunanthus hexapetalus gen. & sp. nov. has been collected from the Wuyun Formation of Palaeocene age, Jiayin County, Heilongjiang Province, north-east China. The flower is bisexual, actinomorphic, and 6-merous. There are six free valvate petals in a whorl inserted on the margin of a hexagonal disk, and six free stamens alternate with petals within it. The pistil consists of fused carpels. Ovary with an apical style is superior. The unique combination of characters of the fossil, such as stamens inside the thick and flattened disk surrounding the ovary and equal in number to the petals, is similar to that of Celastraceae. The fossil is assigned to Celastraceae but distinguished from other extant genera of the family by being 6-merous rather than 4- or 5-merous. It might represent an extinct and early type of Celastraceae. The presence of a disk might provide evidence for insect pollination in the early stages of Celastraceae evolution.     

Manipulus Rubiacearum IV. Kerianthera (Rubiaceae), a new genus from Amazonian Brazil

A new genus,Kerianthera, and its single new species,K. preclara, are presented.Kerianthera is a member of tribe Condamineeae and is easily distinguished from related genera by its 4-merous calyx, 7 or 8-merous corolla, stamens separating from the corolla tube at its apex, anthers with approximately 300 locelli, septicidal capsules, and seeds irregularly biwinged. It is most similar phenetically to the genusPinckneya with which it shares foliar calyx lobes on some flowers, corolla lobes densely pubescent on the inner surface, and winged seeds.     

A New Species and a Newly Recorded Species of Adoxa L. from Xizang

A new species and a new record for China of the genus Adoxa L.are reported from Xizang, China. They are Adoxa xizangensis G.Yao and Adoxa inodora (Falc.ex C.B.Clarke) Nepomn.     

Archamamelis,hamamelidalean flowers from the Upper Cretaceous of Sweden

Lignite fossil flowers (including pollen) and isolated stamens of probable hamamelidalean (possible hamamelidaceous) affinities from the upper Cretaceous (Late Santonian or Early Campanian) of Sweden are described. The flowers are 6–7-merous with probably a double perianth, one whorl of stamens and (2-?)3 carpels. The stamens are disporangiate; each theca opens by a valve towards the centre of the flower. Pollen is tricolpate, tectate-columellate and reticulate; the endexine is lamellated in the apertural region. The gynoecium has free styles and a syncarpous ovary. In the one flower that was serially sectioned the ovary is either non-functional or development of the few (2?) ovules is retarded.     

Taxonomic studies in the Miconieae (Melastomataceae). IX. Calycogonium formonense, a new species from the Massif de la Hotte, Haiti

Calycogonium formonense, a new species, is here described from the floristically diverse Massif de la Hotte of southern Haiti. It is compared to C. hispidulum, to which it is likely related. Although the genus Calycogonium is not monophyletic, C. formonense and C. hispidulum are probably related to other species, e.g., C. calycopteris, C. heterophyllum, and C. reticulatum, that exhibit reduced inflorescences and 4-merous flowers having hypanthia with four conspicuous lobes separated by longitudinal grooves and external calyx lobes that are flattened parallel to the floral radii. These distinctive species may form a clade. Calycogonium formonense is distinguished from C. hispidulum by its smaller leaves with the tertiary veins not raised on the abaxial surface, with entire margins (i.e., margin without elongate multicellular hairs), and usually solitary flowers.