松辽盆地白垩纪介形类生物地层学特征

本文详细论述了松辽盆地白垩系介形类属、种数量及壳饰变化在纵向上的分布规律和介形类组合特征的生物地层学意义。认为介形类属的分布是岩石地层组划分的重要依据,种的分布控制了岩石地层段的划分,并可将萨尔图油层、葡萄花油层和高台子油层进一步细分。同时指出松辽盆地介形类的盛衰与湖盆的发育关系密切,介形类的演化周期与湖盆的发育、收缩和沉积旋回相一致,在湖盆最发育的时期,也是介形类繁盛的高峰,湖盆发育的初期和末期,往往是介形类的发生和衰退期。     

东营凹陷沙河街组介形虫化石保存特征研究及应用

本文对山东东营凹陷沙河街组介形虫化石壳体的保存特征进行了研究。研究结果表明：沙河街组介形虫化石壳体颜色可分为浅色、中色和深色三种类型;按有无充填物,可分为未充填壳和充填壳两种,其中充填壳充填物的成分有方解石、白云石、黄铁矿和泥质四种;按完整程度有双瓣壳、单瓣壳和碎壳。文中对这些保存特征形成的原因进行了讨论,最后以一口井的研究结果为例,探讨了介形虫保存特征在古湖泊学及石油地质方面的应用。     

松辽盆地晚白垩世介形类古生态学

通过化石组合和岩性指标分析松辽盆地上白垩统自下而上青山口组—姚家组—嫩江组一段介形类的古生态学。青山口组盐度具有偏咸性、微咸性到半咸性的变化,32个介形类种以栖居于温暖湿润气候带的浅湖微咸水类型占绝对优势,水体清澈安静,有机质丰富,底质为泥等细粒沉积物(个别的为粉砂沉积物),宜于介形类动物群生长且有利于壳体完好保存;其次为生活于半咸水的深湖—半深湖类型,底质为泥。姚家组—嫩江组一段见23个介形类种,栖居于温暖半湿润气候带,以半咸水的深湖、半深湖—浅湖区的类型为主,多泥底,偶见粉砂质底;其次为微咸水的浅湖—三角洲前缘型,泥或粉砂质底;仅3个种生活于温暖湿润的粉砂质底淡水浅湖区。温暖湿润的淡水三角洲分流平原带则难以产出介形类化石。     

内蒙古海拉尔盆地南部地区晚白垩世青元岗组介形类

内蒙古海拉尔盆地南部探井青元岗组红色碎屑岩中发现较丰富的介形类化石,即下段的Ahanicypris obesa-Talicypridea triangulata组合和上段的Chinocypridea augusta-Talicypridea qingyuangangensis组合,这些介形类化石地方性色彩强烈,表现为以具冠状壳喙类型的Ahanicypris,Talicypridea,Chinocypridea和网纹发育的Harbinia等属繁盛为特征,反映了中国东北地区晚白垩世晚期介形类动物群面貌。根据介形类化石组合特征及其分布规律,可以与松辽盆地晚白垩世四方台组的介形类动物群对比,时代为晚白垩世Maastrichtian期。     

同种介形类壳体的多态性兼及化石介形类的物种鉴定

介形类个体发育过程中壳体形状及各种构造的多态性,以及各种外界环境因素对分形类壳体各种分类特征形态变化的影响是十分明显的。该文较系统地阐述了在介形类个体发育中壳体形状、大小、肌痕、边缘毛细管带、边庭、铰合构造及纹饰的变化,并介绍了成年期雌、雄性壳体及左、右壳瓣在形态上的差异;列举了环境因素(主要是水体温度、含盐度、深度)对介形类壳体各分类特征形态变化的影响。基于以上事实,提出了古生物工作者在介形类化石鉴定中应注意的一些问题并讨论了某些描述术语的具体标准。     

介形类系统分类与起源演化的形态、分子和化石证据

介形类(Ostracoda)因其丰富的化石记录和广布的海陆现生代表类群,而被认为是进化生物学中研究生物多样性产生机制和演变历程的颇具潜力的重要模式生物。介形类在甲壳亚门中的谱系发生位置、起源及其内部各类群间的系统关系还存在诸多争议。基于其体制构造的形态学特征,介形类被归入甲壳亚门下的颚足纲(Maxillopoda),但来自18S rDNA序列数据分析却显示Maxillopoda不是单系群。基于化石记录和壳体形态特征,高肌虫(Bradoriida)长期以来被认为是介形类的一个祖先类群,但保存有软躯体的早寒武世化石的研究表明,Bradoriida不是介形类甚至可能也不属于甲壳类。不同的研究者所强调的壳体和肢体形态特征各异,导致介形类最大的现生类群速足目(Podocopida)的四个超科之间的关系也存在诸多推测。壳体和肢体特征在系统演化意义上的不兼容,需要分子生物学等证据的介入。分子、形态和化石证据的积累及各种信息整合是系统演化研究的必然趋势。     

介形类的形态学及其环境、功能和分类学意义

化石介形类分类依据--壳体形态,包括壳体的大小、形状、装饰和诸如毛孔、叠覆、似犁状构造、喙、铰合构造等构造特征都与介形类的栖息环境,包括地理、气候(季节、纬度等)、水体的深度、化学性质和稳定程度,食物的富集程度和群落的组成及个体数量以及性别息息相关.因此,正确理解介形类的形态学特征及其功能学,不仅能揭示介形类的生活习性,推测化石介形类的生活环境,从而为古环境和古群落生态重建提供依据,而且可以提高化石介形类个体发育、系统分类和演化研究结果的可靠性.     

松辽盆地上白垩统嫩江组一段介形类生物地层研究

在对杏104井,杏1—丁3—检323井和塔3井等3口优选井的介形类化石鉴定的基础上,通过对3口优选井涉及研究层位的87块介形类化石样品进行统计分析和编制介形类主要种的地层分布延限图、开展介形类生物地层学研究,建立松辽盆地晚白垩世嫩江组一段介形类化石带7个,包括新建介形类化石带1个,修订介形类化石带1个,并详细描述这7个介形类化石带。     

渤海海域渤中34-A地区古近系介形类组合特征及沉积环境

在渤海海域渤中34-A地区东营组二段下部,发现了以渤海地方属种具棘绍曾介Shaozengia echinata、腹瘤绍曾介S.ventrinodosa、鼓包绍曾介S.gibbosa等为主的大量介形类化石,指示沉积水体环境为淡水-微咸水的滨湖-浅湖;在沙河街组一段上部发现了指示沙河街组一段上亚段存在的李家广北介Guangbeinia lijiaensis亚组合,亚组合发育有椭圆西营介Xiyingia elliptica、高西营介X.alta等大个体、高梯形的介形类,反映当时浅湖-半深湖的沉积环境。通过东营组二段下部与沙河街组一段上部两个介形类发育段的追踪,重新厘定了研究区东营组的地层界面,并确定了沙河街组一段的连续存在。     

川东中侏罗统新田沟组介形类化石壳体方解石化特征及元素成分初步分析

首次利用扫描电子显微镜观察川东新田沟组介形类化石的保存情况,并利用X射线能谱分析部分化石的元素含量及分布。研究发现新田沟组介形类壳体绝大部分方解石化,不同层位介形类壳质方解石化有所不同,且方解石化特点显示其可能与有机质作用有关。通过与前人研究结果对比发现,新田沟组壳体保存处在介形虫壳体埋藏作用的第二阶段和第三阶段。     

甘肃玉门下沟地区早白垩世下沟组介形类

甘肃玉门下沟地区下沟组介形类化石十分丰富,该地区下沟组介形类化石共计9属4亚属,21种,本文描述了其中4新种,即Cypridea(Cyamocypris)xiagouensissp.nov.,Cypridea(Cypridea)subunicostatasp.nov.,Stenestroemiasubpeculiarissp.nov.和Stenestroemiaxiagouensissp.nov.。该介形类化石组合尤以Cypridea最为繁盛,通过分析介形类属种的形态特征和化石组合特征并结合岩性特征,推断下沟组的地质时代为早白垩世巴列姆期;并认为下沟组为水动力较弱的淡水-微咸水河湖相沉积。     

山东蒙阴盆地常路组二段(古新世——早始新世)介形类化石

本文记述了2011年采自山东蒙阴盆地常路组二段的介形类化石共计6属10种4未定种,可建立一个Metacypris-Paracandona-Cyclocypris-Candona-Candoniella组合。根据介形类化石与国内已知化石组合对比研究,常路组二段的时代大致为古新世——早始新世。     

Brood care in a Silurian ostracod

An exceptionally preserved new ostracod crustacean from the Silurian of Herefordshire, England, preserves eggs and possible juveniles within its carapace, providing an unequivocal and unique view of parental brood care in the invertebrate fossil record. The female fossil is assigned to a new family and superfamily of myodocopids based on its soft-part anatomy. It demonstrates a remarkably conserved egg-brooding reproductive strategy within these ostracods over 425 Myr. The soft-tissue anatomy urges extreme caution in classifying 'straight-hinged' Palaeozoic ostracods based on the carapace alone and fundamentally questions the nature of the shell-based Palaeozoic ostracod record.     

滴水湖现生介形类与水环境因子间的典范对应关系

滴水湖作为人工新建湖泊,其生态系统形成过程中的跟踪数据对重建其他湖泊生态系统具有重要的参考意义。2013年7月和2014年8月对滴水湖现生介形类分布状况及水环境因子分别进行了调查,结果发现,滴水湖湖水盐度范围介于1.4‰—2.0‰之间,属于微咸水湖。12个采样点共鉴定出现生介形类13种,包括非海相类克氏丽星介Cypria kraepelini(G.W.Müller)和无偶斗星介Cypridopsis vidua(O.F.Müller),海相类中华洁面介Albileberis sinensis Hou、向岛薄丽星介Dolerocypria mukaishimensis Okubo、腹结细花介Leptocythere ventriclivosa Chen、东台新单角介Neomonoceratina dongtaiensis Yang et Chen、长新中华花介Neosinocythere elongata(Hu)、闪光似异口介Paradoxostoma nitida Ho、典型中华美花介Sinocytheridea impressa(Brady)、长中华海花介Sinopontocythere elongata(Gou)、古屋刺花介Spinileberis furuyaensis Ishizaki et Kato、美丽刺面介Spinileberis pulchra Chen和丰满陈氏介Tanella opima Chen。应用典范对应分析(CCA)方法对滴水湖12个样点的8种介形类和9个环境因子进行了相关性研究,结果显示:介形类和环境因子间具有明显的相关性;9个环境因子中叶绿素、总磷和电导率对介形类的分布影响最大,总氮和湖水水温对介形类也有一定影响;从物种分布状况看,相对于低分布频度物种,高分布频度(Y>0.01)物种对环境的耐受性更大,其中Cypria kraepelini和Neomonoceratina dongtaiensis对环境因子的敏感性明显高于其它物种。结合历史资料分析发现,不仅自然环境因子(如电导率、水温等),而且人为干扰因素(如总磷、总氮)都对介形类分布的改变起到重要影响。     

Complex controls on ostracod palaeoecology in a shallow coastal brackish‐water lake: implications for palaeosalinity reconstruction

1. The low‐Mg calcite shells of Ostracoda (Crustacea) are often well preserved in the sediments of alkaline lakes. In coastal waters that have undergone large temporal changes in salinity, ostracod assemblages preserved in the sediment record have been used to reconstruct palaeosalinity, often assuming that salinity is the only significant control on the faunas. 2. We evaluate the performance of ostracods as palaeosalinity indicators in Hickling Broad, a shallow brackish coastal lake in Norfolk, U.K., by comparing fossil ostracod assemblages covering two centuries with geochemical inferences and instrumental records of past salinity and water composition along with other palaeolimnological indicators. 3. Despite large changes in the salinity of the lake and the supposed salinity sensitivity of ostracods, the fossil ostracod assemblages do not clearly reflect the salinity trends inferred from the other independent data. Rather, a complex series of changes has occurred in the lake over the past 200 years and factors other than salinity, including eutrophication, toxicity and associated complex alterations in habitat availability have probably influenced ostracod assemblages. In contrast, there is a good broad agreement between inferred or measured salinity and the trace‐element chemistry of ostracod shells. 4. We conclude that ostracod faunas may not always provide unambiguous palaeosalinity records and should therefore not be used to reconstruct salinity changes except as part of a multi‐proxy investigation that includes other palaeoecological and/or geochemical indicators.     

松辽盆地三肇地区泉头组三段上部介形类化石的指相意义

松辽盆地三肇地区白垩系下统泉头组三段上部(杨三组—杨一组)以大套的紫红、红色砂、泥岩沉积为主,在紫红色泥岩中含有较丰富、完整的介形类及其它门类化石。以往认为,这套以红色为基调的地层属河流相沉积。对其中介形类等化石的古生态、埋藏特征及丰度变化的研究表明,该地层的沉积环境应为浅水湖泊环境,而非河流等水上环境。湖相沉积自杨三组就已存在,向上分布范围不断扩大,反映了湖泊水域面积呈现逐渐扩大的趋势。     

On the preservation of carapaces of some limnic ostracods: An exercise in actuopalaeontology

The way in which the ostracod carapace decomposes through biological and physico-chemical processes is illustrated with Recent and Neogene non-marine ostracods (candonines, herpetocyprines and cytherids) from lacustrine and groundwater habitats. The degree of carapace preservation for Recent Candoninae, in the Mondsee (Austria) is shown to be dependent on the sedimentary microenvironment. The carapaces of herpetocyprines, because of their special morphological structure, decay very easily. This could explain the rare occurrrence of this ostracod group in Palaeogene deposits in Europe.     

Land use and ostracod community structure

Ostracods are important members of the benthos and littoral communities of lake ecosystems. Ostracods respond to hydrochemistry (water chemistry) which is influenced by climatic factors such as water balance, temperature, and chemicals in rainfall runoff from the land. Thus, at local scales, environmental preferences of ostracods and characteristics of lakes are used to infer changes in climate, hydrology, and erosion of lake catchments. This study addresses potential drivers of ostracod community structure and biodiversity at multiple spatial scales using NMS, CART^?, and multiple regression models. We identified 23 ostracod species from 12 lake sites. Lake area, maximum depth, spring conductivity, chlorophyll a, pH, dissolved oxygen, sedimentary carbonate, and organic matter all influence ostracod community structure based on our NMS. Based on regression analysis, lake depth, chlorophyll a, and total dissolved solids best explained ostracod richness and abundance. Land uses are also important community structuring elements that varied with scale; locally and regionally agriculture, wetlands, and grasslands were important. Nationally, using regression tree analysis of lakes sites in the North American Non-marine ostracod database (NANODe), row-crop agriculture was the most important predictor of biodiversity. Low agriculture corresponded to low species richness but greater landscape heterogeneity produced sites of high ostracod richness.     

The first non‐marine ostracod fauna from the Lower Barremian dysodiles of Lebanon

Dysodiles are finely laminated sedimentary rocks rich in organic matter and in exceptionally well‐preserved fossils, hence their petroleum and palaeontological interest. However, few studies focused on the dysodiles of Lebanon, leaving their palaeontological content and their depositional environment largely unknown. Our previous investigations show a variety of well‐preserved lacustrine fossils including ichthyofauna, chelonians, insects, plant debris, gastropods, palynomorphs and ostracods. The present study focuses on the ostracods of five sampling localities from the Lower Barremian of Lebanon. Being the most abundant fossils, ostracods raise some taphonomic and palaeoecological questions that may help us understand the depositional environment and enrich the story of these Early Barremian lakes. Faunal differences between the sampled sections provide clues to the specific characteristics of each lake. The abundance of ‘butterfly’ preservation of carapaces, together with the association of adults and juveniles, indicates calm depositional environments without post‐mortem transport. The assemblages comprise ostracods with known swimming ability (Cyprois and Zonocypris) or that are thought to have been swimmers (Cypridea), while non‐swimmers are absent, suggesting that nektobenthonic ostracods must have lived only in the shallow margins of the lakes or among floating vegetation because hypolimnion conditions were hostile to benthos. This study provides different scenarios on how this fauna ended up preserved in the laminated lake sediments.     

珠江三角洲晚第四纪环境演变的微体生物记录

珠江三角洲是主要由西江、北江和东江在珠江口湾内堆积而成的复合三角洲,自晚更新世开始形成。微体生物是珠江河口三角洲晚第四纪沉积记录中的重要组分,是重建第四纪环境变化极为有效的指标。本文样品取自PD孔岩心,钻孔位于珠江三角洲中部(113°28′35″E, 22°53′41″N),孔深16．7 m。代表约25 100年以来的沉积。PD孔中见有孔虫6属10种,其中以Ammonia beccani为优势种。根据地层分布特征,可识别3个有孔虫层位。介形类共7属9种,丰度较低,以Sinocytheridea impressa和Neomonoceratina delicata为代表,在5个层位相对富集。根据微体生物分布特征,并结合有机碳、氮以及磁化率数据,PD孔晚第四纪沉积可划分为6个沉积环境阶段。A．三角洲前缘阶段(-16．7～-13．5 m;25 100～24 000 aB．P．):岩性为粗砂、砂至砂质粉砂。主要含Ammonia属的4个种(A．beccarii、A．sp．、A．tepida、A．annectens),丰度均较低,其中Ammonia beccarii为优势种。不见适应于正常浅海的Elphidium属。介形类以滨岸类型Sinocytheridea impressa为主。出现少量双壳类。代表河口附近水深10 m以内软泥底质、受河流影响大、水体动荡的半咸水环境。有机碳含量和磁化率较低,同样指示低海平面时期的沉积。B．河口湾阶段(-13．5～-10．6 m;24 000～22 400 aB．P．):岩性为粘土质粉砂至粘土。前一阶段出现的Ammonia 4个种的丰度比较高,且出现浅海底栖分子Elphidium advenum和E．hispidulum,以及浮游有孔虫分子Globigerinoides sp．。介形类出现Bicornucythere leizhouensis、B．bisanensis、Alocopocythere goujoni、Pistocythereis bradyformis等适应水深较大的分子,而滨岸浅水分子较少。有机碳含量和磁化率相对升高。代表海平面相对较高时期的沉积。C．三角洲平原阶段(-10．6～-9．2 m;22 400～21 700 aB．P．):底部为一薄层花斑状风化粘土,表明曾短期露出水面。向上变为富含有机质的深灰色粉沙质粘土。不含微体生物和双壳类,反映不稳定的环境条件。由于海平面降低,磁化率表现为持续低值。有机碳和C/N比值很高,表明有丰富的陆源有机质输入,海洋影响较小。D．陆地风化阶段(-9．2～-7．4 m;21 700～3 700 aB．P．):岩性为1．6 m厚的杂色粘土,不含任何生物化石,有机碳含量极低,是长期强烈风化作用的产物,对应于末次冰盛期。E．潮坪沉积阶段(-7．4～-2．5 m;3 700～2 600 aB．P．):互层状深灰色粉沙质粘土、灰色粉沙和细沙,含广盐性有孔虫Ammonia beccarii和Arenoparrella asiatica,以及少量介形类,如Sinocytheridea impressa。典型的半咸水双壳类Potamocorbula amurensis较为丰富,另外还见藤壶和虫管碎片,指示盐度频繁波动的潮间带环境。F．上三角洲平原阶段(2 600 aB．P．以来):未见任何有孔虫和双壳类,顶部见少量陆相介形类Cypria yanchengensis,磁化率变化频繁,C/N比值较高,表明陆相影响不断增强,且环境条件不稳定。研究区晚第四纪古环境的演化是由海平面升降变化控制的。上述沉积环境变化序列表现出两次海进一海退旋回,分别对应阶段A至阶段D和阶段E至阶段F,相应于晚更新世和全新世的两次海平面上升事件。