Effect of delay in a Lotka-Volterra type predator-prey model with a transmissible disease in the predator species

We consider a system of delay differential equations modeling the predator-prey ecoepidemic dynamics with a transmissible disease in the predator population. The time lag in the delay terms represents the predator gestation period. We analyze essential mathematical features of the proposed model such as local and global stability and in addition study the bifurcations arising in some selected situations. Threshold values for a few parameters determining the feasibility and stability conditions of some equilibria are discovered and similarly a threshold is identified for the disease to die out. The parameter thresholds under which the system admits a Hopf bifurcation are investigated both in the presence of zero and non-zero time lag. Numerical simulations support our theoretical analysis.     

一类功能性反应模型在常数收获下的动态特性

具有功能性反应的捕食与被捕食模型具有非常复杂的动态性质．特别是在常数收获下,该模型呈现了各种各样、纷杂多变的动态特性。其中包括正平衡点及其稳定性的变化、各种分叉的产生以及周期解和极限环的出现．本文重点研究了常数收获项对一类功能性反应模型的动态性能的影响,得到了该收获模型存在稳定正平衡点、产生分叉以及在Hopf分叉附近产生周期解和极限环的若干充分条件．     

The role of transmissible diseases in the Holling-Tanner predator-prey model

Transmissible diseases are known to induce remarkable major behavioral changes in predator-prey systems. However, little attention has been paid to model such situations. The latter would allow to predict useful applications in both dynamics and control. Here the Holling-Tanner model is revisited to account for the influence of a transmissible disease, under the assumption that it spreads among the prey species only. We have found the equilibria and analyzed the behavior of the system around each one of them. A threshold result determining when the disease dies out has been identified. We also investigated the parametric space under which the system enters into Hopf and transcritical bifurcations, around the disease free equilibrium. The system is shown to experience neither saddle-node nor pitch-fork bifurcation. Global stability results are obtained by constructing suitable Lyapunov functions.     

Global stability and persistence in LG–Holling type II diseased predator ecosystems

A Leslie–Gower–Holling type II model is modified to introduce a contagious disease in the predator population, assuming that disease cannot propagate to the prey. All the system’s equilibria are determined and the behaviour of the system near them is investigated. The main mathematical issues are global stability and bifurcations for some of the equilibria, together with sufficient conditions for persistence of the ecosystem. Counterintuitive results on the role played by intraspecific competition are highlighted.     

Competing predators for a prey in a chemostat model with periodic nutrient input

A system of ordinary differential equations is used to model the interactions of n competing predators on a single prey population in a chemostat environment with a periodic nutrient input. In the case of one or no predators, criteria for the existence of periodic solutions are given. In the general case, conditions for all populations to persist are derived.Research is in part based on a Ph.D. thesis submitted to the Faculty of Graduate Studies, University of AlbertaResearch is partly supported by the Natural Sciences and Engineering Research Council of Canada, Grant No. NSERC A4823     

Effects of prey refuges on a predator-prey model with a class of functional responses: The role of refuges

In this paper, the effects of refuges used by prey on a predator-prey interaction with a class of functional responses are studied by using the analytical approach. The refuges are considered as two types: a constant proportion of prey and a fixed number of prey using refuges. We will evaluate the effects with regard to the local stability of the interior equilibrium point, the values of the equilibrium density and the long-term dynamics of the interacting populations. The results show that the effects of refuges used by prey increase the equilibrium density of prey population while decrease that of predators. It is also proved that the effects of refuges can stabilize the interior equilibrium point of the considered model, and destabilize it under a very restricted set of conditions which is disagreement with previous results in this field.     

Analysis of a disease transmission model in a population with varying size

An S I R S epidemiological model with vital dynamics in a population of varying size is discussed. A complete global analysis is given which uses a new result to establish the nonexistence of periodic solutions. Results are discussed in terms of three explicit threshold parameters which respectively govern the increase of the total population, the existence and stability of an endemic proportion equilibrium and the growth of the infective population. These lead to two distinct concepts of disease eradication which involve the total number of infectives and their proportion in the population.Partially supported by NSF Grant No. DMS-8703631. This work was done while this author was visiting the University of VictoriaResearch supported in part by NSERC A-8965     

具有庇护所的三种群捕食者-食饵模型

讨论了一类具有庇护所的自治三种群捕食者一食饵模型,运用Liapunov函数方法,得到了该模型持久性的充分条件．对于该模型的周期系统,在一定的条件下,将产生唯一一个全局渐近稳定的周期正解．对更具普遍意义的概周期现象,也得出了概周期正解唯一存在且全局渐近稳定性的充分条件．     

Dynamical behavior of a hepatitis B virus transmission model with vaccination

Hepatitis B virus (HBV) infection is a globally health problem. In 2005, the WHO Western Pacific Regional Office set a goal of reducing chronic HBV infection rate to less than 2% among children five years of age by 2012, as an interim milestone towards the final goal of less than 1%. Many countries made some plans (such as free HBV vaccination program for all neonates in China now) to control the transmission HBV. We develop a model to explore the impact of vaccination and other controlling measures of HBV infection. The model has simple dynamical behavior which has a globally asymptotically stable disease-free equilibrium when the basic reproduction number R₀≤1, and a globally asymptotically stable endemic equilibrium when R₀>1. Numerical simulation results show that the vaccination is a very effective measure to control the infection and they also give some useful comments on controlling the transmission of HBV.     

The impact of mortality on predator population size and stability in systems with stage-structured prey

The relationships between a predator population's mortality rate and its population size and stability are investigated for several simple predator-prey models with stage-structured prey populations. Several alternative models are considered; these differ in their assumptions about the nature of density dependence in the prey's population growth; the nature of stage-transitions; and the stage-selectivity of the predator. Instability occurs at high, rather than low predator mortality rates in most models with highly stage-selective predation; this is the opposite of the effect of mortality on stability in models with homogeneous prey populations. Stage-selective predation also increases the range of parameters that lead to a stable equilibrium. The results suggest that it may be common for a stable predator population to increase in abundance as its own mortality rate increases in stable systems, provided that the predator has a saturating functional response. Sufficiently strong density dependence in the prey generally reverses this outcome, and results in a decrease in predator population size with increasing predator mortality rate. Stability is decreased when the juvenile stage has a fixed duration, but population increases with increasing mortality are still observed in large areas of stable parameter space. This raises two coupled questions which are as yet unanswered; (1) do such increases in population size with higher mortality actually occur in nature; and (2) if not, what prevents them from occurring? Stage-structured prey and stage-related predation can also reverse the 'paradox of enrichment', leading to stability rather than instability when prey growth is increased.     

Persistence and permanence in a metapopulation model with space-limited recruitment

In this paper we analyze a metapopulation model with space-limited recruitment. The model describes the population dynamics of sessile adult and planktonic larvae in a common larval pool. We introduce the basic reproduction number of each species which is the expected number of future larvae reproduced by one larva. We consider the conditions for the persistence of the multi-species and multi-habitats model and the permanence of the single-species model. Subsequently, we consider the conditions for the existence of the non-trivial steady state of the single-species model and its global stability, and the permanence of the two species and two habitats model.     

一类具功能反应的食饵-捕食者两种群模型的极限环的唯一性

考虑具有功能反应的食饵-捕食者两种群模型：x^.=x(a-bx^1/2-h(x))-cyx^1/2,y^.=y(-d ecx^1/2)。对该系统给出了完整的定性分析,证明了该系统极限环的存在与唯一性。     

Analysis of a sterile insect release model with predation

A model for the sterile insect release method of pest control in which the target species is under predatory or parasitic regulation is analyzed. The equations are nondimensionalized and the rescaled parameters are interpreted. There are four types of equilibria, whose existence and stability depend on which of ten regions of parameter space contain the rescaled parameters, and in turn give minimal release rates to achieve eradication of the pest. In at least one region, Hopf bifurcation theory shows the existence of limit cycles, but they are found to be unstable. In addition, the optimal release rate to minimize a total cost functional for pest control by the sterile release method is studied. Both approaches show that when predation accounts for a large fraction of the natural deaths, the necessary release rate and total cost are higher than for weak predation. If the predators are removed without being replaced by any other source of mortality, the cost rises in all cases but rises much more dramatically for cases with strong predation. A definite danger of the sterile release method when some predatory control exists is that the predators are frequently driven extinct before the prey, so that the target species could explode to much higher levels and be more difficult to eradicate again after the sterile release is terminated.     

Global dynamics of a SEIR model with varying total population size

A SEIR model for the transmission of an infectious disease that spreads in a population through direct contact of the hosts is studied. The force of infection is of proportionate mixing type. A threshold sigma is identified which determines the outcome of the disease; if sigma < or = 1, the infected fraction of the population disappears so the disease dies out, while of sigma > 1, the infected fraction persists and a unique endemic equilibrium state is shown, under a mild restriction on the parameters, to be globally asymptotically stable in the interior of the feasible region. Two other threshold parameters sigma' and sigma are also identified; they determine the dynamics of the population sizes in the cases when the disease dies out and when it is endemic, respectively.     

Modelling the effect of treatment and behavioral change in HIV transmission dynamics

In this paper we analyze a model for the HIV-infection transmission in a male homosexual population. In the model we consider two types of infected individuals. Those that are infected but do not know their serological status and/or are not under any sort of clinical /therapeutical treatment, and those who are. The two groups of infectives differ in their incubation time, contact rate with susceptible individuals, and probability of disease transmission. The aim of this article is to study the roles played by detection and changes in sexual behavior in the incidence and prevalence of HIV. The analytical results show that there exists a unique endemic equilibrium which is globally asymptotically stable under a range of parameter values whenever a detection /treatment rate and an indirect measure of the level of infection risk are sufficiently large. However, any level of detection/ treatment rate coupled with a decrease of the transmission probability lowers the incidence rate and prevalence level in the population. In general, only significant reductions in the transmission probability (achieved through, for example, the adoption of safe sexual practices) can contain effectively the spread of the disease.     

A model of a myelinated nerve axon: threshold behaviour and propagation

A model of a myelinated nerve axon is developed on the basis of FitzHugh-Nagumo dynamics under the assumption that the nodes of Ranvier are of small but finite width. It is shown that a periodic excited state may not exist if the width of the nodes is too small and the leakage across the myelin sheath is too great. The propagation of a super threshold pulse is prevented in the absence of nodes. Global stability of the resting equilibrium state is investigated as well as the propagation of wave front, type solutions.     

Global stability of an SEIS epidemic model with recruitment and a varying total population size

This paper considers an SEIS epidemic model that incorporates constant recruitment, disease-caused death and disease latency. The incidence term is of the bilinear mass-action form. It is shown that the global dynamics is completely determined by the basic reproduction number R(0). If R(0)1, a unique endemic equilibrium is globally stable in the interior of the feasible region and the disease persists at the endemic equilibrium.     

Research of population with impulsive perturbations based on dynamics of a neutral delay equation and ecological quality system

This paper studies the global behaviors of a nonlinear autonomous neutral delay differential population model with impulsive perturbation. This model may be suitable for describing the dynamics of population with long larval and short adult phases. It is shown that the system may have global stability of the extinction and positive equilibria, or grow without being bounded under some conditions.     

具有时滞的Hopfield人工神经网络动力系统模型的全局渐近稳定性

Hopfield人工神经网络动力系统模型平衡点的全局渐近稳定性在网络记忆以及最优化等领域具有广泛的应用。本文中,作者研究了一类具有时滞的Hopfield人工神经网络动力系统,通过构造Liapunov泛函的方法,获得了其平衡点全局渐近稳定和局部渐近稳定的充分判定条件。所给出的判定条件只依赖于系统本身的拳数参数和传递函数以及系统中出现的部分时滞。同时,当系统的自身反馈项为负时,此自身反馈项对于系统的稳定性起到稳定化的作用。此外,数值模拟表明时滞的变化对于系统的稳定性具有重要的影响。可破坏系统的稳定性。进而产生周期振动或更为复杂的非线性现象。     

Stability of a Ricker-type competition model and the competitive exclusion principle

Our main objective is to study a Ricker-type competition model of two species. We give a complete analysis of stability and bifurcation and determine the centre manifolds, as well as stable and unstable manifolds. It is shown that the autonomous Ricker competition model exhibits subcritical bifurcation, bubbles, period-doubling bifurcation, but no Neimark–Sacker bifurcations. We exhibit the region in the parameter space where the competition exclusion principle applies.