On the miocene centracanthidae (Teleostei, Perciformes) from the eastern Paratethys

A new genus, Naslavcea, is erected for the Middle Miocene species Spicara fundata Bannikov, 1990 (Centracanthidae) from the Lower Sarmatian of northern Moldova. This genus is characterized by the relatively small number of rays in the dorsal and anal fins. At the same time, Pelates islamdagicus Prokofiev, 2001 from the Lower Miocene of the Apsheron Peninsula (Azerbaijan) is transferred from the family Terapontidae to Centracanthidae and referred to the genus Spicara.     

First record of late middle Miocene Moschidae from Turkey: Micromeryx and Hispanomeryx from Catakbağyaka (Muğla,SW Turkey)

Here, we describe remains of Micromeryx sp. and Hispanomeryx sp. from the middle Miocene locality Catakbağyaka (MN 7/8). This is the first record for Miocene Moschidae for the late middle Miocene in Turkey and verifies the importance of Turkey as a corridor for the dispersal of Eurasian mammals. Furthermore, the record from Catakbağyaka confirms that the sympatric co-occurrence of two Miocene moschid taxa in one locality appears to be a common phenomenon.     

Dabbling ducks (Aves: Anatidae) from the Middle Miocene of Mongolia

A new genus and two new species of ducks (Aves: Anatidae) from the Middle Miocene Sharga locality are described. Mioquerquedula minutissima gen. et sp. nov. is a very small duck. Anas velox Milne-Edwards, 1868 from the Middle Miocene of France is transferred to the genus Mioquerquedula. Aix praeclara sp. nov. described here is the oldest record of the modern genus Aix. A revision of the previously described small duck Anas soporata Kurochkin, 1976 shows that only the specimens from the Sharga locality should be referred to this species. The status of other small ducks from the Neogene of Europe and North America is discussed. The diversity of herbivorous and diving ducks in the Sharga locality indicates that Miocene Shargyn Govi Lake was rich in food resources.     

Palaeoenvironmental significance of Miocene larger benthic foraminifera from the Xisha Islands,South China Sea

We studied 344 samples from Well XK-1 in Xisha Islands, South China Sea, and identified 66 species of larger benthic foraminifera, providing critical evidence for biostratigraphy and palaeoenvironmental interpretation of the Miocene reef carbonate sequence. Three assemblages are recognized, namely, Spiroclypeus higginsi–Borelis pygmaeus Assemblage (Letter Stage Te5, Early Miocene, 1256.28–1180.15 m), Nephrolepidina–Miogypsina Assemblage (Tf, Middle Miocene, 1031.10–577.04 m), and Cycloclypeus–Heterostegina Assemblage (Tg, Late Miocene, 468.13–380.42 m). On the basis of the palaeoecological preference of the larger foraminifera, we interpret that the Miocene carbonate sequence was deposited mainly in a warm tropical shallow water environment, characterized by five stages of continuous long-term evolution: backreef lagoon to shelf in the Early Miocene, normal to frontal reef in the early Middle Miocene, backreef lagoon to shelf in the later Middle Miocene, normal to frontal reef in the early Late Miocene, and proximal forereef shelf in the later Late Miocene.     

A new species of Gaimanophis (Serpentes,Boidae) from the Miocene of northwestern Argentina with remarks on the Neogene boids of South America

Gaimanophis is an extinct boid genus represented so far by a single species (Gaimanophis tenuis) known by isolated vertebrae from the early Miocene of Patagonia. In this paper, a new species of Gaimanophis is described from the India Muerta Formation (late Miocene) of Tucumán province (Argentina). Gaimanophis powelli sp. nov. distinguishes itself from G. tenuis mainly in its larger size, prezygapophyses less slanting, neural spine shorter dorsally than ventrally, and zygosphene straight bearing an anteriorly protruding tongue. This record indicates a wider temporal and geographical distribution of the genus from the early Miocene of Patagonia to the late Miocene of northwestern Argentina. The recognition of a new species of boid in South America increases the known diversity of this group. Although boids have inhabited in this territory since the Paleocene, fossils belonging to the group only show a glimpse of the real past diversity in the continent.     

Kretzoiarctos gen. nov., the Oldest Member of the Giant Panda Clade

The phylogenetic position of the giant panda, Ailuropoda melanoleuca (Carnivora: Ursidae: Ailuropodinae), has been one of the most hotly debated topics by mammalian biologists and paleontologists during the last century. Based on molecular data, it is currently recognized as a true ursid, sister-taxon of the remaining extant bears, from which it would have diverged by the Early Miocene. However, from a paleobiogeographic and chronological perspective, the origin of the giant panda lineage has remained elusive due to the scarcity of the available Miocene fossil record. Until recently, the genus Ailurarctos from the Late Miocene of China (ca. 8–7 mya) was recognized as the oldest undoubted member of the Ailuropodinae, suggesting that the panda lineage might have originated from an Ursavus ancestor. The role of the purported ailuropodine Agriarctos, from the Miocene of Europe, in the origins of this clade has been generally dismissed due to the paucity of the available material. Here, we describe a new ailuropodine genus, Kretzoiarctos gen. nov., based on remains from two Middle Miocene (ca. 12–11 Ma) Spanish localities. A cladistic analysis of fossil and extant members of the Ursoidea confirms the inclusion of the new genus into the Ailuropodinae. Moreover, Kretzoiarctos precedes in time the previously-known, Late Miocene members of the giant panda clade from Eurasia (Agriarctos and Ailurarctos). The former can be therefore considered the oldest recorded member of the giant panda lineage, which has significant implications for understanding the origins of this clade from a paleobiogeographic viewpoint.     

A new duck from the Middle Miocene of Mongolia, with comments on Miocene evolution of ducks

A new relatively large duck, Chenoanas deserta gen. et sp. nov. from the Middle Miocene of the Sharga locality, which is morphologically intermediate between the extant genera Chenonetta and Tachyeres, is described. The diversity of Early and Middle Miocene ducks is discussed. It is noted that some Middle Miocene duck remains are incorrectly referred to the genus Mionetta. The distribution of morphological characters of the humerus in Neogene and extant ducks shows that the present day diversity of ducks apparently results from extinction of some taxa which were formed in the Oligocene-Early Miocene. The distribution of morphological characters in the evolution of diving ducks is evidence that not only the formation of different morphotypes but also so-called ??evolutionary maturation?? of taxa also explains the modern diversity.     

New baleen whale genera (Cetacea, Mammalia) from the miocene of the northern caucasus and ciscaucasia: 1. Kurdalagonus gen. nov. from the middle-late Sarmatian of Adygea

A revision of previously known material and analysis of new records of Miocene baleen whales from the Sarmatian of Adygea (city of Maikop, Blinovo Formation) resulted in the establishment of a new genus, Kurdalagonus with three species: K. mchedlidzei sp. nov. (Middle Sarmatian, Nagiezh-Uashkh locality), K. adygeicus sp. nov. (Upper Sarmatian, Hydroelectric power station locality), and K. maicopicus (Spasskii, 1951) (=Cetotherium maicopicum Spasskii, 1951; Middle Sarmatian, Hydroelectric power station locality).     

Fossil wood of <Emphasis Type="Italic">Laricioxylon erkovetskiense</Emphasis> sp. nov. (Pinaceae) from the Miocene of the Erkovetskii Brown Coal Field,Amur Region,Russia

A new species, Laricioxylon erkovetskiense sp. nov. (Pinaceae) identified on the basis of anatomical features of fossil wood from the Sazanka Formation (upper Middle Miocene–Upper Miocene) of the Erkovetskii Brown Coal Field (Amur Region) is described. The new species shares some characters with the extant larches Larix cajanderi and L. olgensis.     

A new Miocene small-bodied ape from Kenya

A new genus of small-bodied ape from Kalodirr, a Miocene site in northern Kenya, is described. It is compared with other recognized genera of small apes from the Miocene of East Africa and found to show closer similarities with Dendropithecus than with either Limnopithecus or Micropithecus. Among the recognized genera of larger Miocene hominoids it shares most features with Rangwapithecus but is distinguished by details of its dental morphology.     

A Miocene leaf fossil record of Rosa (R. fortuita n. sp.) from its modern diversity center in SW China

Roses (Rosa, Rosaceae) are arguably the most admired ornamental plants in the world. Southwestern China is the center of diversity for many extant native species of Rosa and fossils found in this region are critical for understanding the evolution of this genus. Herein, we report a leaf fossil record with good preservation from the late Miocene of Yunnan Province. The opposite and odd-pinnate leaf is composed of seven elliptical leaflets, with close, crenulate, and regular marginal teeth. The stipules are lanceolate and adnate to the petiole. Additionally, the secondary veins are semicraspedodromous, showing the same venation pattern as most living Rosa species in southwestern China. On the basis of the extensive morphological comparisons, we propose a new species, R. fortuita T. Su et Z.K. Zhou n. sp. This is the first confirmed Rosa leaf fossil record in China, and the discovery of R. fortuita n. sp. indicates that Rosa existed in southwestern China by the late Miocene. It suggests that Rosa was distributed in subtropical or temperate forests and shared a similar ecological niche as Rosa in Europe during the Oligocene and Miocene. The modern diversification of Rosa in southwestern China is thought to have been closely associated with the continuous uplift of the Qinghai-Tibet Plateau since the late Miocene, creating complex topography and a variety of climate conditions.     

Fossil wood of <Emphasis Type="Italic">Pinus priamurensis</Emphasis> sp. nov. (Pinaceae) from the Miocene deposits of the Erkovetskii Brown Coal Field,Amur Region

New species of the pine fossil wood, Pinus priamurensis sp. nov. (Pinaceae) from the Sazanka Formation (upper Middle Miocene–Upper Miocene) of the Erkovetskii Brown Coal Field (Amur Region) is described. The new species shares some wood anatomical features with modern species of the subsection Pinus (section Pinus, subgenus Pinus). This is the first record of fossil wood of Pinus in the Amur Region.     

Creodonts and carnivores from the Middle Miocene Muruyur Formation at Kipsaraman and Cheparawa,Baringo District,Kenya

The Middle Miocene Muruyur Formation (ca 14.5 Ma), Tugen Hills, Kenya, has yielded a huge creodont and a variety of carnivores ranging in size from mongoose-sized viverrids and herpestids to lion-sized amphicyonids. The fauna partly fills what used to be a major gap in our knowledge of Neogene African carnivores, spanning the period between the better known Early Miocene assemblages of western Kenya and eastern Uganda, and the Late Miocene and Plio–Pleistocene faunas of East Africa. Present in the deposits are Megistotherium, two species of Hecubides, one species of Agnotherium, Herpestes, Vishnuictis, and one or two undetermined felids.     

Palaeoreas,Majoreas, and Stryfnotherium gen. nov. (Mammalia: Artiodactyla: Bovidae) from the Late Miocene of Greece

A new comparative study of the two crania, LGPUT RZO-03 and LGPUT RZO-68 attributed to Palaeoreas zouavei Bouvrain, 1980 from the Late Miocene of Axios Valley, Greece reveals great morphological and metric differences between them, which require the exclusion of LGPUT RZO-68 from this species and even from Palaeoreas Gaudry, 1861. This decision leaves unsupported many diagnostic features of P. zouavei applied by previous authors and along with recent data from Bulgaria put in doubt the validity of P. zouavei as a whole. As a result, the genus Palaeoreas is re-defined as monotypic and the cranial morphometric changes of P. lindermayeri (Wagner, 1848) are discussed on a temporal basis. An updated taxonomic analysis confirms the presence of P. lindermayeri in the Late Turolian mammal assemblage of Dytiko-1 (Axios Valley, Greece) but rejects its occurrence at Dytiko-2. Instead, a re-examination of old cranial specimens and a study of new ones suggest the presence in both sites of Majoreas Kostopoulos, 2004 a Late Miocene taxon previously mostly known from the eastern banks of the Aegean Sea. A reassessment of the LGPUT RZO-68 cranium, and its comparison with Palaeoreas and Criotherium Forsyth-Major, 1891, suggests that it deserves taxonomic distinction at genus and species levels: Stryfnotherium exophthalmon.     

The Ramblian-Aragonian boundary and its significance for the European Neogene continental chronology. Contributions from the Ebro Basin record (NE Spain)

In this paper we report on the results of a paleontological and paleomagnetic sampling carried out in the Miocene of the central Ebro Basin. It includes the sections of Albalatillo, Lanaja, La Almolda and San Caprasio in the Sierra de Alcubierre, which delivered six new micromammal levels ranging from the MN3 to MN5 mammal units. The magnetobiostratigraphic study of the Sierra de Alcubierre enables one to shed light on the chronology of the Early to Middle Miocene European mammal biozonation (e.g., the Ramblian-Aragonian boundary). As a result, the first occurrence of the “Miocene Cricetids” and, therefore, the lower boundary of MN4 – zone B, is set back to the lower part of chron C5Cr, thus reducing significantly the supposed discrepancy of the First Occurrence of Democricetodon in the Swiss Molasse and the Aragonian type-section. Another significant result is the persistence of Vasseuromys in the Ebro Basin up to the Middle Miocene (MN5).     

A Late Miocene Paratethyan mollusc fauna from the Denizli Basin (southwestern Anatolia, Turkey) and its regional palaeobiogeographic implications

We report on a Late Miocene mollusc fauna from a single locality in the Denizli Basin in southwestern Turkey that is composed of ten presumably euryhaline species and three freshwater species. The fauna is remarkably distinct from faunas of the adjacent Late Miocene Aegean and Euxinian Basins, and has elements in common with the modern Caspian Sea fauna in the form of Didacna species. The suggested age of the fauna (Late Miocene) would extend the stratigraphic range of the lymnocardiid genus Didacna (hitherto Calabrian-Extant) considerably. The palaeobiogeographic significance of the Denizli fauna is discussed.     

Late Miocene lagomorphs from the Republic of Moldova

The aim of this paper is to enlarge knowledge about still poorly documented and understood record of Lagomorpha (Mammalia) from the Late Miocene of Moldova. The lagomorph material from two new Late Miocene localities, Razeni (MN11/12), Gradishte (MN12), and re-sampling locality Chimislyia (MN12), of Southern Moldova, is described in detail here. The localities yielded small vertebrate fauna including five lagomorph taxa: Prolagus cf. oeningensis, “Proochotona” cf. eximia, Alilepus laskarewi, “Proochotona” sp., and Alilepus sp. P. cf. oeningensis is described for the first time in the studied area. A short review of the lagomorph record from the Republic of Moldova is provided. This contribution outlines the importance of the lagomorphs for biostratigraphic purposes, and highlighs the gaps in our knowledge related to the faunal succession in this area.     

New record of Discradisca (Brachiopoda: Discinidae) from the Early Miocene of the Aquitaine Basin,south-western France

The linguliform inarticulate brachiopod genus Discradisca Stenzel, 1964 has been identified for the first time from the Early Burdigalian (Early Miocene) sandy deposits of the Aquitaine Basin, south-western France. The species studied here, D. multiradiata (de Morgan, 1915), originally described from the Middle Miocene of the Loire Basin, north-western France, is characterized by a small size, ornamentation of numerous (up to 100), fine but distinct and beaded tuberculate ribs, and an apex situated subposteriorly to subcentrally. The ecological and taphonomic aspects of this species are also presented. The attribution of some previously described Discinisca Dall, 1871 species from the Tertiary to the genus Discradisca is discussed.