Upper Devonian shoal-water delta integrated with cyclic back-reef facies off the Mowanbini Archipelago (Canning Basin), Western Australia

The Oscar Range in Western Australia’s Canning Basin exhibits folded Proterozoic, quartzite, quartzite conglomerate, phyllite, and metavolcanic rocks that survive with positive relief. Facies of the Pillara Limestone were deposited around this relief during Late Devonian (Frasnian) time. A segment of the Great Devonian Barrier Reef with a linear reef margin strikes parallel to the outer paleoislands in the Mowanbini Archipelago. A more sheltered strait separates inner islands from the cratonic Devonian mainland on the Kimberley Block. Large fan-deltas emanated from the craton, but locally small shoal-water deltas prograded from a drainage basin on one of the larger paleoislands in the Oscar Range. That island is expressed today by local topography exhumed from beneath a cover of former Devonian, Carboniferous, and Permian strata. The Devonian shoal-water delta rests unconformably on tilted Proterozoic phyllite and incorporates abundant phyllitic debris accumulated under fluvial to shoreface conditions. Some quartzite pebbles and hydrothermal quartz were derived from a source more than a kilometer away. Rare gastropods and stromatoporoid fragments in the deltaic sediments were abraded from the adjacent reef margin. The clast-supported conglomerate in the exposed shoal-water delta is mapped over a distance of 130 m to within 15 m of the inner reef margin, exposed nearby on steeply dipping phyllite. A cyclic succession of mixed clastic and carbonate parasequences, 31.5 m in thickness, follows above a disconformity surface on the delta-top facies. The overall succession represents a minor fall in relative sea level associated with erosion of delta facies and a major transgression characterized by a retrograde parasequence stacking pattern. The succession shifts through siliciclastic-rich shoreface to intertidal distal back-reef facies, ending with a subtidal, siliciclastic-poor proximal back-reef facies. The study demonstrates how variability in sedimentary cycles is influenced by local paleogeographic constraints in an island system dominated by quartzite highlands and phyllite lowlands.     

Earth's oldest liverworts—Metzgeriothallus sharonae sp. nov. from the Middle Devonian (Givetian) of eastern New York,USA

Liverworts are generally regarded as rare elements in Palaeozoic floral assemblages. However, a focus on dark gray to black shales and siltstones in the Middle–Late Devonian Catskill Delta of eastern New York shows that liverworts are locally quite common as well-preserved, apparently parautochthonous specimens in thin, lenticular, dark gray–black shale and siltstone lenses. These lenses are either dysoxic–anoxic lacustrine or estuarine facies deposited under oxygen-stratified water masses or rapidly deposited flood plain deposits that were not oxidized after deposition. Carbonized remains of the upper Middle Devonian (Givetian) liverwort Metzgeriothallus sharonae sp. nov. are locally common in these lenses. Well-preserved thalli (gametophytes) are only evident by projecting polarized light on the shale and siltstone surfaces. An associated sporophyte capsule is the first evidence of a reproductive structure in a Devonian liverwort. Metzgeriothallus sharonae sp. nov. is the oldest known liverwort. The age of the new species helps recalibrate chloroplast DNA studies that have led to proposals of the timing of liverwort diversification by showing that the evolutionary separations of the Jungermanniopsida and Marchantiopsida and of the Metzgeriidae and Jungermanniidae [previously thought to be Late Devonian and Late Carboniferous, respectively] were no younger than late Middle Devonian.     

Geological and geochemical aspects of a Devonian siliceous succession in northern Thailand: Implications for the opening of the Paleo-Tethys

The opening of the Paleo-Tethys are reconstructed, including the depositional setting and redox conditions, based on an analysis of radiolarian fossils and the geochemistry of a Devonian siliceous succession in the Chiang Dao area of northern Thailand. The succession is subdivided into the following five rock types (in ascending stratigraphic order): black shale (Lower Devonian), siliceous shale (Middle Devonian), tuffaceous chert and tuff (Middle/Upper Devonian), and chert (Upper Devonian). The succession was deposited in continental margin and pelagic environments between the Sibumasu Block and the Indochina-North China blocks. The concentrations of terrestrial-derived elements (Al₂O₃, TiO₂, Rb, and Zr) suggest that the succession (except for the chert) was supplied with terrigenous material and volcanic ash from the adjacent continent, deposited within a SiO₂-rich environment. Geochemical indicators of redox conditions (total organic carbon and the Th/U ratio) reveal a gradual change from anoxic to oxic oceanic conditions between the black shale and chert. Taking into account the interpreted depositional setting and redox conditions, the initial Paleo-Tethys developed as a small, closed anoxic-suboxic oceanic basin during the Early to Middle Devonian, located close to the continental margin. Black shale and siliceous shale were deposited in the basin at this time. Opening of the Paleo-Tethys started around the Middle and Upper Devonian boundary, marked by voluminous volcanic activity. The tuffaceous chert was deposited under oxic conditions, suggesting that ash and pumice within the chert were derived from a continental source. After the Late Devonian, the Paleo-Tethys developed as a deep, broad ocean in which pelagic chert was deposited.     

Phanerozoic changes in the high-rank suprageneric diversity structure of brachiopods: Linear and non-linear effects

Phanerozoic evolution of brachiopods produced many linear (established by a comparison of successive geologic time units) and non-linear (established by a comparison of non-successive geologic time units) effects, which can be examined quantitatively by using the similarity coefficients (Czekanowski's Quantified Coefficient and Gower Index) and correlation tools. The high-rank suprageneric diversity structure accounts for a number of superfamilies in each of 26 orders for every epoch of geological time. The intensity of turnovers in this structure was generally low during the entire Phanerozoic. It was slightly stronger during the Early Paleozoic, but close to zero during the Cenozoic, when the high-rank suprageneric diversity structure of brachiopods stabilized finally. Significant turnovers took place at the Middle Cambrian–Early Ordovician, the Late Ordovician–Early Silurian, the Late Silurian–Early Devonian, the Middle Devonian–Mississippian, and the Permian–Triassic transitions. Influences of mass extinctions, both major like those End Ordovician or Permian/Triassic and minor like Early Jurassic or Jurassic/Cretaceous, on the high-rank suprageneric diversity structure of brachiopods is registered. The strongest was the consequences of the Permian/Triassic catastrophe, which perhaps even reset the brachiopod evolution. No evident direct relationships are established between intensity of turnovers and eustatic fluctuations. However, the changes in the diversity structure recorded with the Gower Index provide evidence that eustatic lowstands were more favorable for intensification in these changes.     

Palaeoecology and diversity of endosymbionts in Palaeozoic marine invertebrates: Trace fossil evidence

Endosymbionts are organisms that live within the growing skeleton of a live host organism, producing a cavity called a bioclaustration. The endosymbiont lives inside the bioclaustration, which it forms by locally inhibiting the normal skeletal growth of the host, a behaviour given the new ethological category, impedichnia. As trace fossils, bioclaustrations are direct evidence of past symbioses and are first recognized from the Late Ordovician (Caradoc). Bioclaustrations have a wide geographic distribution and occur in various skeletal marine invertebrates, including tabulate and rugose corals, calcareous sponges, bryozoans, brachiopods, and crinoids. Ten bioclaustration ichnogenera are recognized and occur preferentially in particular host taxa, suggesting host-specificity among Palaeozoic endosymbionts. The diversity of bioclaustrations increased during the Silurian and reached a climax by the late Middle Devonian (Givetian). A collapse in bioclaustration diversity and abundance during the Late Devonian is most significant among endosymbionts of host coral and calcareous sponge taxa that were in decline leading up to the Frasnian-Famennian mass extinction.     

AN ORDOVICIAN LOBOPODIAN FROM THE SOOM SHALE LAGERSTÄTTE, SOUTH AFRICA

Abstract:  The first lobopodian known from the Ordovician is described from the Soom Shale Lagerstätte, South Africa. The organism shows features homologous to Palaeozoic marine lobopodians described from the Middle Cambrian Burgess Shale, the Lower Cambrian Chengjiang biota, the Lower Cambrian Sirius Passet Lagerstätte and the Lower Cambrian of the Baltic. The discovery provides a link between marine Cambrian lobopodians and younger forms from the Silurian and Carboniferous. The new fossil preserves an annulated trunk, lobopods with clear annulations, and curved claws. It represents a rare record of a benthic organism from the Soom Shale, and demonstrates intermittent water oxygenation during the deposition of the unit.     

The Ordovician Biodiversification: revolution in the oceanic trophic chain

The Early Palaeozoic phytoplankton (acritarch) radiation paralleled a long-term increase in sea level between the Early Cambrian and the Late Ordovician. In the Late Cambrian, after the SPICE δ¹³C_carb excursion, acritarchs underwent a major change in morphological disparity and their taxonomical diversity increased to reach highest values during the Middle Ordovician (Darriwilian). This highest phytoplankton diversity of the Palaeozoic was possibly the result of palaeogeography (greatest continental dispersal) and major orogenic and volcanic activity, which provided maximum ecospace and large amounts of nutrients. With its warm climate and high atmospheric CO₂ levels, the Ordovician was similar to the Cretaceous: a period when phytoplankton diversity was at its maximum during the Mesozoic. With increased phytoplankton availability in the Late Cambrian and Ordovician a radiation of zooplanktonic organisms took place at the same time as a major diversification of suspension feeders. In addition, planktotrophy originated in invertebrate larvae during the Late Cambrian–Early Ordovician. These important changes in the trophic chain can be considered as a major palaeoecological revolution (part of the rise of the Palaeozoic Evolutionary Fauna of Sepkoski). There is now sufficient evidence that this trophic chain revolution was related to the diversification of the phytoplankton, of which the organic-walled fraction is partly preserved.     

A myriapod-like arthropod from the Upper Cambrian of East Siberia

Although the fossil record of biramous arthropods commences in the Lower Cambrian, unequivocal uniramous arthropods do not appear until the Upper Silurian, in association with terrestrial biotas. Here we report an Upper Cambrian marine arthropod from East Siberia that possesses some significant myriapodan features. The new arthropod,Xanthomyria spinosa n. gen., n. sp., closely resembles examples of archipolypodans from the Late Palaeozoic. If this resemblance genuinely represents myriapod affinities, this would be the first convincing myriapod from the Cambrian. Suggestions of an early branching point of the myriapods from other arthropods would be consistent with this. Conversely, an as yet poorly known clade of multi-segmented arthropods may exist in the Cambrian, with no close affinities to the myriapods.      

Significance of Devonian-Carboniferous radiolarians from accretionary terranes of the New England orogen, eastern Australia

Radiolarians provide age constraints for many previously undated terranes in the New England Orogen (NEO), a tectonic collage developed along the eastern margin of Australia.Djungati terrane, the age range of which was previously unknown, contains two distinctive siliceous sedimentary lithofacies. The oldest is a thick sequence of red, ribbon-bedded cherts which probably accumulated in a deep ocean-floor setting far from land. Middle Silurian through Late Devonian radiolarians have been recovered from these cherts. Green tuffaceous cherts which contain a latest Devonian (Famennian) radiolarian fauna depositionally overlie the lower red ribbon-bedded chert sequence. These cherts are intercalated with volcaniclastic sediments and the fauna which they contain can be used to constrain the timing of accretion of older rocks into a subduction complex.Anaiwan terrane, which was also previously undated, contains thin ribbon-bedded cherts which are depositionally overlain by tuffaceous chert, siliceous siltstones and volcaniclastic sediments. Latest Devonian (?late Famennian) and Early Carboniferous radiolarians have been recovered from these cherts and tuffaceous siltstones.Radiolarians also occur in fine-grained siliceous sediments of the Yarrimie Formation, part of the Gamilaroi terrane. These radiolarians are of Late Devonian (Frasnian) affinity and their presence indicates that blocks of limestone, which contain Givetian conodonts and corals and were previously thought to indicate the age of the Yarrimie Formation, are allochthonous.     

A Middle and Late Cambrian age for the Booley Bay Formation,County Wexford,Ireland: New acritarch data and its implications

A well preserved Ediacara-type fauna was recorded for the first time a few years ago from the bases of seven sandstone beds in the type section of the Lower Palaeozoic Booley Bay Formation in County Wexford, Ireland. Four microfossil samples from mudstones interbedded with the sandstones yielded an acritarch microflora considered indicative of a late Late Cambrian age for the Ediacara fauna, thus suggesting extension of its range from the Neoproterozoic to the Late Cambrian, rather than to the Middle Cambrian as had been previously suggested. The current study examines the microfossils in more detail, with 18 samples, all of which were productive. One additional Ediacara bed approximately 100 m above those recorded earlier was also observed. The exceptionally well preserved acritarch assemblages indicate a clear Middle Cambrian age for the Ediacara beds, thus predating the previously suggested upper limit for Ediacara faunas. The uppermost part of the formation, above known occurrences of Ediacara beds, is younger; that is, early Late Cambrian, probably coeval with the Olenus trilobite zone. In addition, the age now proposed for the Booley Bay Formation shows that it is part of the Lower Palaeozoic Ribband Group of southeast Ireland, and not part of the Cahore Group as has recently been suggested by the Irish Geological Survey. Thirty-seven acritarch species (or species groups) are distinguished and illustrated. One new species, Stellechinatum ? mariatheresae Vanguestaine, is established.     

SCAPHOPODIZATION IN PALAEOZOIC MOLLUSCS

Abstract:  The relationship of a variety of problematic Early Palaeozoic fossils to the true scaphopods (Late Palaeozoic–Recent) is clarified by the recognition that a trend towards the development of a tubular shell is a recurrent theme within Palaeozoic benthic molluscs. This trend is here termed scaphopodization and can be recognized already in the Cambrian in helcionelloids such as Yochelcionella and Eotebenna . The problematic Early Palaeozoic tubular fossils Janospira , Jinonicella and Rhytiodentalium are interpreted as pseudo-scaphopod derivatives of the apparently exogastric ribeirioid rostroconchs. The direction of coiling of the protoconch indicates that they are not closely related to the true scaphopods, which were derived from endogastric concocardioidean rostroconchs in the Devonian or Carboniferous. Scaphopods show a comparable underlying morphological blueprint to these conocardioideans, but they represent a distinct stream-lining to the infaunal habitat.     

Sedimentary environment of Devonian pelagic limestones in the Southern Alps

Primary sedimentary structures combined with the geometry of Devonian and Lower Palaeozoic lithosomes in the Southern Alps (Austria and Italy) suggest many doubts about the published environmental and bathymetric interpretation of some Lower to Middle Devonian pelagic nodular limestones as 'deep-water' abyssal deposits. Every graded bed is not necessarily aturbidite; a single or some turbiditic beds are not necessarily deep-water or abyssal deposits; carbonate dissolution does not necessarily take place only below the carbonate compensation depth (CCD). Data show that graded allodapic beds, like those of the Lower to Middle Devonian of the Alps, may have been deposited as storm layers at a depth not exceeding some hundreds of metres. Accordingly the margins between Lower Devonian shallow-water platform and basins were characterized by low gradient and transitional sedimentary conditions. They became very steep only at the Devonian–Carboniferous transition because of synsedimentary block faulting.     

Silicified Middle Cambrian trilobites from Kyrgyzstan

Abstract:  A medium diversity silicified Middle Cambrian faunal assemblage occurs in an olistolith of bioclastic limestone within Silurian siliciclastic turbidites on the slopes of Arpatektyr Mountain in the eastern Alai Range, Kyrgyzstan. The fauna includes the trilobites Raragnostus cf. mirus , Altikolia kurshabica sp. nov., Corynexochina weberi , Dorypyge richthofeniformis , aff. Jincella sp., Olenoides comptus , Skreiaspis aff. spinosus and Suluktella mambetovi sp. nov. Three of these taxa ( C .  weberi , O .  comptus and R . cf.  mirus ) are considered locally to be characteristic of the Pseudanomocarina Beds, although Pseudanomocarina itself is not present. Altikolia and Suluktella are endemic to early Palaeozoic terranes of Central Asia, Corynexochina also occurs in South China and Skreiaspis is known elsewhere exclusively from the Middle Cambrian of 'West' Gondwana. Because of preservation in full relief, the silicified trilobite fauna allows a detailed understanding of some morphological structures, related especially to trilobite enrolment, otherwise inadequately known for many Mid-Cambrian polymerid trilobites.     

Three-dimensional preservation of arthropod integument from the Middle Cambrian of Australia

Three-dimensional preservation of arthropod soft integument occurs in Middle Cambrian sediments of the Georgina Basin, western Queensland, Australia. The beds are referred to the Monastery Creek Phosphorite Formation, Gowers Formation, Inca Shale Formation and Devon-court Limestone Formation. The finds include arthropod type-A larvae previously described by Müller & Walossek and several complete appendages, possibly of early Palaeozoic 'ostracodes', as well as indeterminable cuticular remains. The Australian sediments were in part deposited under high water-energy conditions, whereas previously known occurrences of three-dimensional soft-integument preservation have been from environments of lower water energy. Such preservation may thus be more widespread in the Early Paleozoic than hitherto known. □ Phosphatization, arthropods, type-A larvae, appendages, Middle Cambrian, Geogina Basin, Australia, three-dimensional preservation.     

巴西中部元古代Paranoá和Bambui群叠层石的古生态和构造背景

叠层石的形态和它们与陆源沉积的关系反映了沉积盆地的构造背景.在巴西中部地区,叠层石出现在中元古和新元古代的Paranoa群、Vazante群和Bambui群.根据沉积学和矿物学的研究分析,Paranoá群形成于被动边缘盆地,而另外两个群形成于前陆盆地.Vazante群沉积在形成巴西褶皱带的山脉附近,然而Bambui群的大部分都形成于克拉通地区.通常远离山脉褶皱带的Paranoá群和B2mbuí群内的叠层石建造的形态和规模的改变与环境的变化有关.在柱状生长方向的频繁变化和不规则的丰富陆源物质的夹层进入叠层石,表征活跃的构造背景,例如在Vazante,Paranoá群和Bambuí群的西部.     

Stratigraphy, geometry, and facies of a Middle Devonian ramp-to-basin transect (Eastern Anti-Atlas, SE Morocco)

Summary In the castern Anti-Atlas (SE Morocco), a small sedimentary basin (Mader Basin) evolved during the late Palacozoic. The Middle Devonian deposits consist of shales and limestones with a thickness up to 700 m in the depocentre. Sedimentary structures and sole marks of Middle Devonian limestones indicate transport from the northwest and the south towards the basin centre, located in the central Mader area. Lithostratigraphic, biostratigraphic, and dynamic stratigraphic approaches were applied to correlate stratigraphic sections. Five correlatable large-scale base-level hemicycles were recognized in the Middle Devonian succession. Thickness trends of Middle Devonian deposits, regional correlations, and facies-trends reveal the geometry of a carbonate ramp. The carbonate ramp was slightly inclined (<1°) to the NE. A sedimentary wedge, consisting of limestones and limestone/marl alternations, was deposited during the Eifelian and marks the transition from the ramp to the adjacent basin. Middle Devonian water depths are estimated as close to and within the storm wave-base at the southern area of the ramp and far below storm wave-base in the northeastern part of the ramp. Shallowest conditions (inner-ramp environment), close or within the fairweather wave-base, existed during the early Givetian as documented by the abundance of recfal fauna (stromatoporoids, corals) and calcimicrobes (lumps, micritic envelopes) in the eastern and southeastern area of the ramp.     

Permian and Triassic radiolarians from chert breccia in the Nong Prue area,western Thailand: its origin and depositional setting in the Paleotethys

Bedded chert and siliceous shale successions previously regarded as the Silurian–Devonian rock units, distributed in the Nong Prue area, northwest of Kanchanaburi, western Thailand, yielded Lopingian (upper Permian) and Lower–Middle Triassic radiolarians. We found chert breccia layers in northern Nong Prue area, mainly consisting of angular to sub-angular chert clasts with matrices of silt-sized chert grains and clay minerals. We discriminated uppermost Pennsylvanian–Lopingian (upper Carboniferous–upper Permian) and Middle Triassic radiolarian-bearing chert clasts from four different levels of the chert breccia; 28 species of 15 genera with one radiolarian gen. et sp. indet. are identified. On the basis of sedimentary characteristics of the chert breccia, we suggest that the chert breccia is of sedimentary origin. The radiolarian assemblages reported here, together with previously known lithological and paleontological evidence, further indicate that the chert breccia was deposited in the Paleotethys with chert clasts derived from fine grained siliceous rocks in the continental margin to deep ocean basin of the Sibumasu Terrane.     

An Odontogriphid from the Upper Permian of Australia

A problematic organism, Bowengriphus perphlegis gen. et sp. nov., is described based on two specimens from the Late Permian Rangal Coal Measures of eastern central Queensland. It displays a double-looped feeding apparatus bearing small conical elements, considered homologous with that of the supposed lophophorate Odontogriphus omalus Conway Morris, 1976, from the Middle Cambrian Burgess Shale of British Columbia, Canada. Odontogriphids are thus interpreted as a group that survived through the Palaeozoic and made the transition from marine to freshwater environments. Recent proposals that odontogriphids are stem-group chordates are not well established.     

A Late Devonian isoetalean lycopsid, Otzinachsonia Beerboweri, gen. et sp. nov., from north-central Pennsylvania, USA

Compressions and impressions of an isoetalean lycopsid, comprising lower portions of stems, lobed bases, attached rootlets, and rounded rootlet scars, discovered in Late Devonian (Famennian) rocks of Clinton County, north-central Pennsylvania, Appalachian Basin, USA, are here described as Otzinachsonia beerboweri, gen. et sp. nov. These specimens demonstrate unequivocally the existence of the isoetalean lobe-and-furrow rhizomorphic growth pattern as early as the Late Devonian. They were found in an Archaeopteris- and Rhacophyton-dominated flora at Red Hill, an outcrop of the Duncannon Member of the Catskill Formation. The fossils were found in a dark-gray to greenish-gray lenticular siltstone layer that has an average thickness of 1.0 m. This deposit is interpreted as a floodplain pond. The low-energy nature of the deposit and the fine preservation of the intact rootlets of the specimens imply little or no transport. The plants were probably growing along the edge of the floodplain pond with their lower portions submerged for at least part of the year.     

Reassessment of the Devonian problematicum Protonympha as another post‐Ediacaran vendobiont

Protonympha is an enigmatic fossil represented by two species from the Middle Devonian (Protonympha transversa) and Late Devonian (Protonympha salicifolia) of New York. Although interpreted in the past as a polychaete worm or starfish arm, Protonympha is not found with marine fossils, but with fossil plants. This fossil plant community was a swamp woodland of Lepidosigillaria whitei, with ground cover of Haskinsia colophylla, fringing brackish to freshwater coastal lagoons of the Catskill Delta. Protonympha shares with Ediacaran Vendobionta a quilted body of unskeletonized biopolymer that is unusually resistant to burial compaction. In overall form, Protonympha is most like the Ediacaran genus Spriggina. Protonympha has branching and tapering tubular structures radiating from the bottom. These rhizine‐like structures, thallus stratification and internal chambers revealed by petrographic thin sections suggest affinities with lichenized fungi. As for Cambrian Swartpuntia and Ordovician–Silurian Rutgersella, Protonympha may have been a post‐Ediacaran vendobiont.