Stone Age to recent land-use history at Pegrema, northern Lake Onega, Russian Karelia

The village of Pegrema in Karelia may be regarded as a Stone Age innovation centre in the large Lake Onega area. Two pollen and plant macrofossil diagrams are presented which represent the first contribution to the study of human impact in the area using anthropogenic pollen indicators. A continuous but sporadic human presence from the Mesolithic onwards is demonstrated. While there is no archaeological evidence relating to the period 4200–3000 B.P., the pollen data suggest continuous, though rather sparse human presence. The data do not support any natural catastrophes in Pegrema as has been suggested elsewhere. Cerealia pollen is recorded earlier than expected (c. 5000 B.P.=. In the Bronce Age and Iron Age, the settlement of the Zaonezhye peninsula is reflected by a slight increase in herb pollen representation, sporadic Cerealia pollen and several periods of regression in Picea. The long introductory period of agriculture to the area, as well as the similarities and discrepancies between different sources of evidence (palaeoecological, archaeological and historical) are discussed at some length. The start of land clearance for permanent cultivation in the profile Pegrema S was dated to the late 13^th century. The beginning of more intensive field cultivation in the 15^th century is clearly seen in the pollen succession at both localities. The fluctuation in anthropogenic indicators can be related to population density based on historical data. The village of Pegrema was depopulated in 1956 which is reflected in a distinct decline in settlement indicators. Received February 17 / Accepted May 19, 2000     

Late Glacial and early Holocene environment in the middle Lahn river valley (Hessen,central-west Germany) and the local impact of early Mesolithic people—pollen and macrofossil evidence

The Late Glacial to early Holocene river valley landscape of the middle Lahntal in Hessen, central-west Germany, is reconstructed by means of pollen and macrofossil analyses. AMS ¹⁴C dating combined with pollen, macrofossil and geomorphological mapping provide a detailed chronology of the floodplain sediments of the river Lahn. Archaeological evidence for early Mesolithic settlements in the middle Lahntal is backed up by pollen and macrofossil evidence, which indicates an increase in light demanding plants, together with ones indicating nutrient-rich and disturbed environments and more macroscopic charcoal and charred pieces of pine. Different phases of human impact were dated; a first phase was dated at ca. 9,270 uncal b.p. and a second phase at ca. 9,120 uncal b.p. Comparison of the palynological data from different fluvial channel fills demonstrates that during this second phase, between ca. 9,120–8,700 uncal b.p., there were several cycles of woodland clearance. The outcome of combined archaeological, palynological and macrofossil data is discussed in terms of the impact of early Mesolithic people using fire on the vegetation during the early Holocene in the middle Lahntal.     

Palaeoecological studies at the Mesolithic site at Bedburg-Königshov near Cologne, Germany

The Late Glacial and early Holocene palaeovegetation is reconstructed by analysis of pollen and macroscopic plant remains from sediments of the lower river Erft valley, near Cologne, exposed by the Garzweiler opencast lignite mine. The study was carried out in parallel with the archaeological excavation of the Mesolithic site of Bedburg-Königshoven, located within a former meander of the Erft. During the Younger Dryas period, the study region was thinly wooded. Relatively open pine woods existed during the Preboreal, becoming more closed during the Boreal. Later in the Boreal,Corylus and taxa of the mixed oak woodland (Ulmus, Quercus, Tilia andFraxinus) appeared. The Atlantic period was characterised by mixed oak woods with dominantUlmus. At the beginning of the Subboreal,Ulmus declined andTilia became the predominant element of the woods. The Mesolithic settlement existed during the early and middle part of the Preboreal period. At that time, the woodland of the Erft valley was mainly composed ofPinits, Belula and somePopulus. The pollen results do not show any evidence of the activity of Mesolithic people. Local vegetational changes of the filling process of the Erft meander were described and interpreted. The vegetational development of the study region is compared with that of neighbouring regions.     

The significance of the oil palm (Elaeis guineensis Jacq.) in the late Holocene environments of west and west central Africa: A further consideration

Abundant archaeological evidence of the occurrence of the endocarp of the oil palm,Elaeis guineensis, in the rain forest and woodland savanna zones of west and central Africa from about 5000 B.P. has shown the tree to be an important element in the subsistence economy of the region; its pollen also has been recorded in most of the regional terrestrial sediments studied so far. The distinct and consistently sudden and more marked increases in this pollen during the late Holocene when compared with the late Tertiary and late Pleistocene frequencies strongly indicate that the late Holocene upsurges were due to both natural and human factors favourable for the expansion of this heliophytic tree. Reasons are given for suggesting that upsurges in oil palm pollen during the late Holocene period in this region can be used as indices of plant cultivation. While the oil palm is known from early Tertiary deposits in west Africa, its earliest palynological record from terrestrial sediments in the west central part dates back only to the early Holocene. More palynological studies of Tertiary and Quaternary terrestrial cores are required to establish with more certainty the antiquity ofE. guineensis in west central Africa.     

Evaluation of Holocene pollen records from the Romanian Plain

This study is a critical review of pollen analyses carried out on Holocene sequences from 15 sites in and near the Romanian Plain. Three sites come from natural sediments, 10 sites are from anthropogenic deposits and two are from both anthropogenic and natural settings. The general reconstruction is of a steppe-forest-steppe vegetation through the Holocene. The nature of the deposits, however, casts doubts on this reconstruction. Deposits of archaeological sites generally yield pollen spectra that are influenced by human activities and thus unsuitable for vegetation reconstructions. Loess deposits are also unfavorable for pollen preservation because of high pH and porosity. Consequently, pollen spectra from loess deposits are strongly biased by selective pollen destruction. Research and experiments carried out by several authors suggest that spectra dominated by Asteraceae, Poaceae, Chenopodiaceae or Pinus pollen in soils and loess are a result of selective pollen destruction, especially if low pollen concentrations, progressive pollen deterioration or high frequencies of deteriorated or unidentifiable pollen are evidenced. The fact that pollen records from the Romanian Plain come from loess, alkaline peat or archaeological sites reduces their reliability for reconstructions of vegetation. The vegetation history of similar regions in Hungary, Bulgaria and Turkey suggests that early Holocene steppe vegetation was gradually replaced by forest or forest-steppe vegetation in the late Holocene. Records from lake sediments are required to find out whether the Holocene vegetation history of the Romanian Plain was similar.     

The late Holocene palaeoecological sequence of Serranía de las Villuercas (southern Meseta,western Spain)

New pollen results and radiocarbon dating from a valley mire in south-western Spain are presented. This is a region where few palaeoecological records have been preserved and the sequence yields important new palaeobotanical evidence for the late Holocene. The landscape is shown as having been largely open woodland, but more wooded than at present. The vegetation history of the last four millennia in these montane territories of south-western Iberia is discussed in the light of anthropogenic indicators, archaeological and documentary archives; fire incidence and climate change. Alnus is the predominant pollen type, although a decline is noted during the last couple of centuries. Its presence is connected with local topography behaving as a phreatophyte, that is, a plant which obtains a significant amount of water from the zone of saturated soil. A framework is also provided for the age and ecological dynamics of some major woodland taxa—Betula, Corylus, Ilex, evergreen and deciduous oaks, Ericaceae and Pinus. In addition, the natural status of several pollen taxa and local trends in biodiversity are discussed. We consider that the results of our work will have important implications for the understanding of the vegetation history in a floristically very rich area, with a noticeable diversity of woody taxa, and a relatively well preserved ecosystem structure.     

Late-glacial and Holocene woodland dynamics and land-use history of the Lower Oder valley, north-eastern Germany, based on two, AMS14C-dated, pollen profiles

High-resolution Holocene pollen profiles from lakes Großer Krebssee and Felchowsee, in the Lower Oder valley, north-eastern Germany, are presented. The Großer Krebssee profile includes a Late-glacial sequence. These investigations have been carried out in the context of a programme of archaeological excavation. AMS radiocarbon dates (26 in all) based on pollen concentrates have been used to provide a chronology for the pollen records. Holocene forest history and human impact are reconstructed for contrasting landscapes, namely, the Neuenhagener Oderinsel in the Lower Oderbruch (Großer Krebssee profile) and the more fertile Uckermärker Hügelland (Felchowsee profile) that lies immediately to the north. Both landscapes were glaciated during the Pomeranian stage of the Weichselian. New information on the spread of trees, includingTilia, Fagus and Carpinus, at both regional and local level, is presented. Five major phases of intensive human activity are recognised, the most intensive activity of the prehistoric period occurring in the Neolithic (Großer Krebssee profile). Differences between the records is explained in terms of local habitat, especially edaphic conditions, settlement history and also the pollen source area, the profile from the much larger Felchowsee lake providing a record of environmental change that is more regional in character.     

Holocene vegetation history of the Sahel: pollen, sedimentological and geochemical data from Jikariya Lake, north-eastern Nigeria

Aim This study aims to separate regional and local controls on Holocene vegetation development and examine how well pollen records reflect climate change in a semi‐arid region. The relative importance of climate and human activity as agents of vegetation change in the Sahel during the late Holocene is also considered. Location Jikariya Lake, an inter‐dune depression in the Manga Grasslands of north‐eastern Nigeria. Methods Pollen and charcoal were used to provide a record of Holocene vegetation history. Palaeoclimate and hydrological changes were reconstructed from sedimentary and geochemical data. Regional and local influences were separated by comparing the evidence obtained from Jikariya Lake with previously published data from the Manga Grasslands. Results The Manga Grasslands experienced a prolonged wet period during the early and mid‐Holocene, during which swamp forest vegetation with Guinean affinities (Alchornea, Syzygium, Uapaca) occupied the inter‐dune depressions. However, variation in the pollen records between sites suggests that their establishment was dependent on conditions being locally favourable, rather than being directly coupled to regional climate. The pollen records from the Manga Grasslands are more consistent in suggesting the colonization of the dunefields by trees associated with Sudanian savanna (Combretaceae, Detarium) c. 8700 cal. yr bp . The Jikariya Lake pollen data are in accordance with the sedimentological and geochemical data from the region in indicating that the onset of arid conditions occurred progressively during the late Holocene (from c. 4700 cal. yr bp ). Abrupt changes in pollen stratigraphy, recorded at other Manga Grasslands sites 3500 cal. yr bp , appear to be the product of the local passing of ecological thresholds. The dunefield vegetation (Sahelian savanna) appears to have been resilient to (or at least palynologically silent regarding) to the climatic variability of the late Holocene. Main conclusions While climate appears to have been the primary control on vegetation development in the Manga Grasslands during the Holocene, local conditions (particularly depression size and sand influx) had a strong influence on the timing of pollen stratigraphic changes. Anthropogenic influences are difficult to detect, even during the late Holocene.     

Plant subsistence and environment at the Mesolithic site T?gerup, southern Sweden: new insights on the “Nut Age”

Pollen was analysed from a sediment sequence collected in the close vicinity of the Mesolithic settlement T?gerup, southern Sweden. Macroremains were also retrieved from numerous samples taken at the site of the archaeological excavations of Kongemose and Erteb?lle settlement phases, 6700–6000 b.c. and 5500–4900 b.c. respectively. Plants and other organic remains were well preserved in the refuse layers from the settlements embedded in the gyttja. The pollen record includes no clear indications of human impact on the vegetation during the Mesolithic. The occurrence of charcoal particles and pollen of grass and herbs associated with nutrient-rich soils are contemporaneous with the Kongemose settlement. The Erteb?lle settlement phase, although characterised by considerable dwelling activities less than a hundred metres from the pollen sampling site, is scarcely seen in the pollen data. Numerous finds of crushed dogwood stones from the Kongemose phase, often partly carbonised, suggest that these stones were used for the extraction of oil. Other plants found in the Kongemose refuse layers that may have been used are apples, cherries, raspberries, acorns and rowan-berries. Based on the abundance of hazelnut shells found at the studied site and in other studies of Mesolithic sites in southern Scandinavia it is proposed that these remains may testify to an important food supply rather than just use as a supplement to animal protein. It is also hypothesised that a regional decrease in hazel populations and thus hazelnut availability at the end of the Mesolithic may have motivated the adoption of Neolithic subsistence.     

Settlement and landscape history at the Federsee,south-west Germany,as reflected in pollen diagrams

The Federsee mire in the Alpine Foreland of south-western Germany contains a record of a remarkable archaeological landscape. Since the first excavations in the 1920's, botanists and mire geologists have studied the relationship between landscape development and settlement at this site. In a new study, funded by the Deutsche Forschungsgemeinschaft, various disciplines embracing both archaeology and the natural sciences have come together to address outstanding questions and problems. Pollen analysis can only be carried out within the Federsee mire since no other suitable mires are found in the vicinity. Because of the size of the Federsee basin (30 km² at the end of the last glaciation), the regional pollen component, consisting predominantly of arboreal pollen, prevails over the herbaceous component which mainly reflects activity associated with settlements. Nevertheless, phases of settlement are clearly reflected in the radiocarbon-dated pollen diagrams and can be correlated with Neolithic and Bronze Age settlements that are dated either by dendrochronology or radiocarbon. In addition, some settlement phases were identified for which no archaeological evidence is yet available. As a consequence of human impact during the Atlantic and Subboreal periods, a gradual opening-up and change in structure of the forests is recorded. There is evidence for an exceptionally high level of human impact associated with two Bronze Age settlements that were present in the central part of the Federsee mire. Each of the five transgressions of the Federsee so far identified occurred at the end of a settlement phase. These may have resulted from anthropogenic activity rather than climatic change. A contribution to the 8th IPC, Aix-en-Provence, Sept. 1992     

The presence of the brown bear Ursus arctos in Holocene Britain: a review of the evidence

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Late Mesolithic environmental change at Black Heath, south Pennines, UK: a test of Mesolithic woodland management models using pollen, charcoal and non-pollen palynomorph data

The recognition of Mesolithic impacts in mid Holocene pollen diagrams of the British Isles has led to the development of models describing sophisticated woodland management, particularly through the use of fire, by Mesolithic populations. However, the significance of human agency in creating mid Holocene woodland disturbances is unclear, with natural and human-induced clearings arguably indistinguishable in the pollen record. Analysis of non-pollen palynomorphs (NPPs) should aid the identification of events and processes occurring within these woodland disturbances and provide more precise palaeoecological data. In this paper we present pollen, charcoal and NPP analyses from a potentially critical location in the Mesolithic impacts debate. NPP types aid significantly in the reconstructions, suggesting periods of dead wood, grazing, local burning and wetter ground conditions. The results indicate that between 7700 and 6800 cal b.p., a predominantly wooded environment periodically gave way to phases of more open woodland, with inconsistent evidence for animal grazing. From 6800 cal b.p., a phase of open woodland associated with high charcoal concentrations and indicators of grazing was observed. This probably represents the deliberate firing of vegetation to improve grazing and browse resources, although it remains unclear whether fire was responsible for initially creating the woodland opening, or if it was part of an opportunistic use of naturally occurring woodland clearings.     

Multi-profile fine-resolution palynological and micro-charcoal analyses at Esklets,North York Moors,UK, with special reference to the Mesolithic-Neolithic transition

Multi-proxy palaeoecological data from two peat profiles at Esklets on the North York Moors upland provide a record of vegetation changes for much of the Holocene. Possible vegetation disturbance in the late Mesolithic and activity in the Neolithic and Bronze Age are recognised. In both profiles fine resolution analyses have been applied to the period leading up to the mid-Holocene Elm Decline which in this upland has been dated to ca. 4,800 bp (uncalibrated ¹⁴C years). Disturbance impacts at the Esklets Elm Decline are low scale, but phases of woodland disturbance, which include cereal (Hordeum)-type pollen, occur in both profiles ca. 5,200 bp, some centuries before the Elm Decline on the North York Moors, but similar to dates for this key palynological horizon in nearby lowland areas. A protocol is presented for the separation of Hordeum (cultivated species) and Glyceria (wild grass) pollen. The Esklets sites record disturbances during the late Mesolithic-Neolithic transition. These pre-Elm Decline disturbance phases represent either early penetration of neolithic cultivator-pastoralists into this upland or the activities of final mesolithic foragers. No neolithic archaeological sites occur nearby, but a ‘Terminal Mesolithic’ flint site dominated by microlith ‘rod’ forms occurs close to the palaeoecological sites. Such rod sites are dated in northern England to the centuries leading up to 5,000 bp and so are contemporary with the disturbance phases that included Hordeum-type pollen at Esklets. The cultural context of these disturbance phases and the role of ‘rod’ microlith sites during the Mesolithic-Neolithic transition require further focused research to clarify all issues relating to this important period.     

Vegetation changes and human occupation in the Patagonian steppe, Argentina, during the late Holocene

Vegetation changes during the late Holocene are interpreted from four fossil pollen sequences from two caves at the Los Toldos archaeological locality, Santa Cruz province, Argentina. Taphonomic processes are particularly taken into account in order to analyze the effects on the fossil pollen records of biotic factors such as human occupation and animals, and abiotic ones such as volcanic ash fall. Fossil pollen assemblages are interpreted using local modern pollen data. The main vegetation change occurred at ca. 3750 uncal b.p., when a shrub steppe of Asteraceae subf. Asteroideae with Schinus, Ephedra frustillata and a high proportion of grasses was replaced by a shrub steppe of Colliguaja integerrima and Asteraceae subf. Asteroideae. This change is synchronous with an archaeological record change and could be related either to moderate climatic variations or the effects of ash fall on the environment. Plant communities similar to the present-day ones were established in the Los Toldos area from ca. 3750 uncal b.p.     

New evidence of possible crop introduction to north-eastern Europe during the Stone Age

Long term (from the Mesolithic to the Bronze Age) habitation of the Akali settlement on a clearly defined bog-island in East Estonia is used as an example of transitional development from a prosperous foragers’ habitation centre to a hinterland of established farming cultures, taking place through availability, substitution and consolidation phases of crop farming in the boreal forest zone. The pre-Neolithic finds of Triticum and Cannabis t. pollen at c. 5600 b.c. are interpreted as possible indications of the acquaintance of foragers with farming products, through contacts with central European agrarian tribes during the availability phase. The substitution phase is marked by more or less scattered pollen finds of various cereals and hemp and, at Akali, is connected with Neolithic period 4900–1800 b.c. An increasing importance of crop farming in the economy is characteristic of the consolidation phase, but because natural conditions are unfavourable for arable land-use, a regression of human presence is recorded during the second part of the Neolithic. The settlement was abandoned during the Bronze Age at the time when crop farming become the basis of the economy in Estonia. The re-colonisation of the area, traced to ca. a.d. 1200, took place for political reasons rather than through increasing suitability of the landscape.Editorial responsibility: Felix Bittmann     

Early and Middle Holocene Hunter-Gatherer Occupations in Western Amazonia: The Hidden Shell Middens

We report on previously unknown early archaeological sites in the Bolivian lowlands, demonstrating for the first time early and middle Holocene human presence in western Amazonia. Multidisciplinary research in forest islands situated in seasonally-inundated savannahs has revealed stratified shell middens produced by human foragers as early as 10,000 years ago, making them the oldest archaeological sites in the region. The absence of stone resources and partial burial by recent alluvial sediments has meant that these kinds of deposits have, until now, remained unidentified. We conducted core sampling, archaeological excavations and an interdisciplinary study of the stratigraphy and recovered materials from three shell midden mounds. Based on multiple lines of evidence, including radiocarbon dating, sedimentary proxies (elements, steroids and black carbon), micromorphology and faunal analysis, we demonstrate the anthropogenic origin and antiquity of these sites. In a tropical and geomorphologically active landscape often considered challenging both for early human occupation and for the preservation of hunter-gatherer sites, the newly discovered shell middens provide evidence for early to middle Holocene occupation and illustrate the potential for identifying and interpreting early open-air archaeological sites in western Amazonia. The existence of early hunter-gatherer sites in the Bolivian lowlands sheds new light on the region’s past and offers a new context within which the late Holocene “Earthmovers” of the Llanos de Moxos could have emerged.     

Traces of human impact reflected in the pollen diagram from Tarnowiec mire near Jasło (Jasło-Sanok depression), SE Poland

The pollen diagram from Tarnowiec concentrates on human impact on vegetation, and is supported by the archaeological background. The earliest evidence of human activities dated at ca. 6500 B.P. was probably connected with the Neolithic Linearbandkeramik culture. Further periods of settlement are recorded at ca. 5200 B.P. and at 4800 B.P. The next very distinct period of intensification in human impact at ca. 4200-3800 B.P. was most certainly connected with the presence of the Corded Ware culture. The next period of settlement occurred during the late Bronze Age, in connection with the development of the Lusatian culture from ca. 3200 B.P. but is recorded only faintly in the pollen diagram. The next colonization phase at ca. 2600 B.P. is possibly the result of Scythian invasions. The widest expansion of human impact came during the period of Roman influence. After this, anthropogenic indicators decreased during the Migration period. The youngest part of the pollen diagram reveals an intensification of anthropogenic indicators connected with early Medieval settlement.     

From foraging to farming in the Great Mazurian Lake District: palynological studies on Lake Miłkowskie sediments, northeast Poland

Early and mid Holocene local vegetational history, with special reference to woodland communities, was revealed by pollen analysis of a radiocarbon dated lake sediment profile from Lake Miłkowskie (Jezioro Miłkowskie) in northeastern Poland. The main factor controlling the dynamics of woodland composition changes until ca. 1950 b.c. was climate. After that, the role of human activity became increasingly important. The results of high-resolution pollen analyses provide evidence for early woodland disturbances caused by Mesolithic people at ca. 6950 b.c. Several episodes of human impact, differing in scale, and separated by subsequent episodes of woodland regeneration/stabilization were noted. The first traces of local crop farming, shown by the presence of Cerealia pollen, were recorded at ca. 3800 b.c. in the Paraneolithic/Neolithic period. Animal husbandry as well as cereal cultivation played only a marginal role in the economy, which was traditionally based on hunting, fishing and gathering through the Neolithic and the early Bronze Age. The change in economic strategies from foraging towards farming, starting around 3750 b.c., was a long-lasting process. An increase of productive economy took place in the middle Bronze Age at ca. 1400 b.c.     

Earliest Holocene vegetation history and island biogeography of the Isle of Man, British Isles

Aim To present radiocarbon dated early Holocene pollen analytical data from two sites on the northern plain of the Isle of Man and to discuss the implications of the vegetation history in relation to severance of the island from the British Isles and to identify further evidence for divergent biogeographical development previously exemplified by the survival and apparent dwarfism of late glacial Megaloceros giganteus (Giant Deer). Location The Isle of Man, British Isles. Methods Pollen analysis and AMS radiocarbon dating of late glacial to early Holocene lake sequences at Pollies and Curragh‐y‐Cowle on the northern plain of the Isle of Man. Results The pollen data indicate a prolonged period of pre‐woodland vegetation after the late Glacial/Holocene transition, which lasted for most of the first post‐glacial millennium. This persistence of pre‐forest environments meant that the expansion of Betula woodland occurred later in this part of the Isle of Man than in adjacent areas of Britain and Ireland. Conclusions The Isle of Man, in the northern Irish Sea, was isolated from Britain during the late Glacial period perhaps explaining the delayed arrival of tree species. Delayed rise of the Holocene forest compared with surrounding regions probably reflects severance of the land‐bridge with Cumbria, but also could be a function of climate changes during the early Holocene and local environmental conditions. Late survival and the dwarfism of the Megaloceros giganteus (Giant Deer) fauna is another example of biogeographical divergence during the early Holocene/late Glacial of the Isle of Man. The delayed afforestation and absence of human hunters in the Manx early Holocene offers a permissive environmental context for the as yet unproven survival of Megaloceros into the early Holocene.     

Reconstruction of environment and history of plant use during the late Mesolithic (Ertebølle culture) at the inland settlement of Bökeberg III,southern Sweden

Pollen, plant macrofossils and charcoal were analysed from a lake-sediment sequence, including a refuse layer, from the Late Mesolithic settlement at Bökeberg III, southern Sweden. The chronology was established by means of AMS-dated plant macroremains. The results of the biostratigraphical studies indicate two settlement phases (A and B), at ca. 6650-6400 B.P. (5560-5320 cal. B.C.) and ca. 6150-5800 B.P. (5200-4680 cal. B.C.), respectively. The two settlement phases are associated with periods of low lake-level contemporaneous with the second major period of low lake levels during the Holocene in southern Sweden, and thus with a period of generally drier climate. The pollen analytical data suggest only minor human impact on the local vegetation during the two settlement phases. Three elm declines at ca. (1) 6200 B.P. (5200-5100 cal. B.C.), (2) 5450 B.P. (4340 cal. B.C.), and (3) 5150 B.P. (3980 cal. B.C.) are discussed. Elm decline 3 is synchronous with the classical north-west European elm decline. Elm declines 2 and 3 may be due to outbreaks of elm disease rather than to strong human impact or climate change. The charcoal analyses show that wood of a wide range of species was collected for fuel or other purposes. During phase A, plants used included acorns, hazelnuts and, possibly, Cornus sanguinea, and also Prunus spinosa, Sorbus aucuparia and Rubus idaeus. There is convincing evidence that Cladium mariscus was used for thatching. The second occupation phase, B, is characterised by the use of hazelnuts for food. The possible use of several other identified species is discussed.