Two new European species of <Emphasis Type="Italic">Docosia</Emphasis> (Diptera: Mycetophilidae)

Two new species of Docosia Winnertz, 1863 are described, Docosia chandleri sp. n. from Austria and Italy and Docosia matilei sp. n. from France and Italy. Both species resemble Docosia montana Laštovka et Ševčík, 2006 and they are characterized by bare laterotergites, yellow legs and long flagellomeres.     

First fungus gnats of genus Manota Williston (Diptera: Mycetophilidae) from Japan

The genus Manota is recorded from Japan for the first time. Three new species, Manota satoyamanis, Manota indahae and Manota tunoae spp. nov., are described, based on specimens collected in an ecological sampling program of arthropods in the “satoyama” landscape of Ishikawa Prefecture. “Satoyama” represents the traditional rural landscape of Japan, which is characterized by a mosaic of secondary forests, plantations, ponds and rice paddy fields. The new species raise the number of Palearctic Manota species from five to eight.     

Molecular phylogeny of the Paleogene fungus gnat tribe Exechiini (Diptera: Mycetophilidae) revisited: Monophyly of genera established and rapid radiation confirmed

The phylogeny of the fungus gnat tribe Exechiini (Diptera: Mycetophilidae) is reconstructed based on the combined analysis of five nuclear (18S, two parts of 28S, CAD, EF1α) and two mitochondrial (12S, COI) gene markers. According to known fossil record, and recent higher‐level phylogenies, the tribe constitutes the most apomorphic, distinctly monophyletic clade of the family Mycetophilidae. The tribe originated in the Paleogene and apparently quickly diversified in the Neogene with an unusual rapid radiation of complex male terminalia. Earlier attempts to reconstruct the phylogeny of the tribe, based on both morphology and molecular methods, have not yielded reliable hypotheses, neither in terms of resolution nor in terms of support for major clades. Increased taxon sampling and wider gene sampling have been suggested to achieve better phylogenetic resolution. Aiming at this, we present new phylogenies, for the first time with all known genera and subgenera of Exechiini represented. While many terminal intergeneric relationships are well supported, both in maximum likelihood and in Bayesian analyses, most of the major, deeper clades remain poorly supported. We suggest that a rapid radiation event close to the root may be causing the low resolution at this level in the phylogeny. This contrasts parallel phylogenies of the older subfamilies and tribes of the family Mycetophilidae, where traditional clades have usually been recovered with high support. Further in‐depth studies into the evolutionary history of the tribe are needed to enlighten and coalesce the specific phenomena driving their unique morphological, genetic and phylogeographic histories.     

Amiota sinuata species group from eastern Malaysia (Diptera, Drosophilidae)

Eight species of the Amiota sinuata species group are reported from eastern Malaysia, including six new species, A. bispinula, A. cerata, A. curvibacula, A. lambirensis, A. parviserrata and A. quadrifoliolata spp. nov. A key to Asian species of the group is provided.     

A New Species of Chelipoda Macquart,with a Key to the Species from China (Diptera: Empididae)

Chelipoda nigraristata sp. n., a new aquatic dance fly of the subfamily Hemerodromiinae (Diptera, Empididae) is described from China. Relationships with other described species are discussed. A key to the species of the genus Chelipoda from China is presented for the first time.     

Six new species of Acomoptera from North America (Diptera, Mycetophilidae)

Six new species are described, raising the number of North American Acomoptera species to seven and the genus total to ten, and nearly doubling the number of species within the putative clade containing Acomoptera, Drepanocercus, and Paratinia. These novel species forms have implications for the concept of Acomoptera that in turn, may impact our understanding of its generic relationships and the evolution and composition of Gnoristinae and Sciophilinae. The new species, Acomoptera crispa, Acomoptera digitata, Acomoptera echinosa, Acomoptera forculata, Acomoptera nelsoni,and Acomoptera vockerothi, are compared with the type species of the genus, Acomoptera plexipus (Garrett), whose diagnostic features are imaged and illustrated for the first time. The European species, Acomoptera difficilis (Dziedzicki) is also illustrated and compared. Acomoptera spinistyla (Søli) comb. n. is transferred from Drepanocercus. A key to species is provided. Future work will seek to incorporate this knowledge into a systematic phylogenetic study of relationships between these species and their sister taxa.     

The genus Apsiphortica from China with DNA barcoding information (Diptera: Drosophilidae)

Two new species of the genus Apsiphortica are described from China: A. orthophallos n. sp. and A. sinuatipenis n. sp. Species delimitations are improved by integrating morphological and DNA barcoding information. The intra- and interspecific pairwise p-distances (proportional distance) are summarized for five Apsiphortica species from China. Furthermore, nucleotide sites with fixed status in the alignment of the COI sequences (639 nucleotide sites in length) are used as “pure” molecular diagnostic characters to delineate the five species. A key to all the Chinese species of the genus Apsiphortica is provided.     

Fungus gnats of the genera Anatella,Allodia and Cordyla in Japan (Diptera: Mycetophilidae)†

Six new mycetophilid species from three genera belonging to the tribe Exechiini are described: Anatella prominens on a fungus of the Polyporaceae, Allodia bipexa and Allodia conifera on fungi of the Russulaceae, Allodia quadrata on fungi of the Hygrophoraceae, Tricholomataceae, Rhodophyllaceae, Russulaceae and Fistulinaceae, Allodia laccariae on a fungus of the Tricholomataceae, and Cordyla bidenticulata on a fungus of the Boletaceae. Their characteristic male genitalia are illustrated. Cordyla flaviceps Staeger, Cordyla fusca Meigen and Cordyla pusilla Edwards are newly recorded from Japan.     

The sequoia-loving sprite,a new genus and species of fungus gnat (Diptera,Mycetophilidae) from California

California is one of the most biologically diverse regions of the world, yet the diversity of fungus gnats (Mycetophilidae) remains largely undocumented within the state. A modest survey of these flies has led to the discovery of a new genus and species of gnat that lives alongside one of the most iconic trees in the world, the giant sequoia (Sequoiadendron giganteum). Spritella sequoiaphilagen. et sp. n. is described and illustrated and its status among other mycetophilid genera is analyzed and discussed.     

Description of a new species of the genus Glenea from Tibet, China (Coleoptera, Cerambycidae, Lamiinae, Saperdini)

A new species, Glenea jinisp. n. is described from Tibet, China. It can be separated from other species of the genus Glenea Newman by the complicated black and ochre markings as well as characters of the genitalia.     

Revision of the Chinese species of the genus Coenochilus Schaum (Coleoptera,Scarabaeidae, Cetoniinae)

Summary

The Chinese species of the genus Coenochilus Schaum, 1841 are revised, including a new species, C. pilosus n. sp. described from southern China (also from Vietnam and Laos). The lectotype for Coenochilus uncinatipes Moser, 1915 is designated, and this name is recognized as a junior synonym of C. bifoveolatus Fairmaire, 1888. Previously published synonymy of Coenochilus apicalis Westwood, 1873 and C. armiger Westwood, 1873 is confirmed based on the examination of type specimens. The neotype of C. tonkinensis Moser, 1910 is designated, and C. clinteroceroides Jákl, 2015 is synonymized with this species. Coenochilus tonkinensis is newly recorded from China, while Chinese records of C. nitidus Arrow, 1910 are verified as misidentification of C. striatus Westwood, 1873, thus C. nitidus is removed from the Chinese fauna. Diagnoses and illustrations are provided for all five Chinese species. A key to Chinese species and new distribution records are also presented.??     

Review of the Palaearctic Acomopterella Zaitzev (Diptera,Sciaroidea, Mycetophilidae)

The distribution of Acomopterella species in the Palaearctic region has been re-examined in this study, using recently collected material. The European species was found to be distributed in the eastern Palaearctic as well. A second Palaearctic species from Honshu (Japan) is herein described. The morphology of adult specimens was studied by light microscopy and scanning electron microscopy. The shape of functional specialized setae on mid tibiae in Acomopterella and seven further fungus gnat genera is described and the suitability of this character for systematic studies is discussed. Details of a “hind tibial organ” are described.The position of Acomopterella in the tribe Gnoristini is briefly discussed. Acomopterella is found to be more closely related to Speolepta Edwards, 1925, than to any other recent genus.     

浙江龙王山滑菌蚊三新种(双翅目：菌蚊科)

报道浙江省龙王山自然保护区滑菌蚊属Leia三新种：针尾滑菌蚊L.aculeolusa sp.nov.,长尾滑菌蚊L.ampulliforma sp.nov.和龙王山滑菌蚊L. longwangshana sp.nov.。模式标本保存在中国农业大学昆虫标本馆。     

Taxonomic study on the Phytosciara genus group (Diptera: Sciaridae) in Korea,including the description of a new species

The Phytosciara genus group (Diptera, Sciaridae: Phytosciara Frey, Dolichosciara Tuomikoski, and Prosciara Frey) are studied in the Korean peninsula. Dolichosciara koreaensis Shin & Menzel sp. n. is described and illustrated as new to science. Additionally, four species are reported as new to Korea: Dolichosciara orcina (Tuomikoski, 1960), Dolichosciara semiferruginea (Menzel, 1995), Prosciara producta (Tuomikoski, 1960), and Prosciara ussuriensis (Antonova, 1977). Identification keys are given for the genera of the Phytosciara genus group and for the Korean species of the genera Dolichosciara Tuomikoski and Prosciara Frey. The COI barcode for the new species is studied.     

浙江龙王山滑菌蚊三新种 (双翅目：菌蚊科)

 报道浙江省龙王山自然保护区滑菌蚊属Leia三新种： 针尾滑菌蚊L.aculeolusa sp.nov.,长尾滑菌蚊L.ampulliforma sp.nov.和龙王山 滑菌蚊L.longwangshana sp.nov.。模式标本保存在中国农业大学昆虫标本馆。     

The Megophthalmidia (Diptera,Mycetophilidae) of North America including eight new species

Megophthalmidia Dziedzicki is a small leiine genus (Mycetophilidae) with seven species described from the Neotropics and ten species from the Palearctic region. Two species of Megophthalmidia have been reported for North America. Recent collecting of Mycetophilidae in California and Arizona, however, shows current North American diversity of Megophthalmidia is at least on par to other regions of the world. Eight new species of Megophthalmidia are described here, increasing the number of Nearctic Megophthalmidia species to nine. Included is a particularly atypical member of the genus, M. saskia sp. n., which expands the genus concept of Megophthalmidia. Of the two species previously recorded for North America, only one actually belongs in the genus. Megophthalmidia occidentalis Johannsen, is fully described and illustrated. The other named species, M. marceda (Sherman) is illustrated and transferred to the genus Ectrepesthoneura Enderlein. A lectotype is designated for this species. A key to the species of Megophthalmidia of North America is provided. The biology of these flies is not yet known. Three of the new Megophthalmidia species – M. lenimenta, M. misericordia, and M. radiata – are only known to occur within small protected areas within the California State Park and UC Natural Reserve systems.