Ostracodes et incursions marines dans l’Holocène terminal du lac Retba (Cap Vert, Sénégal)

The study of marine incursions based on ostracods is carried out for the first time in the late Holocene, with core sediments of the Lac Retba edge. Twenty-four species are identified along the core; most of them are encountered in the Gulf of Guinea shelf and its adjacent lagoons. For the first time Buntonia olokundudui Reyment and Van Valen, Hermanites foveolata Omatsola and Soudanella africana Omatsola are recorded in Senegal. Four ostracods assemblages are evidenced along the core, corresponding to different stages of the lake evolution. Cyprideis nigeriensis Omatsola, Neomonoceratina iddoensis Omatsola and Aglaiocypris gambiensis Witte dominate the first assemblage; they indicate a lagoon slightly open to the sea and bordered with mangrove swamps under relatively humid climate conditions. The second association yields the most diversified ostracod fauna with C. nigeriensis, N. iddoensis and high proportion of reworked “marine species” which evidence a marine little gulf bordered by mangrove under a humid climate. The third, monospecific association (C. nigeriensis) is typical of a closed and hypersaline lagoon under dry climate conditions. An oligospecific association with C. nigeriensis, Pseudoconcha omatsolai Witte and very scarce reworked “marine species” occurs at the top of the core and characterises an open and hypersaline lagoon under arid climate conditions. Ostracods evidence two marine incursions. The first and the stronger one is dated here for the first time between 1420 and 1250 years B.P. The more recent is the weaker and is not yet dated. Evidence of the beginning of the Retba Lake closure towards 680 years B.P. appears at the top of the core, as indicated by high reduction of ostracods abundance, predominance of reticulate Cyprideis and disappearance of the “marine species”, while salts precipitated at the bottom of the lake.     

Évolution du trait de côte à l’Holocène supérieur dans la Sebkha El-Guettiate de Skhira (Golfe de Gabès, Tunisie) à travers sa faune d’ostracodes et de foraminifères

A 122 cm-long core was taken in the El-Guettiate Sebkha of Skhira (southeastern part of Tunisia) in order to investigate the recent palaeoenvironmental evolution of this region. The quantitative and qualitative analysis of ostracod and benthic foraminifera assemblages coupled with a correspondence analysis allows the reconstruction of palaeoenvironmental changes during the Holocene in this area. Four typical associations of ostracods (open marine, coastal marine, lagoonal and estuarine brackish) and two associations of benthic foraminifera (coastal and lagoonal) were distinguished. The onset of restricted lagoonal environments linked to the building-up of sand spit led to the onset of restricted lagoon and brackish environment at cal. 5408 years BP. The Shannon and equitability index of diversity were used to decipher the structural variations of the populations of ostracods and benthic foraminifera along the sampled core. We note a reduction in the Shannon index from the bottom to the top, which indicates a progressive isolation of the biotope. The open lagoonal episode is characterized by high values of diversity. During restricted lagoonal episodes the Shannon and equitability index are reduced. The correspondence analysis reveals an environmental gradient related to the marine influence. It shows an antagonism between the widely opened estuarian lagoonal species and those of restricted lagoon. The less opened estuarian lagoonal taxa occupy an intermediate position. Based on these evidences, microfauna carried out in the El-Guettiate Sebkha allows us to recognize four phases beginning with a widely opened estuarian lagoon (ca. cal. 7460 years PB), followed by a restricted lagoon (ca. cal. 5408 years BP) and finally a brackish lagoon evolving towards the present-day sebkha environment. The opened estuarine lagoon is characterized by high values of species richness and diversity indices.     

Des Miliolacea aux Alveolinacea (grands foraminifères): la faune du Crétacé supérieur et du Tertiaire inférieur du Chiapas (S.E. Mexique)

Found in Chiapas (South-East of Mexico), Praealveolina michaudi nov. sp. and Chubbina jamaicensis (of Campanian-Maestrichtian age) are described, both evolved from Nummoloculina sp. while Raadshoovenia guatemalensis (upper Paleocene-lower Eocene) arises from Quinqueloculina. We precisely analyze and modelize the nepionic coiling during the evolutionary change from Miliolacea to Alveolinacea. During ontogenesis as well as during phylogenesis, quinqueloculine coiling progresses to streptospiral and then to planispiral chamber arrangement with the increase of volume of embryo and of chambers. Appearance of these large foraminifers are understood as a consequence of the rise of K-strategies in their mode of life, following the development of photosynthetic symbioses.     

Les foraminifères du Burdigalien moyen à supérieur de la région sud-aquitaine (golfe de Saubrigues, SW France)

Many exposures assigned to the Miocene formation of “Marnes de Saubrigues” (southwestern Aquitaine Basin) have been studied. Their revised stratigraphical allocation has made it possible to recognize mainly Middle to Late Burdigalian age deposits which have been widely sampled. They have been correlated with planktonic zones N6 pars-N7 and NN3-NN4 and locally dated 17.7 Ma with Sr isotopes. It is noteworthy that this chronological interval is only known in the southern Basin. An inventory of the rather abundant microplankton is provided, including series crossed by drillings in the Gulf of Biscay. These deposits yield a large diversity of benthic foraminifera, nearly 400 taxa, which can be considered as a reference fauna for western Europe Burdigalian. From a geographic point of view, it has been possible to subdivise the exposures into groups according to their bathymetry. Let us point out the first discovery of Borelis in the Burdigalian of France and in the northeastern Atlantic Miocene as well. Very rare specimens of Cycloclypeus are also present in the Aquitaine Basin. Among smaller foraminifera, the occurrence of Rosalina aguayoi, Pavonitina styriaca, the last appearance of Falsocibicides aquitanicus and the persistance of the genus Almaena must be emphasized. Several assemblages supply diatoms, rare radiolaria and sometimes abundant sponge spicules; the presence of these organisms, seldom reported from the Aquitaine Tertiary, might be linked to upwelling-type currents. Bachmayerella (inc. sed.) is uncommon and cited for the first time in the Burdigalian. A few taxonomical remarks and short notes are given for some species of foraminifera, and palaeoecological details as well. Numerous taxa are represented and comparisons with the foraminifera faunas mainly from the Lower-Middle Miocene of Western Mediterranean and Paratethys are sketched.     

Les grands mammifères du Villafranchien supérieur des travertins du Bassin de Denizli (Sud-Ouest Anatolie,Turquie)

The large mammals from travertine deposits in the Denizli basin include the following species: Archidiskodon meridionalis meridionalis, Equus cf. altidens s. l., E. cf. apolloniensis, Stephanorhinus cf. etruscus, Metacervoceros rhenanus, Cervalces (Libralces) ex gr. minor-gallicus, Palaeotragus sp., Bovinae gen. and sp. indet. This association resembles those from the late Villafranchian of Southern and Eastern Europe, and, to some extent, fromWestern Asia, and could be older than 1.2 Ma.     

Un nouveau Bovini dans la faune du Pléistocène supérieur d’Algérie

During the study of the Quaternary Maghrebian bovines in 1985, the review of the Bovini (aurochs and buffalos) found in Upper Pleistocene sites of Algeria showed that in comparison with most of the African bovine assemblage the attribution of genus Pelorovis to the species antiquus was inappropriate. The biometric and cladistic data revealed a close kinship with genus Syncerus. Later, other authors’conclusions confirmed its relevance. The antique buffalo of the Aterian site of the Phacochoeri has all the characteristics of genus Syncerus, but is different in that it shows a certain number of specific characteristics described in this article.     

Le bassin de l’homme du Pléistocène supérieur de Liujiang, Sud de la Chine : taille du corps, forme du corps et encéphalisation

The Late Pleistocene hominin fossil assemblage from Liujiang, South China include a fairly well-preserved cranium, a right os coxa, a complete sacrum, and other postcranial elements all belonging to a single individual. This rare discovery offers us a unique and singular opportunity in understanding this Late Pleistocene hominin's body proportion and relative cranial capacity (encephalization quotient [EQ]), and also pelvic morphology. Using the available right innominate and its mirror-imaged left side, we reconstruct Liujiang hominin's pelvis. Our analysis of the pelvis indicates that the Liujiang hominin has a very gracile and modern-like pelvic morphology. Indeed, all of the pelvic dimensions are smaller than those of other Pleistocene hominins. Moreover, the pelvic characteristics typical of Middle and Late Pleistocene hominins including Neanderthals cannot be identified in the Liujiang pelvis. In contrast, both Liujiang's metric and non-metric features indicate affinities to more recent human populations including our modern Chinese collections from Guangxi of south China. Further support of this assessment comes from the EQ value of 5.754 for Liujiang which is closer to Minatogawa 2 and modern Chinese populations than to Middle and Late Pleistocene fossil hominins. Our analysis of body shape shows that Liujiang has body proportion (i.e. body height relative to body breadth) typical of warm-adapted populations. Based on these findings, we reason that the modern physical characteristics of Liujiang may allude to a more recent geological age. Alternatively, its morphological “modernity” could also point to a much higher degree of skeletal variation within Late Pleistocene hominins in East Asia.     

La transgression thanétienne (Paléocène supérieur) dans l’Aurès occidental (Algérie), d’après les associations de Foraminifères de la coupe d’El Kantara

In the W-Aures (Algeria), the El Kantara pass displays about 50 m of Red Marls overlying the Maastrichtian limestones with Laffitteina. The Red Marls are, in their turn, overlain by the Thanetian marly limestones of the river El Haï. The micropaleontological inventory of these Red Marls establishes the co-occurrence of subaerian (Microcodium = Paronipora), fresh-water (Charophyta) and marine microfossils (Foraminifers). The stratigraphical significance of these microfossils is discussed. The upper part of the Red Marls, yielding abundant Valvulina and scarce Glomalveolina, is of Thanetian age. The age of the lower part is less established, but the occurrence of Microcodium suggests a Thanetian age too. Located at the margin of the northern opening of the trans-Saharan epeiric sea, the El Kantara section establishes a Thanetian age for the beginning of the Paleocene transgression. Danian deposits are missing, in spite of the absence of an obvious hiatus on the field. After general emersion during the early Paleocene, the Thanetian transgression starts with the set up of lakes at the depositional area of the Red Marls. Marine influences, limited at first, become gradually prevailing, and end with the deposition of neritic marly limestones of river El Haï.     

Biostratigraphie d’après les foraminifères et paléoenvironnements des séries post-Éocène du sondage ASHTART 28, golfe de Gabès (Tunisie)

The Cenozoic sequence of Ashtart 28 well drilled in the Gulf of Gabes (Tunisia) is the subject of a biostratigraphical study. The samples recovered in cuttings from 390 m and downwards allowed to recognize, above the Late Eocene sediments, a sedimentary series, lithologically diversified, nearly 1600 m thick. Marine Pliocene deposits, generally attesting a low bathymetry, lie unconformably above the Messinian (Oued Bel Khedim formation), which shows the usual features of the Mediterranean confinement. The underlying Messinian pre-evaporitic platform series (Melqart formation), that is over 250 m thick, is typical of a perireefal environment. The sediments assigned to the Tortonian (Somâa Sands formation) are continental and occur unconformably above the approximately 500-metres-thick Middle Miocene strata (Saouaf, Mahmoud, Aïn Grab and Salammbô pars formations). The marine Lower Miocene and Oligocene sediments (Salammbô pars and Ketatna formations), that are more than 300 m thick, lie in continuity under the Middle Miocene. The infralittoral Chattian sequence has especially supplied a diversified assemblage of larger foraminifera recovered in other west-mediterranean basins. Datings were obtained based on planktonic and larger benthic foraminifera (Miogypsinidae, Nummulitidae, Lepidocyclinidae) and by correlations obtained by means of well loggings and lithostratigraphy. Benthic foraminifera, mainly listed for the Miocene and Oligocene, are studied from a systematic, stratigraphic and paleogeographic point of view. The paleoenvironments of deposits are defined for each considered stratigraphic interval. Comparisons are sketched with other drillings of the Gulf of Gabes. Thanks to the numerous data obtained by this detailed study, the Ashtart drilling can serve as a reference for the Tertiary sequence of this part of the Mediterranean domain.     

L’industrie du site néandertalien de la grotte de Vindija (Croatie) : une révision des matières premières lithiques

The new analysis of the Middle Paleolithic industry from the cave Vindija (Croatia), showed a necessity of revision of the previous obtained data especially considering the use and origin of the raw materials (Kurtanjek and Marci, 1990). The results presented in this study pointed out some new aspects of interpretation. First, the significance of quartz in the Middle Paleolithic was underestimated. At the same time it is obvious that the major change in the use of the raw material was at the beginning of the Upper Paleolithic and of the Epigravettian, especially in the use of chert. The second change is registered in the use of chert between Middle and Upper Paleolithic: the chalky nodules of chert are more frequent in the Upper Paleolithic then the river pebbles used in the production of Middle Paleolithic artefacts. Still, the origin of this new raw material is at the moment unknown.     

Identification des zones biostratigraphiques méditerranéennes dans le Pliocène du Cap-Bon (Tunisie)

The Nabeul-Hammamet series analysis allows to recognize the six biozona known in the Mediterranean realm, as specified by Cita (1975a). These are: the Sphaeroidinellopsis subdehiscens BANNER & BLOW acme biozone equivalent to MPL1; the Globorotalia margaritae margaritae BOLLI & BERMUDEZ interval zone (= MPL2); the Globorotalia margaritae BOLLI & BERMUDEZ, Globorotalia puncticulata (DESHAYES), concurrent range zone (= MPL3); the Sphaeroidinellopsis subdehiscens BANNER & BLOW interval zone (= MPL4); the Globigerinoides elongatus (D’ORBIGNY) interval zone (= MPL5); the Globorotalia inflata (D’ORBIGNY) interval zone (= MPL6). There is no similarity between these biozones and those established by Berggren and al. (1995) for the atlantic and the indo-pacific realms. The MPL1, MPL2, MPL3 and MPL4 zones are especially englobed in argillaceous and yellow sandy lithological units specified in north-eastern Tunisia, such as ‘Argiles des Potiers’, ‘Sables Jaunes de Nabeul’ and ‘Argiles de Sidi Barka’. These units are together equivalent to the Raf-Raf Formation developed towards the North, in the Bizerte area. Besides, the MPL5 and the MPL6 zones are included in sands and sandstones composing the Hammamet unit, equivalent to the Porto-Farina Formation.     

La Grotte des Félins, site paléolithique du Pléistocène supérieur à Dar Bouazza (Maroc)

In the continental sediments filling a cave that developed in Middle Pleistocene deposits near Casablanca, Morocco, a very rich mammal fauna from the last climatic cycle was discovered. It was associated with a Middle Palaeolithic poor and atypical lithic assemblage manufactured on blocks and pebbles. The lower fossiliferous layer contained abundant carbonized remains of felids. Unfortunately, the site was destroyed by quarrying before adequate excavations could be carried out.     

Les Schackoïnes (Foraminifères planctoniques) de l’Albien supérieur du Blake Nose (marge atlantique de la Floride, Leg ODP 171B)

A study of rather rich and well-preserved material from Upper Albian levels of Site 1050 of ODP Leg 171B led us to review the criteria used in the subdivision of the schackoinids into two genera and to describe a new morphotype, Schackoina leckiei n. sp., the phylogenetic position of which is examined within the group.     

Biostratigraphie,paléoécologie et zones d’acmé des foraminifères planctoniques au passage Crétacé-Paléogène dans la Téthys (Tunisie et Espagne) et l’Atlantique (France)

Biostratigraphical high-resolution analyses and quantitative data confirm that deposition is continuous across the K-Pg transition in several sections in Tunisia (El Kef, stratotype section) and Spain (Agost and Caravaca sections) located in the Tethyan realm and the Bidart sections in the Atlantic realm, without any relevant hiatus. The Upper Maastrichtian assemblages of planktic foraminifera from these sections are largely dominated by small biserial heterohelicids. They are associated to common species having planispiral test (i.e. globigerinelloids), trochospiral test (i.e. hedbergellids, rugoglobigerinids globotruncanids), to rare triserial heterohelicids (i.e. guembelitriids) and trochospiral species showing tubulospines (i.e. schackoinids). Stratigraphical ranges of these diverse taxa through the late Maastrichtian in the Tethyan and Atlantic realms show very few changes in the planktonic foraminiferal assemblages and most of the species are present in the Abathomphalus mayaroensis biozone. By our high-resolution sampling and the intensive research for the A. mayaroensis index species in the uppermost Maastrichtian samples, we confirm that this species is omnipresent up to the top of the Maastrichtian. Therefore, A. mayaroensis is present in almost all samples which are late Maastrichtian in age, but this species became very scarce in the uppermost Maastrichtian samples. This scarceness could be due to a climate cooling. A sharp decrease in relative abundance of the deep dwellers species, like as Abathomphalus intermedius and A. mayaroensis as well as in other keeled globotruncanids is observed at the studied sections from the Tethyan realm (indicative of low latitude) across the latest Maastrichtian. At the K/Pg boundary, all the globotruncanids disappeared. They are considered specialists living in tropical-subtropical deep seawater habitat. At this boundary, large and ornate heterohelicids also disappeared. Therefore, all the studied sections show that about 90 % of the Maastrichtian species became extinct according to a catastrophic mass extinction pattern. Only about 10 % crossed the K/Pg boundary and survived during the earliest Danian. The minor difference in the number of disappeared taxa is related to their latitude location or environment paleodepth. The changes in the species relative abundance, observed in the successive planktic foraminiferal assemblages, make it possible to recognize the Acme-stage 0 typical of the upper Maastrichtian interval. It is characterized by the highest species richness of Globotruncanids and heterohelicids specialists of tropical to subtropical marine conditions, the Acme-stage 1 typical of the Guembelitria cretacea Zone, and in particular of the Hedbergella holmdelensis Subzone dominated by “opportunists” species belonging to Guembelitria, the Acme-stage 2 which corresponds to the Pv. eugubina Zone dominated mainly by specimens belonging to Palaeoglobigerina and Parvularugoglobigerina and the Acme-stage 3 which characterizes mainly the Ps. pseudobulloides Zone dominated by biserial species belonging to Chiloguembelina and Woodringina.     

Les stratégies de débitage dans les industries lithiques archaïques des premiers habitants de l’Europe

Barranco León and Fuente Nueva 3 sites at Orce in the south of Spain, and Vallonnet cave in the south of France, are some of the oldest sites in Europe, testimonies of the first settlements of these territories. The Orce sites, dated around 1.3-1.2 Ma, have given away numerous lithic artifacts principally issued from an important débitage activity. The beginning of flake production processes is few represented into these two consummation sites. In return, full débitage flakes are in high proportion, with generally short dimensions, in consequence of which, in some case an intense core reduction. Two percussion methods are identified: with direct percussion or on anvil. Sometimes, there was débitage on flake, taking advantage of the inferior face natural convexity of the core-flake, permitting the maximal exploitation and the economy of the best raw materials. The bipolar percussion on anvil permitted the maximal exploitation of the short dimensions supports, often in naturally cubic or sub-cubic form. A lot of the producted flakes, sometimes called “pièces esquillées”, show some scales on the distal and/or proximal transversal edges, issued from this bipolar knapping method. In the Vallonnet cave, where the occupation levels are dated around 1 Ma, the lithic industry show a lot of percussion tools and hammerstone flakes, but also some core reduction flakes and some very rares cores, especially in flint. This good raw material was chosen for the débitage, and the flakes show on superior face several anterior removals, and a butt which could reveal a striking platform preparation. At the gates of Europe, in Georgia, Dmanisi site give away an abundant lithic industry associated with faunal and human remains, dated around 1.81 to 1.77 Ma. Amid the lithic assemblages, cores are rather frequent and are characterized by a poor degree of exploitation, most often unifacial, and high frequency of cortical striking platforms. Raw materials, essentially volcanic cobbles, were from local origin, and the knapped supports were often chosen for their angular morphology adequate to begin the débitage in easier way, débitage which was essentially by direct percussion.     

La phalange distale du pouce d’Orrorin tugenensis (Miocène supérieur du Kenya)

The lion's share of articles dealing with the thumb anatomy of Plio-Pleistocene hominids has focussed on the capacity to manipulate and manufacture tools, and has largely neglected the locomotor aspects. However, in these hominids, the forelimb was still employed in locomotion. Certain of the anatomical characters classically associated with manipulation and/or fabrication of tools are already present in the Late Miocene species Orrorin tugenensis as shown by the terminal thumb phalanx BAR 1901'01. This specimen reveals crucial information suggesting that thumb morphology is not exclusively related to such tool using and manufacturing activities but reflects in a frequently bipedal creature a deeper adaptation concerning the precision grip essential for climbing and balancing, different from that of apes.     

Foraminifères benthiques complexes du Bathonien supérieur au Kimméridgien inférieur de la marge atlantique marocaine (« Bassin » d’Agadir) : stratigraphie et paléobiogéographie

The carbonate sediments of the Agadir Basin (Morocco) deposited during the Upper Bathonian-Lower Kimmeridgian p.p. interval are rich in agglutinated complex benthic foraminifera. The detailed inventory of several field-sections allows to identify and to describe these benthonic foraminifera, some of them being recorded for the first time in Morocco. They belong to the main genera Archaeosepta, Andersenolina, Praekurnubia, Pseudocyclammina, Everticyclammina, Alveosepta, Kurnubia, Parurgonina, Neokilianina, Labyrinthina and Rectocyclammina. The stratigraphic and paleobiogeographic distribution of these foraminifera seem to be related to the geodynamic events occurring on the Moroccan Atlantic margin during this time-interval. The stratigraphic distribution of the principal species is generally controlled by the occurrence of ammonites, brachiopods and of some foraminifera which are considered as good stratigraphic markers.     

La macrofaune du Pléistocène supérieur d’El Harhoura 2 (Témara, Maroc) : données préliminaires

The El Harhoura 2 cave is situated in the commune of El Harhoura in the Temara region (Morocco). The entrance faces west, towards the ocean, around 200 m away from the current shoreline and about 16 m above sea level. The stratigraphy has currently 11 levels numbered from top to bottom. Level 1 is attributed to the early Neolithic. The archaeological material from level 2 places it within the Upper Palaeolithic (Iberomaurusian). Levels 3 to 11 are assigned to the Middle Palaeolithic (Aterian). Since 1977, a surface of 37 m² has been excavated, yielding particularly abundant remains of a large mammal fauna study in this paper.     

Apport des grands mammifères de la grotte du Vallonnet (Roquebrune-Cap-Martin, Alpes-Maritimes, France) à la connaissance du cadre biochronologique de la seconde moitié du Pléistocène inférieur d'Europe

The large mammals of Vallonnet cave, found into a level contemporary of Jaramillo event, are characteristics of the second half of the Lower Pleistocene. Some differences with the faunas of Fuente Nueva 3 and Barranco Leon, in Spain, which datation is the middle of the Lower Pleistocene, show that Vallonnet is younger. On the other hand, the differences are clear with Soleilhac faunas, which datation is the beginning of the middle Pleistocene.