The millipede family Polydesmidae in Taiwan, with descriptions of five new species (Polydesmida, Diplopoda)

Polydesmidae are represented in Taiwan by seven species in two genera. Neither of the genera is endemic to Taiwan, but six of the species are, including five new: Nipponesmus minorsp. n., Epanerchodus bispinosussp. n., Epanerchodus curtigonopussp. n., Epanerchodus flagellifersp. n. and Epanerchodus pinguissp. n. In addition, the diagnosis of the hitherto enigmatic genus Nipponesmus Chamberlin & Wang, 1953 is refined vis-à-vis the especially similar, Central Asian, Siberian and Eastern European genus Schizoturanius Verhoeff, 1931, chiefly based on new material of the type-species Nipponesmus shirinensis Chamberlin & Wang, 1953; this species is adequately redescribed and represents still another Taiwanese endemic. A key to all three currently known species of Nipponesmus Chamberlin & Wang, 1953 is given. The highly speciose Central to East Asian genus Epanerchodus Attems, 1901 is represented in Taiwan by five species, all keyed, including Epanerchodus orientalis Attems, 1901, which is long known to be highly variable in Japan and found particularly polymorphous and apparently allochthonous in Taiwan. The following synonymy is formalized: Epanerchodus orientalis orientalis Attems, 1901 = Epanerchodus orientalis takakuwai Verhoeff, 1913, syn. n. The genus Usbekodesmus Lohmander, 1932 is formally synonymized with Epanerchodus Attems, 1901, syn. n., resulting in the following new formal transfers: Epanerchodus redikorzevi (Lohmander, 1932), Epanerchodus swatensis (Golovatch, 1991), Epanerchodus varius (Geoffroy & Golovatch, 2004), Epanerchodus anachoretus (Golovatch, 1986), Epanerchodus buddhis (Golovatch, 1986), Epanerchodus occultus (Golovatch, 1986), Epanerchodus sacer (Golovatch, 1987), Epanerchodus theocraticus (Golovatch, 1990) and Epanerchodus theosophicus (Golovatch, 1986), all comb. n. ex Usbekodesmus. The distributions of all seven species of Polydesmidae occurring in Taiwan are mapped and discussed.     

Revision of the Taiwanese millipede genus Chamberlinius Wang, 1956, with descriptions of two new species and a reclassification of the tribe Chamberlinini (Diplopoda, Polydesmida, Paradoxosomatidae, Paradoxosomatinae)

The millipede genus Chamberlinius is basically confined to Taiwan, with only one of the four known species presumably introduced to southern Japan. Both previously known species are redescribed, based on new material: Chamberlinius hualienensis Wang, 1956 (the type species) and Chamberlinius piceofasciatus (Gressitt, 1941), the latter being a new subjective senior synonym of Chamberlinius shengmui Wang, 1957, syn. n. Two further congeners are described as new: Chamberlinius pessior sp. n. and Chamberlinius sublaevus sp. n. The genus is re-diagnosed, all of its four species are keyed, and their distributions mapped. The tribe Chamberlinini is reclassified and, based on gonopod traits, shown to comprise the following five genera: Chamberlinius Wang, 1956, Haplogonosoma Brölemann, 1916, Riukiupeltis Verhoeff, 1939, Aponedyopus Verhoeff, 1939 and Geniculodesmus Chen, Golovatch and Chang, 2008.     

Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)

The large genus Orthomorpha is rediagnosed and is shown to currently comprise 51 identifiable species ranging from northern Myanmar and Thailand in the Northwest to Lombok Island, Indonesia in the Southeast. Of them, 20 species have been revised and/or abundantly illustrated, based on a restudy of mostly type material; further 12 species are described as new: Orthomorpha atypicasp. n., Orthomorpha communissp. n., Orthomorpha isarankuraisp. n., Orthomorpha picturatasp. n., Orthomorpha similanensissp. n., Orthomorpha suberectasp. n., Orthomorpha tuberculiferasp. n.,Orthomorpha subtuberculiferasp. n. and Orthomorpha latitergasp. n., all from Thailand, as well as Orthomorpha elevatasp. n.,Orthomorpha spiniformissp. n. and Orthomorpha subelevatasp. n., from northern Malaysia. The type-species Orthomorpha beaumontii (Le Guillou, 1841) is redescribed in due detail from male material as well, actually being a senior subjective synonym of Orthomorpha spinala (Attems, 1932), syn. n. Two additional new synonymies are proposed: Orthomorpha rotundicollis (Attems, 1937) = Orthomorpha tuberculata (Attems, 1937), syn. n., and Orthomorpha butteli Carl, 1922 = Orthomorpha consocius Chamberlin, 1945, syn. n., the valid names to the left. All species have been keyed and all new and some especially widespread species have been mapped. Further six species, including two revised from type material, are still to be considered dubious, mostly because their paraterga appear to be too narrow to represent Orthomorpha species. A new genus, Orthomorphoidesgen. n., diagnosed versus Orthomorpha through only moderately well developed paraterga, coupled with a poorly bi- or trifid gonopod tip, with at least some of its apical prongs being short spines, is erected for two species: Orthomorpha setosus (Attems, 1937), the type-species, which is also revised from type material, and Orthomorpha exaratus (Attems, 1953), both comb. n. ex Orthomorpha.     

Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera,Heptageniidae)

Based on historic collections and new material from Sumatra and Java, the species Rhithrogeniella ornata Ulmer, 1939, type species of the genus Rhithrogeniella, is reinvestigated. The nymph is described for the first time and is closely related to the continental Southeast Asian species Rhithrogeniella tonkinensis Soldán and Braasch, 1986. Rhithrogeniella belongs to the subfamily Ecdyonurinae, and is related to the genera Nixe Flowers, 1980 and/or Paracinygmula Bajkova, 1975 based on characters of the nymphal stage. Species described from Taiwan in the genus Nixe are transferred to the genus Rhithrogeniella: Rh. littoralis (Kang and Yang, 1994) comb. n., Rh. mitifica (Kang and Yang, 1994) comb. n. and Rh. obscura (Kang and Yang, 1994) comb. n.     

Revision of the endemic Taiwanese millipede genus Aponedyopus Verhoeff, 1939, with descriptions of two new species (Diplopoda, Polydesmida, Paradoxosomatidae). Advances in the systematica of Diplopoda III

The millipede genus Aponedyopus is endemic to Taiwan and contains three species. All previously described nominal species are considered to represent one species: Aponedyopus montanus Verhoeff, 1939 (the type species), including Aponedyopus reesi (Wang, 1957) and Aponedyopus maculatus Takakuwa, 1942, syn. n. Two further species are described as new: Aponedyopus similissp. n. and Aponedyopus latilobatussp. n. The genus is re-diagnosed, all of its three species are keyed, and their distributions mapped.     

A revision of the Oriental stonefly genus Phanoperla (Plecoptera: Perlidae)

The genus Phanoperla (=Dyaperla Banks, 1939) (Plecoptera: Perlidae: Perlini) is revised and generic diagnoses are provided for adults and larvae. Diagnostic and constitutive characters of the tribe Neoperlini are discussed, and Chinoperla Zwick, 1980, is shown to be the closest relative of Phanoperla. Many past misidentifications of Phanoperla species have been corrected by the use of characters recently recognized as important, namely the structure of the internal genitalia of male and female specimens and particularly the complex pattern of spines on the male penial sac made visible by eversion of this structure, and details of sculpturing of the egg chorion.
The following species of Phanoperla are recognized: amorpha sp.n.; anomala (Banks, 1939); bakeri (Banks, 1924); ceylonica Kawai, 1975; comuta sp.n.; flaveola (Klapálek, 1921), comb.n. ( =clarissa (Banks, 1913), syn.n.), (=Neoperla hageni Banks, 1920, syn.n.), (= N.consimilis Banks, 1924, syn.n.); guttata sp.n.; himalayana Zwick, 1977 (= siwalika Harper, 1977); limosa (Hagen, 1858); maculata sp.n.; maindroni (Navas, 1926), comb.n.; malayana sp.n.; minutissima (Enderlein, 1909); nana sp.n.; nervosa Banks, 1939; nuwara Kawai, 1975; omega sp.n.; pallipennis (Banks, 1938); parva sp.n.; pumilio (Klapálek, 1921), comb.n.; peniculus Kawai, 1969a; schmidi sp.n.; sertispina Jewett, 1975; srilanka sp.n.; sumatrae sp.n.; testacea (Hagen, 1858); wedda sp.n. P.claggi (Banks, 1938) is a nomen nudum.
All species are (re-)described and figured; all primary types have been examined. A key to species is provided. Most species can be assigned to one of seven species groups which are defined.
Phanoperla is endemic to the Oriental Region. Species groups are generally widespread, but individual species are in most cases known only from restricted areas.     

Stenotaenia Koch, 1847: a hitherto unrecognized lineage of western Palaearctic centipedes with unusual diversity in body size and segment number (Chilopoda: Geophilidae)

Based on morphological evidence, we newly define the genus Stenotaenia Koch, 1847 (=Scnipaeus Bergsøe & Meinert, 1866; =Simophilus Silvestri, 1896; =Onychopodogaster Verhoeff, 1902; =Insigniporus Attems, 1903; =Notadenophilus Verhoeff, 1928; =Bithyniphilus Verhoeff, 1941; =Schizopleres Folkmanova, 1956; =Euronesogeophilus Matic, 1972; all syn. nov. ) as including the following 15 species: Stenotaenia linearis (Koch, 1835) (=Geophilus simplex Gervais, 1835; =Geophilus brevicornis Koch, 1837; =Scnipaeus foveolatus Bergsøe & Meinert, 1866; =Himantarium caldarium Meinert, 1886 syn. nov. ; =Geophilus (Geophilus) linearis var. polyporus Verhoeff, 1896 syn. nov. ; =Geophilus ormanyensis Attems, 1903 syn. nov. , after lectotype designation; =Insigniporus acuneli C?pu?e, 1968 syn. nov. ) from central and northern Europe; Stenotaenia frenum (Meinert, 1870) from northern Africa; Stenotaenia romana (Silvestri, 1895) (=Geophilus silvestrii Verhoeff, 1928 syn. nov. ) and Stenotaenia sorrentina (Attems, 1903) (=Geophilus forficularius Fanzago, 1881 syn. nov. ; =Geophilus linearis abbreviatus Verhoeff, 1925 syn. nov. ) from the Italian peninsula and Sardinia; Stenotaenia antecribellata (Verhoeff, 1898) (=Simophilus albanensis Attems, 1929 syn. nov. ), Stenotaenia cribelliger (Verhoeff, 1898), Stenotaenia palpiger (Attems, 1903), Stenotaenia rhodopensis (Kaczmarek, 1970), and Stenotaenia sturanyi (Attems, 1903) from the Balkan peninsula; Stenotaenia naxia (Verhoeff, 1901) (=Geophilus graecus Verhoeff, 1902) from the Aegean islands; Stenotaenia asiaeminoris (Verhoeff, 1898) and Stenotaenia bosporana (Verhoeff, 1941) from Anatolia; Stenotaenia giljarovi (Folkmanova, 1956) from western Caucasus; Stenotaenia fimbriata (Verhoeff, 1934) and Stenotaenia palaestina (Verhoeff, 1925) from Palestine; with the only exception of S. linearis, all of these binomens are comb. nov. In Stenotaenia, a strongly conserved overall morphology is matched by an unusual interspecific variation in both the body size of fully grown specimens (from 1.7 cm in S. romana to 7.7 cm in S. sturanyi) and the number of leg‐bearing segments (from 43 in male S. romana to 115 in female S. sturanyi). The number of segments correlates with maximum body size. © 2008 The Linnean Society of London, Zoological Journal of the Linnean Society, 2008, 153 , 253–286.     

The taxonomic status of Architipula fragmentosa (Bode) and the family Eoasilidae (Diptera) from the Lower Jurassic

The fossil species Architipula fragmentosa (Bode) comb. nov. (Diptera: Nematocera: Limoniidae) from the German Upper Lias (Lower Jurassic) is redescribed. This species was originally described by Bode (1953) in a new monotypic genus as Eoasilidea fragmentosa and placed in the Brachycera, with Eoasilidea as the type-genus of a new family Eoasilidae. This family is here synonymized with the Limoniidae (Nematocera), and the genus Eoasilidea is synonymized with Architipula Handlirsch .     

New species of the tenebrionid genus <Emphasis Type="Italic">Menimus</Emphasis> sharp, 1876 (Coleoptera,Tenebrionidae) from southern Palaearctic

Three new species of the genus Menimus are described from Yunnan Province, China. Over 70 species of Menimus are distributed in the tropical and subtropical zones from India to Oceania and New Zealand. A new subgenus, Sinomenimus subgen. n., is erected with the type species M. kabaki sp. n. characterized by the absence of eyes and by the presence of the spermatheca and sharp depression at the posterior margin of the thoracic metasternite. The following new synonymy is established: Menimus Sharp, 1876 = Neomenimus Kaszab, 1939, syn. n., with the following new combinations resulting: Menimus clavatus (Kaszab, 1939) = Neomenimus clavatus Kaszab, 1939, comb, n.; Menimus biroi (Kaszab, 1939) = Neomenimus biroi Kaszab, 1939, comb, n.; Menimus setosus (Kaszab, 1939) = Neomenimus setosus Kaszab, 1939, comb, n.; Menimus brevissimus (Kaszab, 1939) = Neomenimus brevissimus Kaszab, 1939, comb. n.     

The family Ctenobelbidae (Acari,Oribatida), with description of a new species and discussion on systematic placement and taxonomic status of the genus Berndamerus Mahunka, 1977

The oribatid mite genus Berndamerus Mahunka, 1977 is transferred into the family Ctenobelbidae as the subgenus Ctenobelba (Berndamerus) Mahunka, 1977, stat. n. from the family Amerobelbidae. The known species of Berndamerus combined: C. (B.) bicostata (Berlese, 1910), comb. n., C. (B.) eremuloides (Berlese, 1910), comb. n., C. (B.) hellenica (Mahunka, 1977), comb. n. A new species, Ctenobelba (Berndamerus) bugiamapensis sp. n., is described from soil, Bu Gia Map National Park, southern Vietnam. It differs from the other species of the subgenus by the heterotrichy of notogastral setae, presence of adanal neotrichy and localization of adanal lyrifissures. Ctenobelbidae is recorded in Vietnam for the first time. A new diagnosis of the family Ctenobelbidae and the identification keys to the known subgenera of the genus Ctenobelba and species of the subgenus Ctenobelba (Berndamerus) are provided.     

The earliest Mesopsychidae and revision of the family Mesopanorpodidae (Mecoptera)

The family Mesopanorpodidae is revised. Most taxa referred to this family are not related to the type genus Mesopanorpodes Tillyard, 1918. The type species of the latter, Mesopanorpodes wianamattensis,is closely related to Mesopsyche Tillyard, 1917. Therefore Mesopanorpodes is transferred to Mesopsychidae Tillyard, 1917 (= Mesopanorpodidae Tillyard, 1918, syn. n.).The earliest Mesopsychidae are described from the Upper Permian of European Russia (Severodvinian; Isady locality, Vologda Province): Permopsyche issadensisgen. etsp. n. (type species) and Permopsyche rasnitsynisp. n. Two species described under Mesopanorpodes from the Upper Permian of Australia are also included into Permopsyche: Permopsyche belmontensis (Riek, 1953), comb. n., Permopsyche robustus (Riek, 1953) comb. n. The first pre-Triassic Mesopsyche, Mesopsyche incompletasp. n. is described from the uppermost Permian (the town of Vyazniki, Vladimir Province). Bittacopanorpa javorskii Zalessky, 1935 from the uppermost Permian or basal Triassic of Kuznetsk Basin is identified as a hindwing of Mesopsyche: Mesopsyche javorskii (Zalessky, 1935) comb. n. The origin, evolutionary history, and stratigraphic occurrence of Mesopsychidae are discussed.     

Taxonomic revision of the genus Phryganogryllacris (Orthoptera: Gryllacrididae) from China

A taxonomic revision of the genus Phryganogryllacris Karny, 1937 is provided. Twelve species of the genus from China are recorded, including 5 species described as new to science, namely P. sichuanensis Li, Liu Li, sp. nov., P. parva Li, Liu Li, sp. nov., P. truncata Li, Liu Li, sp. nov., P. interrupta Li, Liu Li, sp. nov. and P. longicerca Li, Liu Li, sp. nov. Two new combinations are also reported here, P. gialai(Gorochov, 2004)(Vietnam) comb. nov. and P. alia(Gorochov, 2004)(Vietnam) comb. nov. A key to Chinese species of the genus was provided.     

A review of the catfish genus Pterocryptis (Siluridae) in Vietnam, with the description of two new species

Of the silurid catfish genus Pterocryptis in Vietnam, a total of four species are recognized as valid, of which Pterocryptis crenula and P. verecunda are described here as new. The type locality of P. cochinchinensis is restricted to central Vietnam, and the species is redescribed from topotypic material. Pterocryptis crenula (from northeastern Vietnam) can be differentiated from its congeners in having visibly confluent anal and caudal fins while P. verecunda (from Cat Ba Island in northeastern Vietnam) can be differentiated from its congeners in having the genital papilla concealed behind the anus. Notes on the identity of other nominal Pterocryptis species are also provided.     

The genus Alphitobius Stephens (Coleoptera,Tenebrionidae, Alphitobiini) in Africa and adjacent islands

All species of the genus Alphitobius Stephens, 1829 (Alphitobiini Reitter, 1917, subfamily Tenebrioninae Latreille, 1802) from Africa and adjacent islands are revised. New species: Alphitobius capitaneus sp. n. from Kenya. New synonyms: Cryptops ulomoides Solier, 1851, syn. n. of Alphitobius diaperinus (Panzer, 1796); Alphitobius rufus Ardoin, 1976, syn. n. of Alphitobius hobohmi Koch, 1953); Peltoides (Micropeltoides) crypticoides Pic, 1916, syn. n. of Peltoides (Micropeltoides) opacus (Gerstaecker, 1871), comb. n. Homonym: Alphitobius ulomoides Koch, 1953 = Alphitobius arnoldi nom. n. New combinations from Alphitobius: Ulomoides basilewskyi (Ardoin, 1969), comb. n.; Peltoides (Micropeltoides) opacus (Gerstaecker, 1871), comb. n. Figures of all examined species are added and a species key is compiled.     

The millipede genus Orthomorpha Bollman, 1893 in Laos (Diplopoda,Polydesmida, Paradoxosomatidae), with descriptions of new species

The genus Orthomorpha is currently represented in Laos by nine species, including three, O. paviei Brölemann, 1896, O. communis Likhitrakarn, Golovatch & Panha, 2011 and O. cambodjana (Attems, 1953), which are new to the fauna of the country, and further three new to science: O. suberectoides sp. n., O. gladiata sp. n. and O. sutchariti sp. n.     

Two new genera of Nanophyidae with six desmomeres (Coleoptera, Curculionoidea)

A new genus Lyalia is described in Nanophyidae and three species are included in it: Lyalia curvatasp. n. (Vietnam), Lyalia robusta (Pic, 1921), comb. n. (from Nanophyes) (Java, Bali, Laos) and Lyalia albolineata (Pajni & Bhateja, 1982), comb. n. (from Ctenomerus) (India: Assam). Ctenomerus lagerstroemiae G. A. K. Marshall, 1923 is a syn. n. of Lyalia robusta. Thus, the genus Ctenomerus Schoenherr, 1843 is restricted to the Afrotropical Realm. Kantohiagen. n. is erected for Kantohia taiwana (Kantoh & Kojima, 2009) (from Shiva) (Taiwan). A key to the Nanophyinae genera with six desmomeres is presented.     

New or little-known species of the genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) from Vietnam

Twelve new species of the genus Amphimenes Bates, 1873 are described from Vietnam: Amphimenes maculatussp. n., Amphimenes bidoupensissp. n., Amphimenes gracilissp. n., Amphimenes montanussp. n., Amphimenes giganteussp. n., Amphimenes mediussp. n., Amphimenes minutussp. n., Amphimenes rufipessp. n., Amphimenes reflexicollissp. n., Amphimenes planicollissp. n., Amphimenes nitidussp. n., and Amphimenes kabakovisp. n.. Amphimenes rugulipennis Bates, 1892, comb. n., is transferred from the genus Brachichila Chaudoir, 1869 and is redescribed from fresh material. A key to all congeners is provided, these being arranged into five new species-groups. Adults of the rugulipennis- and piceolus-groups show subcortical habits, while those of the medius-, rufipes- and planicollis- groupsare herpetobiotic, resulting in such morphological adaptations as partly reduced eyes, missing wings and adnate elytra.     

A new species of genus Pseudaspidapion Wanat, 1990 (Coleoptera, Apionidae) from China

Pseudaspidapion botanicumsp. n. from China is described and figured. Its host plant is Grewia biloba G. Don var. parviflora (Bunge) Hand.-Mazz (Malvaceae: Grewioideae). The genus Harpapion Voss, 1966 is recorded as new for China and Vietnam and two comb. n. are proposed: Harpapion vietnamense (Korotyaev, 1985) (from Aspidapion) and Harpapion coelebs (Korotyaev, 1987) (from Pseudaspidapion). A key to the known species of the genus Pseudaspidapion from China is presented.