Historical and contemporary evidence of a mutton snapper (<Emphasis Type="Italic">Lutjanus analis</Emphasis> Cuvier, 1828) spawning aggregation fishery in decline

Scientific information on reef fish spawning aggregation fisheries is sparse in light of numerous regional declines and extirpations from overexploitation. Fisher interviews of the small-scale commercial mutton snapper (Lutjanus analis) spawning aggregation fishery at Gladden Spit, Belize, suggests a historic decadal decline. The reported trend is supported by analysis of inter-seasonal catch and effort and yield (2000–2002) that reveals a 59% decline in catch per unit effort (CPUE) and a 22% decrease in mean landings per boat. Declining population-level trends are also supported by a significant decrease in inter-annual median lengths of mutton snappers (2000–2006). These findings demonstrate the need for additional life history information that includes length-associated age and details on growth to provide clearer support of the effects on, and responses by, populations following fishing. In view of the historical changes to mutton snapper CPUE and landings at Gladden Spit and the fishery-associated declines in fish spawning aggregations observed globally, a precautionary approach to spawning aggregation management is warranted that provides full protection from fishing to enhance population persistence. The findings also highlight the need for substantially greater enforcement and long-term fisheries monitoring under a comprehensive regional management strategy.     

The use of optimal foraging theory in the understanding of fishing strategies: A case from Sepetiba Bay (Rio de Janeiro State,Brazil)

The community of Gamboa is located on Itacuruçá Island, Sepetiba Bay (State of Rio de Janeiro, Brazil) and includes 26 families, mostly of artisanal fishermen who use paddled or motor canoes, and encircling nets for fishing. In this study, predictions from optimal foraging theory (patch model), in particular of patch residence time, are compared to the observed behavior of fishermen on fishing trips. Fishermen's strategies differ depending on their intended prey. They spend more time in patches and use fewer patches for shrimp than for fish. Gamboa's fishermen tend to leave a patch later than predicted by the model. The difficulty in evaluating stock availability, the comparatively few patches available for shrimp, and the competitive aspects of fishing contribute to the explanation of this behavior.     

Utilization of seagrass habitats by juvenile groupers and snappers in Banten Bay,Banten Province,Indonesia

Coastal development in Banten Bay, Indonesia, decreased seagrass coverage to only 1.5% of its surface area. We investigated the importance of seagrass as habitat for juvenile groupers (Serranidae) and snappers (Lutjanidae), by performing beam trawl hauls on a weekly basis in two seagrass locations and one mudflat area, and monthly trawl hauls in three different microhabitats (dense, mixed and patchy seagrass) in one of the seagrass locations. We studied the effects of location and microhabitat, as well as temporal patterns (diel, weekly and monthly) on the probability of occurrence and abundance of the most abundant grouper (Orange-spotted grouper, Epinephelus coioides) and snapper (Russell’s snapper, Lutjanus russellii). We found that both species were almost exclusively found in seagrass locations, with a preference for microhabitats of high complexity (dense and mixed microhabitats). L. russellii had a higher probability of catch and abundance during the night, most probably because of its ability to avoid the beam trawl during daytime sampling. In addition there was an effect of week and month on the presence and abundance of both species, but patterns were unclear, probably because of high fishing pressure on juvenile groupers and snappers by push net fishermen. Groupers and snappers mainly fed on abundant shrimps, and to a lesser extent on fish. Moreover, juveniles find protection against predators in seagrass, which confirmed the critical role of quantity and quality of seagrass areas for juvenile groupers and snappers in Banten Bay.     

Consensus of the 'Malasars' traditional aboriginal knowledge of medicinal plants in the Velliangiri holy hills, India

Dusky grouper (garoupa, Epinephelus marginatus) is an important catch for several artisanal small-scale fisheries along the Brazilian coast. It is a sedentary, monandric, and late maturing protogynous species, which makes it vulnerable to overharvesting even though it is mainly caught through hook and line or spear fishing through free diving. Lack of information on the ecology and biology of this species in Brazil is astonishing. Much of the information found in the literature concerns Mediterranean dusky groupers. Studies compiling local knowledge (ethnoecology) about fish species complement biological data, and have been fundamental for effective fisheries management. In this study, our objectives are to obtain data about dusky grouper through fish catches and analysis of stomach contents and gonad maturation (macroscopic analyses), along with interviews from fishermen from six small-scales communities from the southern (Pântano do Sul, Florianópolis, Santa Catarina State) to the northern Brazilian coast (Porto Sauípe, Bahia State). We conclude that precautionary approaches and 'data-less' management approaches are needed in the coast of Brazil. Research on this species and on the potential of aquaculture for its cultivation, are urgent, due to the apparent vulnerability and decrease of dusky grouper along the coast of Brazil.     

Genetic and morphometric evidence that the jacks (Carangidae) fished off the coast of Rio de Janeiro (Brazil) comprise four different species

Individuals of jacks that were captured off the Rio de Janeiro coast were characterized using multivariate morphometrics and allozymes molecular markers. The main objectives were to determine the number of fish captured and their species. Classical taxonomic classification and multivariate morphometry indicated the existence of three species in the landings from Rio de Janeiro State (Caranx crysos, Carangoides bartholomaei and Caranx latus). The results of the genetic analyses performed were consistent with these results. However, Bayesian analysis revealed the possible presence of a fourth species, which could not be discriminated by classical taxonomy. Caranx crysos possibly comprises two different species. Other population indexes, such as unique alleles, degree of structuring and genetic identity, also pointed to different species. Therefore, based on evidences from classical taxonomy, geometric morphometric analyses together with genetic data from allozymes molecular markers it is possible to support that at least four different Carangidae species are present in fish landings from Niterói, Rio de Janeiro State, Brazil, whereas fish capture records for the same location have recorded only two species. These results are important for not only developing fisheries management strategies but also providing general information regarding the ecology and evolution of the genus.     

An interdisciplinary evaluation of fishery production systems off the state of Pará in North Brazil

The performance of 20 fishery production systems off the state of Pará in the northern region of Brazil was compared using the 'RAPFISH' methodology, with 57 identified attributes distributed among five evaluation fields: economics, sociology, ecology, technology and politics. The results indicated the existence of three large groups of fishery sectors: (i) industrial (red snapper with traps, the Laulao catfish, shrimp trawl) and semi-industrial (lobster) fisheries; (ii) large-scale artisanal fisheries (acoupa weakfish, red snapper with lines, king mackerel, Spanish mackerel, coco sea catfish); and (iii) small-scale artisanal fisheries (shellfish, crab, estuarine longline, fish traps, etc.). While the industrial and large-scale artisanal systems demonstrated greater sustainability from an economic and social standpoint, small-scale fisheries appeared to be more ecologically sustainable. Based on the results, a reduction in industrial fishing efforts is recommended, along with the establishment of licensing quotas for fishing vessels, as well as an increased investment in research on proper guidance and management of the semi-industrial and large-scale artisanal fisheries sectors. For small-scale artisanal fisheries, economic incentives are suggested for the aggregate value of the products and to assist fishers in the development of an appropriate social organization. Finally, it is believed that a greater stakeholder involvement in the decision-making process would improve management actions for all modalities.     

When Does this Fish Spawn? Fishermen’s Local Knowledge of Migration and Reproduction of Brazilian Coastal Fishes

Fishermen’s local knowledge of fishing resources may be an important source of information to improve artisanal tropical fisheries management, such as those found in Brazil, where most data on fish biology is lacking. We aim to study the local ecological knowledge that Brazilian coastal fishers have about reproductive aspects (season, places and migration) of 13 coastal fish species of commercial importance. We selected fishermen with more than 30 years of fishing practice and we interviewed a total of 67 fishermen: 29 from the southeastern coast, from the communities of Puruba, Almada, Picinguaba and Bertioga, and 38 from the northeastern coast, from the communities of Valença, Arembepe and Porto Sauípe. In the interviews, we used standardized questionnaires and showed photos of fish species. Our results indicate some general patterns in fishes’ reproduction according to fishermen knowledge: fish species spawn in open ocean, near reefs or in coastal rivers (estuaries); some fishes reproduce during the summer and others in winter, while some have more defined spawning months. The main fish migratory patterns mentioned by interviewees were: long migrations along the coast, usually in the South to North direction, short migrations among reefs, fishes that do not migrate, migrations between the shore and open ocean and migrations between the sea and coastal rivers. Fishermen’s knowledge differed among fish species: most fishermen did not know spawning places or seasons of large pelagic fishes, which raised concerns of their possible depletion. We compared such ethnoichthyological information with available scientific data, indicating promising insights about reproduction and migration of Brazilian coastal fishes. Data gathered from local fishermen may provide inexpensive and prompt information, potentially applicable to fisheries management. Our approach might be useful to several other small-scale fisheries, especially the tropical ones, where there is a high diversity of target species and a low biological and ecological knowledge about these species.     

The paraty artisanal fishery (southeastern Brazilian coast): ethnoecology and management of a social-ecological system (SES)

ABSTRACT: This study intends to give recommendations to the management of Paraty fishery in Brazil through an interplay of local and scientific knowledge. In particular, the objectives are the following: 1) to describe the Paraty fishery; 2) to compare the fishermen's local ecological knowledge with recorded fish landings and previous studies in Paraty; 3) to combine the data on local fishing and on local/Caicara livelihoods with the SES (social-ecological systems) Model. The methods include a systematic survey of fishing in Tarituba and Praia Grande, which are located in the northern end and the central part of the Paraty municipality, respectively. For four days each month, systematic data on catches at landing points were collected, as well as macroscopic gonad analysis data for the fishes Centropomus parallelus and C. undecimalis (snook, robalo), Epinephelus marginatus (grouper, garoupa), Scomberomorus cavalla (King mackerel, cavala), and Lutjanus synagris (Lane snapper, vermelho). Spring and summer are important seasons during which some species reproduce, and the integration of fishing periods for some target species could assist in fishing management through the use of closed seasons. Fishermen could obtain complementary earnings from tourism and from the "defeso system" (closed season including a salary payment) to conserve fishing stocks. The SES model facilitates an understanding of the historical context of fishing, its economic importance for local livelihoods, the constraints from conservation measures that affect fishermen, and the management processes that already exist, such as the defeso. If used to integrate fishing with complementary activities (tourism), such a system could improve the responsibility of fishermen regarding the conservation of fish stocks.     

Local Ecological Knowledge and Scientific Data Reveal Overexploitation by Multigear Artisanal Fisheries in the Southwestern Atlantic

In the last decades, a number of studies based on historical records revealed the diversity loss in the oceans and human-induced changes to marine ecosystems. These studies have improved our understanding of the human impacts in the oceans. They also drew attention to the shifting baseline syndrome and the importance of assessing appropriate sources of data in order to build the most reliable environmental baseline. Here we amassed information from artisanal fishermen''s local ecological knowledge, fisheries landing data and underwater visual census to assess the decline of fish species in Southeastern Brazil. Interviews with 214 fishermen from line, beach seine and spearfishing revealed a sharp decline in abundance of the bluefish Pomatomus saltatrix, the groupers Epinephelus marginatus, Mycteroperca acutirostris, M. bonaci and M. microlepis, and large parrotfishes in the past six decades. Fisheries landing data from a 16-year period support the decline of bluefish as pointed by fishermen''s local knowledge, while underwater visual census campaigns show reductions in groupers'' abundance and a sharp population decline of the Brazilian endemic parrotfish Scarus trispinosus. Despite the marked decline of these fisheries, younger and less experienced fishermen recognized fewer species as overexploited and fishing sites as depleted than older and more experienced fishermen, indicating the occurrence of the shifting baseline syndrome. Here we show both the decline of multigear fisheries catches – combining anecdotal and scientific data – as well as changes in environmental perceptions over generations of fishermen. Managing ocean resources requires looking into the past, and into traditional knowledge, bringing historical baselines to the present and improving public awareness.     

Shared Stocks of Snappers (Lutjanidae) in Australia and Indonesia: Integrating Biology, Population Dynamics and Socio-Economics to Examine Management Scenarios

Joint Australia–Indonesia scientific workshops on the fisheries of the Arafura Sea, held in 1992 and 1994, concluded that the two countries might share stocks of the red snappers Lutjanus malabaricus and L. erythropterus and the gold-band snapper Pristipomoides multidens. At that time, no information concerning stock structure, distribution and movements of these species was available. Moreover, data on the population biology and on commercial catches were inadequate. Such data are crucial for stock assessment and for managing the stocks. Clearly, if the stocks being fished were shared, joint management would be appropriate. In order to answer the questions related to managing shared stocks, a collaborative research project was initiated by Australia (CSIRO as the lead agency) and Indonesia in 1999. The objectives were firstly, to describe the population dynamics, stock structure and biology of snappers relevant to the management of stocks shared between Australian and Indonesian fisheries; secondly, to characterize the social and financial structures of the Indonesian fishery so they could be taken into account in the development of management strategies; and thirdly, to explore ways of developing complementary management for the long term sustainability of the snapper fisheries. This project finished in 2003 and in this paper we bring together the results of the biological, genetic, population dynamics and socioeconomic research in relation to managing shared stocks in the context of managed versus unmanaged fisheries, small scale and industrial fisheries, and in both developed and developing country regulatory environments. Severe data limitations necessitated an innovative approach making use of comparative analyses, often data-poor values, and the drawing together of fishery dependent and independent data to evaluate the status of the stocks.     

Folk taxonomy and scientific nomenclature: Working together for conservation of fishery resources in Brazil

Folk taxonomies of fishers are important for understanding the dynamics and diversity of fishes throughout the world. This study analyzed the folk taxonomy of artisanal fishers of the coast of the state of Paraíba, Brazil. Data were collected through free lists, semi-structured interviews and participant observations. A total of 308 local names were recorded, corresponding to 127 scientific taxa. The broad folk categories ‘pescada’ (croakers) and ‘cação’ (shark) were the most common. The Cognitive Salience Index (CSI) was used to assess which fishes the fishers knew best; the mutton snapper (Lutjanus analis) had the highest CSI score. Four hierarchical levels of folk taxonomy (kingdom, life form, generic and specific) were recorded. The local ichthyofauna contained 16 species at risk at the national level and 34 at the global level. Our results suggest that fisher folk taxonomy can complement scientific knowledge, improve and update threatened species lists and in doing so benefit fisheries management and conservation actions.     

Spatial and Dietary Overlap Creates Potential for Competition between Red Snapper (Lutjanus campechanus) and Vermilion snapper (Rhomboplites aurorubens)

Understanding the complex nature of direct and indirect species interactions is a critical precursor to successful resource management. In the northern Gulf of Mexico fisheries ecosystem, red snapper (Lutjanus campechanus) and vermilion snapper (Rhomboplites aurorubens) are two commercially harvested species within a larger reef fish complex. These two species share similar habitats and diets; however, little is known about how these species partition habitat and dietary resources. In this study we examined the extent of spatial and dietary overlap between red snapper and vermilion snapper, and experimentally compared their feeding behavior. Field data from multiple gear types demonstrates that red snapper and vermilion snapper frequently cohabited reefs in the northern Gulf of Mexico, and Pianka’s niche overlap indices suggest significantly overlapping diets. Experimental manipulations show that red snapper are the dominant forager of the two species, as red snapper foraging alone ate more shrimp per fish than vermilion snapper in both the single species (p = 0.003) and mixed species (p = 0.02) treatments. In addition, red snapper ate significantly more shrimp per fish in the mixed species treatment than in the single species treatment (p = 0.04). Vermilion snapper shrimp consumption per fish did not differ significantly between mixed and single species treatments. Cumulatively, our results suggest that spatial and dietary overlap could lead to competition between red and vermilion snapper in the study area; however, conclusively determining the existence of such competition would require further research.     

Partially protected areas as a management tool on inshore reefs

Partially Protected Areas (PPAs) are a widely-used management tool, yet comparatively little is known about their effectiveness compared to more commonly studied No-Take Marine Reserves (NTMRs). Here, we examine the efficacy of two kinds of PPAs (with and without spearfishing) within the Great Barrier Reef Marine Park (GBRMP) that are subject to a range of fishing limitations, and assess their utility as a marine park zoning and fisheries management tool. Fish abundance, size, and habitat composition were compared inside PPAs and NTMRs on inshore reefs of the central GBR. Fish abundances were lower inside PPAs relative to adjacent NTMRs for primary fishing targets, with no detectable effects for secondary targets and non-targets, or for species richness. Fish assemblages differed amongst zones, but these variations were minor compared to regional variations in species composition. Partially Protected Areas supported 46%–69% of the relative abundance of total primary targets compared to adjacent NTMRs, with no evident increase in abundance in zones where spearfishing was prohibited. There were no reductions in the size of two key target species: coral trout (Plectropomus spp.) and stripey snapper (Lutjanus carponotatus) inside PPAs, and only stripey snapper had significant reductions in abundance inside PPAs compared to NTMRS. Habitat and biophysical characteristics (especially topographic complexity) were strong drivers of fish abundance, but the relative influence of zone was greater for target species compared to non-targets. This study provides novel data on PPAs and highlights their utility as a spatial management tool in contributing to conservation and fisheries management goals.

     

Isolation and characterization of microsatellites in lane snapper (Lutjanus synagris), mutton snapper (Lutjanus analis), and yellowtail snapper (Ocyurus chrysurus)

Polymerase chain reaction primer pairs for a total of 25 nuclear‐encoded microsatellites (loci) were developed from genomic DNA libraries of lane snapper (Lutjanus synagris), mutton snapper (Lutjanus analis), and yellowtail snapper (Ocyurus chrysurus). The microsatellites include 24 perfect (21 dinucleotide and three trinucleotide) and one imperfect (combination tetranucleotide/tetranucleotide) repeat motifs. A total of 32 individuals of each species were assayed for allelic variation at all 25 microsatellites; reliable amplification products were generated for lane snapper (25 loci), mutton snapper (21 loci), and yellowtail snapper (24 loci). Significant deviations from Hardy–Weinberg expectations, following Bonferroni corrections, were found for one microsatellite in lane and yellowtail snappers, and for three microsatellites in mutton snapper. All pairwise comparisons of microsatellites (all three species) did not deviate significantly from genotypic equilibrium.     

Fishing at Rio Grande (Brazil): Ecological niche and competition

This study is an ecological analysis of the use of resources by two potentially competitive groups, the commercial and recreational fishermen of Rio Grande, located on the border of the States of SÃo Paulo and Minas Gerais. Tourism at Rio Grande was intensified after the construction of the hydroeletric plant of Marimbondo in 1974, and local fishermen have complained about its interference. Values of niche dimensions, such as fishing periods, grounds, and prey caught were obtained through data gathered at landing points. Local and recreational fishermen usually fish in different parts of the river and catch different fish, because they use different gear. However, in transition months between the wet and dry seasons, when fish are less abundant, there is conflict or competition related to fishing grounds, reinforced by the territorial fishing rights of local longline fishermen in this period. We show how ecological concepts, such as niche and competition, may help to understand strategies of exploitation by human groups, fundamental information for the management of tropical areas.     

Species Distribution Models of Tropical Deep-Sea Snappers

Deep-sea fisheries provide an important source of protein to Pacific Island countries and territories that are highly dependent on fish for food security. However, spatial management of these deep-sea habitats is hindered by insufficient data. We developed species distribution models using spatially limited presence data for the main harvested species in the Western Central Pacific Ocean. We used bathymetric and water temperature data to develop presence-only species distribution models for the commercially exploited deep-sea snappers Etelis Cuvier 1828, Pristipomoides Valenciennes 1830, and Aphareus Cuvier 1830. We evaluated the performance of four different algorithms (CTA, GLM, MARS, and MAXENT) within the BIOMOD framework to obtain an ensemble of predicted distributions. We projected these predictions across the Western Central Pacific Ocean to produce maps of potential deep-sea snapper distributions in 32 countries and territories. Depth was consistently the best predictor of presence for all species groups across all models. Bathymetric slope was consistently the poorest predictor. Temperature at depth was a good predictor of presence for GLM only. Model precision was highest for MAXENT and CTA. There were strong regional patterns in predicted distribution of suitable habitat, with the largest areas of suitable habitat (> 35% of the Exclusive Economic Zone) predicted in seven South Pacific countries and territories (Fiji, Matthew & Hunter, Nauru, New Caledonia, Tonga, Vanuatu and Wallis & Futuna). Predicted habitat also varied among species, with the proportion of predicted habitat highest for Aphareus and lowest for Etelis. Despite data paucity, the relationship between deep-sea snapper presence and their environments was sufficiently strong to predict their distribution across a large area of the Pacific Ocean. Our results therefore provide a strong baseline for designing monitoring programs that balance resource exploitation and conservation planning, and for predicting future distributions of deep-sea snappers.     

Spatial and temporal variation in artisanal catches of dolphinfish Coryphaena hippurus off north‐eastern Brazil

The sampling of fish from the artisanal fleet operating with surface lines off north‐eastern Brazil was carried out between 1998 and 2000. Generalized linear models (GLMs) were used to standardize mean abundance indices using catch and fishing effort data on dolphinfish Coryphaena hippurus and to identify abundance trends in time and space, using 1215 surface line deployments. A standard relative abundance index (catch per unit effort, CPUE) was estimated for the most frequent vessels used in the sets, employing factors and coefficients generated in the GLMs. According to the models, C. hippurus catches are affected by the operating characteristics and power of different fishing vessels. These differences highlight the need for standardization of catch and effort data for artisanal fisheries. The highest mean abundance values for C. hippurus were off the state of Rio Grande do Norte, with an increasing tendency in areas with greater depths and more distant from the coast, reaching maximal values in areas whose depths range from 200 to 500 m. The highest mean abundance values occurred between April and June. The higher estimated abundance of C. hippurus in this period off the state of Rio Grande do Norte and within the 200–500 m depth range may be related to a migration pattern of food sources, as its main prey, the flying fish Hirundichthys affinis, uses floating algae as refuge and to deposit its pelagic eggs.     

Isolation and characterization of microsatellite loci from mangrove red snapper Lutjanus argentimaculatus

Lutjanus argentimaculatus, also called mangrove red snapper, is a commercially important fish in East Asia. A proper understanding of population structure is primarily linked with the management of genetic resources in exploiting marine fisheries. Herein, seven microsatellite loci, which showed high polymorphism (observed heterozygosity per locus ranging from 0.3571 to 0.7857 and expected heterozygosity per locus ranging from 0.6236 to 0.8821), were isolated and characterized from L. argentimaculatus. Cross‐species amplifications also indicate that primers designed for these loci may be useful for further studies about other closely phylogenetic species of the family Lutjanidae.     

Fishing spots and sea tenure: Incipient forms of local management in Atlantic forest coastal communities

Recent work has dealt with the local management of aquatic resources as an alternative to Hardin's (1968) tragedy of the commons. In communities with no formal management of resources, informal ownership of fishing spots or conflicts with outside competitors may determine the basis for future local management. In this study, I analyze the use of aquatic resources by five fishing communities on the Atlantic Forest coast of southeast Brazil: Búzios Island, Puruba, and Picinguaba in São Paulo State, and Jaguanum and Itacuruçá Islands at Sepetiba Bay in Rio de Janeiro State. Informal ownership of fishing spots, used for set gillnet fishing, is regulated by kin ties at Búzios Island. The artisanal fishers of Sepetiba Bay, especially those from Jaguanum Island, have a conflict with Bay intruders, such as the shrimp and herring trawlers. Two coastal communities, Puruba and Picinguaba, have conflicts with fishing regulations from a State Park (Parque Estadual da Serra do Mar),created in 1977. The transformation of populated areas of the Atlantic Forest to Extractive Reserves might be a way to avoid conflicts with intruders and with governmental agencies, and to involve local populations in management. Kinship rules at Búzios Island and the territorial behavior of fishers at Sepetiba Bay may form a basis for local organization.     

Reproductive cycle of female yellow snapper Lutjanus argentiventris (Pisces, Actinopterygii, Lutjanidae) in the SW Gulf of California: gonadic stages, spawning seasonality and length at sexual maturity

The yellow snapper ( Lutjanus argentiventris ) in the SW Gulf of California is being heavily fished and little information exists on status of this exploited population. From August 2001 to May 2003, 440 specimens in the size range of 10.8–59 cm were collected from four fishing areas: Huizache-Caimanero, Mazatlan, Marmol and Santa María la Reforma in the SW Gulf of California. Specimens were collected from fishermen and obtained by sampling with gill nets, hook and line and spear gun fishing. Gonadal stages, spawning season and length-at-first maturity were estimated for female yellow snapper through histological analyses and relative gonadosomatic (GSI) and hepatosomatic (HSI) indexes. Six gonadal stages were identified and gonadal development was asynchronous. Gonads in advanced stage VI of vitellogenesis occurred during summer and winter together with the highest values of the GSI and HSI, indicating that spawning occurred during these seasons. Mean maturity length was determined to be 32.6 cm total length. The yellow snapper population was being adversely affected considering that the species was exploited throughout the year, including the reproductive seasons, and that organisms as small as 10 cm were found in fishermen's catches and at fish markets. Minimum capture sizes of 33 cm and a closed season during the reproductive periods of summer and winter are suggested as measures to reduce the adverse affects of this exploitation.