土壤养分空间异质性与植物根系的觅食反应

植物在长期进化过程中,为了最大限度地获取土壤资源,对养分的空间异质性产生各种可塑性反应．包括形态可塑性、生理可塑性、菌根可塑性等．许多植物种的根系在养分丰富的斑块中大量增生,增生程度种间差异较大,并受斑块属性(斑块大小、养分浓度)、营养元素种类和养分总体供应状况的影响．植物还通过调整富养斑块中细根的直径、分枝角、节问距以及空间构型来实现斑块养分的高效利用．根系的生理可塑性及菌根可塑性可能在一定程度上影响其形态可塑性．生理可塑性表现为处于不同养分斑块上的根系迅速调整其养分吸收速率,从而增加单位根系的养分吸收,对在时间上和空间上变化频繁的空间异质性土壤养分的利用具有重要意义,可在一定程度上弥补根系增生反应的不足．菌根可塑性目前研究较少,一些植物种的菌根代替细根实现在富养斑块中的增生．菌根增生的碳投入养分吸收效率较高、根系增生对增加养分吸收的作用较复杂,取决于养分离子在土壤中的移动性能以及是否存在竞争植物;对植物生长(竞争能力)的作用因种而异,一些敏感种由此获得生长效益,而其它一些植物种受影响较小．植物个体对土壤养分空间异质性反应能力和生长差异,影响其在群落中的地位和命运,最终影响群落组成及其结构．     

Spatial heterogeneity in soil nutrient supply modulates nutrient and biomass responses to multiple global change drivers in model grassland communities

Changes in the atmospheric concentration of carbon dioxide ([CO₂]), nutrient availability and biotic diversity are three major drivers of the ongoing global change impacting terrestrial ecosystems worldwide. While it is well established that soil nutrient heterogeneity exerts a strong influence on the development of plant individuals and communities, it is virtually unknown how nutrient heterogeneity and global change drivers interact to affect plant performance and ecosystem functioning. We conducted a microcosm experiment to evaluate the effect of simultaneous changes in [CO₂], nutrient heterogeneity (NH), nutrient availability (NA) and species evenness on the biomass and nutrient uptake patterns of assemblages formed by Lolium perenne, Plantago lanceolata and Holcus lanatus. When the nutrients were heterogeneously supplied, assemblages exhibited precise root foraging patterns, and had higher above‐ and belowground biomass (average increases of 32% and 29% for above‐ and belowground biomass, respectively). Nutrient heterogeneity also modulated the effects of NA on biomass production, complementarity in nitrogen uptake and below: aboveground ratio, as well as those of [CO₂] on the nutrient use efficiency at the assemblage level. Our results show that nutrient heterogeneity has the potential to influence the response of plant assemblages to simultaneous changes in [CO₂], nutrient availability and biotic diversity, and suggest that it is an important environmental factor to interpret and assess plant assemblage responses to global change.     

额济纳绿洲土壤养分的空间异质性

利用地统计学方法研究了额济纳绿洲土壤有机质、全氮、全磷、全钾的空间异质性.结果表明:土壤养分含量分布均有较大的空间异质性,土壤有机质的理论模型为指数模型,全氮、全磷和全钾的理论模型均为球状模型;额济纳绿洲土壤有机质、全氮、全钾主要受结构性因子的影响,全磷受结构性因子和随机因子的共同影响;全氮、全磷和全钾的变程在7.68～7.86 km,土壤有机质的变程较小,为5.4 km.土壤养分的空间分布表现为南北高,中间低的分布格局.     

土壤保肥-供肥机理及其调节Ⅱ.棕壤型菜园土的腐殖质结合形态及其肥力学意义

采自辽宁省不同地区的１５对棕壤型菜园土肥、瘦地及其各粒级微团聚体的有机质储量和腐殖质结合形态的研究表明,肥地及其各粒级微团聚体的有机质和各结合形态腐殖质的含量与松结态腐殖质占有机质总量的比例均比瘦地及其各相应粒级微团聚体的高,稳结态腐殖质的比例较小,紧结态腐殖质的比例肥瘦地大体相当．无论肥地或瘦地,小粒级微团聚体的有机质和各结合形态腐殖质的含量及松、稳结态腐殖质占有机质总量的比例均较大粒级的高,而紧结态腐殖质则相反,表明大、小粒级微团聚体具有不同的肥力学意义     

绿洲-荒漠过渡带景观的植被与土壤特征要素的空间异质性分析

采用变异函数的理论和方法,通过空间异质性特征比较,研究了阜康绿洲-荒漠过渡带景观中的植被盖度和土壤环境因子的空间异质性、空间格局及其关系．在此基础上,应用Kriging局部块段插值法,以三维图形显示出各个要素的空间分布格局．结果表明,除土壤全盐量空间相关性较弱外,其他各要素具有中等程度以上的空间相关性,且空间变异主要发生在较小尺度上．其中,植被盖度的空间异质性程度较高,在3km的尺度内具有明显的空间格局．从空间分布格局看,植被盖度、土壤表层含水量和土壤pH值沿样带相邻点差异对比明显,高值区和低值区交错．草本盖度与土壤表层含水量、土壤pH间有相关关系,灌木平均盖度主要依靠深层次地下水维系．灌木和草本盖度的块金值具有明显差异,可能因为二者要求不同的生态条件,因而在同等尺度上具有不同的生态学过程．     

Spatial and temporal heterogeneity of superficial muscle strain during in situ fixed-end contractions

Numerical models of contracting muscle offer a powerful tool to study local mechanical load. For validation of these models, the spatial and temporal distribution of strain was quantified in fixed-end contracting rat tibialis anterior muscle in situ at optimal muscle length (L(o)) and at 120 degrees plantar flexion as well as at 125 and 33Hz stimulation frequency. We studied the hypothesis that after termination of stimulation in situ muscle segments near the motor endplates elongate while segments away from the endplates shorten. We show that both spatial and temporal inhomogeneities in muscle deformation occurred during contraction. Muscle plateau shortening strain equalled 4.1%. Maximal plateau shortening of a muscle segment was much larger (9.6%) and occurred distally (at 0.26 of the scaled length of the muscle). Manipulating torque levels by decreasing the stimulation frequency at the same muscle length induced a decrease in torque ( approximately 20%) with a smaller effect on the level and no effect on the pattern of muscle deformation. During relaxation, distal segments actively shortened at the expense of proximal muscle segments, which elongated. The segments undergoing lengthening were nearer to motor endplates than segments undergoing shortening.In conclusion, the present study provides experimental data on magnitude of contraction-induced deformation needed for validation of numerical models. Local muscle deformation is heterogeneous both temporally and spatially and may be related to proximity to the motor endplates.     

Effects of wind erosion on the spatial heterogeneity of soil nutrients in two desert grassland communities

Wind is known to affect the spatial heterogeneity of soil resources in arid and semiarid systems, but multi-year, quantified observations are largely absent. We studied the effects of wind erosion on the spatial distribution of soil organic carbon (SOC) and other soil nutrients at the Jornada Experimental Range, in southern New Mexico. Enhanced wind erosion was encouraged by grass cover reduction in a Sporobolus-mesquite dominated site (SM) and a Bouteloua-mesquite dominated site (BM). The scale and magnitude of spatial dependence for the soil analytes were quantified using geostatistical analyses. Results of this study show that soil organic matter related analytes such as SOC, TN, N_avail, and SO₄ ^2- are among the first to be eroded and redistributed; cations such as Ca²⁺ and Mg²⁺ may not be removed and redistributed significantly; and other ions such as K⁺, Na⁺ and Cl⁻ showed no discernible pattern of change. Geostatistics show that wind appeared to increase the scale of spatial autocorrelation, but decrease the scale of spatial dependence of most soil analytes over 2–3 windy seasons. In the wind enhanced plot of the SM site, up to 99% of the spatial dependence of SOC was autocorrelated at the distance of 1.45 m before the initiation of wind erosion, but the spatial dependence dropped significantly to only 60% at a larger autocorrelation distance of 2.76 m after three windy seasons. Similar but less significant changes were observed for SOC in the BM site. Despite the differential effects of wind on the soil analytes, we conclude that the overall results of wind on the grass cover reduction plots are the disappearance of small, strong fertile islands, which may be related to grasses; and the reinforcement of large fertile islands, which are likely related to mesquite shrubs. In addition, the change of the spatial patterns of SOC and other soil nutrients induced by enhanced wind erosion may persist and reinforce soil islands associated with shrubs, thus allowing a positive feedback for further desertification in this arid grassland.     

土壤保肥-供肥机理及其调节 Ⅰ.棕壤型菜园土的P素保持与供应

对辽宁省北部、中部及南部的棕壤型菜园土15对肥、瘦地的保P-供P性能的测定表明,在绝大多数供试土样中,肥地及其各粒级微团聚体的全P量、P解吸量及P解吸率均大于瘦地及其各相应粒级微团聚体的,P吸附量则相反。不论肥地或瘦地,<10μm微团聚体均比>10μm微团聚体具有更大的P吸附能力。供试菜园土的<10μm微团聚体亦具有较大的供P潜势,其P解吸率比其他农用土壤相同粒级微团聚体的大1个数量级。     

Prairie vegetation and soil nutrient responses to ungulate carcasses

The impact of large ungulate carcasses on grassland dynamics was investigated by monitoring vegetation and soil nutrients in 50-cm circular zones around the center of bison (Bos bison), cattle (B. taurus), and deer (Odocoileus virginianus) carcasses. An ungulate carcass creates an intense localized disturbance that varies with animal size and the season of death. Unlike other natural disturbances, carcasses deposit a concentrated pulse of nutrients into the soil. One year after death, inorganic nitrogen concentrations were significantly higher in the inner 50 cm at both adult and juvenile carcass sites than in surrounding prairie. Areas around a carcass became zones of fertility that favored different components of the vegetation and stimulated biomass production. Species richness and diversity at the center of carcass sites were lowest 1 year after death, but increased significantly in subsequent years. However, warm-season perennial grasses declined near the center of carcass sites and did not recover. Five years after death, ungulate carcass sites remained disturbed patches that harbored vegetation characteristically different in composition and stature from surrounding prairie. By providing a niche for species not normally found in undisturbed prairie, carcasses increased community heterogeneity and may play an important role in adding spatial complexity to grassland ecosytems. Received: 12 April 1999 / Accepted: 7 September 1999     

沙坡头人工固沙植被土壤水分空间异质性

利用传统统计学和地统计学相结合的方法对沙坡头人工固沙植被区0～200 cm 之间的各层土壤水分的空间变异性进行研究.结果表明,1）土壤水分相对变异较大的层是在160～180 cm层和180～200 cm层,变异系数分别为0.72和0.73.表层0～5 cm层的相对变异也较大,变异系数为0.662）半方差函数分析结果表明,各层土壤水分均具有明显的空间变异性,各层土壤水分自相关部分的空间异质性占总空间异质性的程度很高,所占比例在87.7%～99.9%范围内.各层土壤水分的有效变程大小有较大差异,最小值出现在60～80 cm层(7.04 m),最大值出现在20～40 cm层(19.71 m),土壤水分有效变程从表层到深层没有明显的变化规律;3）土壤水分插值图反映出0～140 cm之间相邻各层土壤水分变化较大,140～200 cm各层土壤水分变化较小;4）土壤水分在0°、45°、90°、135°四个方向的半方差函数基本上是一条直线,表明在这四个方向上半方差和距离的相关性较低,土壤水分变化是独立、随机的,是同质性的.     

黄土高原土壤水分变化的时空特征分析

对晋西北砖窑沟流域荒地7年土壤水分定位观测表明,生长期土壤水分动态主要受降水量及其分配的影响．一般年份生长期土壤水分动态可分为3个时段：1)春末夏初土壤水分消耗期;2)夏末秋初土壤水分蓄积期和3)秋末冬初土壤水分消耗期．0-300cm土层水分含量的季节变化与测定序号的关系方程为：1)直线型(1988,1990),2)指数型(1992,1995)和3)抛物线型(1993)．以150cm为界,下部土壤水分与上部的比大于1．不同降水年土壤湿度与土层深度的关系方程均为三次抛物线型．土壤水分的变异系数随深度的增加而减小．0-300cm内的土壤水分剖面可分为活跃层(200cm以上)、过渡层(200-250cm)和稳定层(250cm以下)．各层深度随降水量及其分配的不同而变化．不同降水年荒地土壤水分的方差分析总体差异显著,但各组间的均值配对比较结果不同．特涝年的荒地土壤水分与其它年份差异均显著．荒地与柠条林地的土壤水分差异显著,但与河北杨林地和小叶杨林地的土壤水分差异不显著．本文采用的定量研究方法有助于黄土高原不同地区和不同植被条件下土壤水分的定量比较．     

土壤保肥—供肥机理及其调节Ⅰ.棕壤型菜园土的P素保持与供应

对辽宁省北部、中部及南部的棕壤型菜园土１５对肥、瘦地的保Ｐ供Ｐ性能的测定表明,在绝大多数供试土样中,肥地及其各粒级微团聚体的全Ｐ量、Ｐ解吸量及Ｐ解吸率均大于瘦地及其各相应粒级微团聚体的,Ｐ吸附量则相反．不论肥地或瘦地,＜１０μｍ微团聚体均比＞１０μｍ微团聚体具有更大的Ｐ吸附能力．供试菜园土的＜１０μｍ微团聚体亦具有较大的供Ｐ潜势,其Ｐ解吸率比其他农用土壤相同粒级微团聚体的大１个数量级．     

Impact of mineral fertilizers on mineral nutrients in the ginger rhizome and on soil enzymes activities and soil properties

Ginger is used as one of the important ingredients in traditional as well as modern medicine besides as a spice. It boosts immunity and is a rich source of many biologically active substances and minerals. Although it is a medicinally important crop, its productivity is, however, affected due to poor nutrient management and therefore it requires an adequate supply of nutrients in the form of inorganic fertilizers or organic manuring, or a mixture of both. In this context, the present study was aimed to investigate the effect of mineral fertilizers on the content of mineral elements in the ginger rhizome, on soil enzyme activity, and soil properties. Lysimeter experiments were conducted at the Institute of Genetics and Plant Experimental Biology, Kibray, Tashkent region, Uzbekistan. The experiment comprised of four treatments T1 – Control, T2 - N₇₅P₅₀K₅₀ kg/ha, T3 - and T4 - N₁₀₀P₇₅K₇₅ + B₃Zn₆Fe₆ kg/ha. The results showed that the application of N₁₂₅P₁₀₀K₁₀₀ kg/ha increased rhizome K content by 49%, P content by 20%, and Na content by 58% as compared to control without fertilizer. While the application of N₁₀₀P₇₅K₇₅ + B₃Zn₆Fe₆ kg/ha showed a significant enhancement in rhizome K, Ca, P, Mg, Na, Fe, Mn, Zn, Cu, Cr, Mo, and Si contents over the control. This treatment also improved active P content by 29%, total P content by 80%, total K content 16%, and N content by 33% content, and the activities of urease, invertase, and catalase activities as compared to control of without mineral fertilizer and control respectively. Thus the application of NPK + BZnFe at the rate of 100:75:75:3:6:6 kg/ha helps in improving macroelements and microelements in the ginger rhizome and activities of soil enzymes that helps in mineral nutrition of the rhizome.     

Adaptations and biomass production of two grasses in response to waterlogging and soil nutrient enrichment

We analysed the response of two grass species, Danthonia montevidensis and Paspalum dilatatum to waterlogging, soil-nutrient enrichment and the combination of both factors. Waterlogging did not affect total biomass of D. montevidensis, but it slightly promoted growth of P. dilatatum. Most analysed variables showed no significant interaction between fertilization and waterlogging. Therefore, waterlogging does not produce a detrimental effect either in the growth of these species or in their response capacity to stimulating growth factors, such as fertilization.     

黄土半干旱区坡地土壤水分、养分及生产力空间变异

通过田问取样,分析黄土半干旱丘陵区陡坡面土壤水分、养分及其生产力空间变异性及其相关关系．结果表明,坡面0～20cm土层土壤各养分含量均不同程度地高于20～40cm土层,但变异程度却明显低于20～40cm土层,且除全磷外,土壤养分变异程度均明显高于水分．20～40cm土层土壤养分沿坡面向下逐渐增大,而0～20cm沿坡面纵向差异较小．坡面浅沟沟槽处的土壤水分、养分条件最优,但其地上生物量却低于坡顶．虽然纵向坡度(35°～45°)明显大于横向(5°～10°),但横向坡位对土壤养分的影响却明显大于纵向(除20～40cm土层速效磷含量除外),而对水分的影响纵向大于横向．相关分析表明,0～120cm土壤水分与20～40cm土壤养分以及土壤养分之间(0～20cm速效磷除外)均呈极显著相关．坡位对土壤水分、养分及地上生物量有很大影响,但土壤水分、养分对生物量影响不显著．     

作物种植前后土壤有机质及养分因子的空间变异分析

以作物土壤试验数据为基础,结合多年的农户调查数据,采用Moran's I指数分析、Pearson相关系数分析和Kriging插值3种地统计学方法,探讨了浙江省浦江县作物种植前后土壤有机质及氮磷钾等养分因子的空间演化规律。研究发现Moran's I指数分析证实随着区域范围的加大,各因子间自相关程度产生较大差异。这一点在Pearson相关系数分析中也得到证实,并在kriging插值图中直观表达。结果还表明:有机质和碱解氮的形成与水的关系较为密切,水源区两因子的含量较周边高,且会促进前者向后者转化。养分投入后,过多的养分在土壤中降解过程与区域自然环境条件的关系较密切。同一作物的农户养分投入在不同区域差异较大,不同作物的农户养分投入在区域内却存在量的相关。     

新疆农田作物覆膜地温极值的时空变化

土壤温度的上限和下限值极大地影响农田作物的生长发育．根据对新疆农田2年度不同观测时刻、不同覆膜条件及不同作物的最高和最低地温的分析,可知2年度全生育期地温极值均发生在0cm位置．地温极值依观测时刻的不同而变动．覆膜最低温度始终高于裸地,说明覆膜对提高低温段土壤温度有明显作用．最低温度的日变化可用二次函数表达;14：00和20：00时最高地温可分别用深度的椭圆曲线和线性关系表达．不同条件下的地温极值和气温具有线性关系,其中裸地最低温度与气温的相关性高于覆膜;玉米最低温度与气温的相关性高于棉花;最高温度与气温的相关性比最低温度与气温的相关性差．     

异龙湖湖滨带不同环境梯度下土壤养分空间变异性

随着环境梯度的变化,生态系统的特征会发生明显的变异,土壤是其重要的表征要素。以纵向岭谷区红河流域异龙湖湖滨带为案例,运用多种统计分析方法对其不同环境梯度下土壤单个养分指标和土壤养分综合质量的空间分布特征进行了研究,结果表明：在北部湖滨坡地,海拔对土壤养分空间分布的影响并不十分显著,而坡位对其影响较大;北部湖滨农田土壤,水分梯度对钾的影响较大,全氮和有机质则一部分受水分梯度影响,另一部分受径流作用的影响;异龙湖湖滨带土壤养分综合质量指数大小的排序为：南部湖演农田〉北部湖滨农田〉北部湖滨坡地）南部湖滨坡地。还探讨了不同环境因子对单个土壤养分指标和土壤养分综合质量的影响,其中,气温、湿度、坡向和坡型对土壤中磷空间分布的影响较大,土壤含水量、坡度和海拔对土壤有机质、全氮和全钾的影响较大;土壤含水量即水分梯度对土壤养分综合质量影响最大。     

贵州麻若平台泥炭藓沼泽中泥炭藓持水特性及其与土壤营养元素关系研究

泥炭藓属（Sphagnum）植物的持水特性在泥炭湿地形成过程中起着重要作用,研究其与土壤营养元素之间的关系有助于地区湿地保护和退化湿地的修复。以贵州麻若平台分布的泥炭沼泽为对象,采用方差分析、相关性分析和冗余分析等方法对沼泽内泥炭藓的生物量、蓄水量、吸水率和土壤营养元素含量等进行研究。结果显示,该区域泥炭藓物种组成主要包括狭叶泥炭藓（Sphagnum cuspidatum Ehrh.）、多纹泥炭藓（Sphagnum multifibrosum X.J.Li&M.Zang）和卵叶泥炭藓（Sphagnum ovatum Hampe.）3种,狭叶泥炭藓为优势种,占泥炭藓总盖度的87.2%。沼泽中泥炭藓的生物量为（0.62 ±0.01）kg/m²,自然蓄水量为（9.42 ±0.45）kg/m²,饱和吸水率达1827.41%±34.56%,说明泥炭藓具有很强的持水能力。RDA分析表明,泥炭藓的生物量、饱和吸水量和饱和吸水率主要受沼泽土壤总钾、总磷、有效磷、有效氮、有效钾的影响,泥炭藓的鲜重、自然蓄水量和自然吸水率受土壤pH和水位的影响较大。泥炭藓的生物量、饱和蓄水量、饱和吸水率与土壤总钾含量呈正相关,与土壤总磷、有效磷、总钾、有效氮、总氮、有机质含量呈负相关,说明土壤总磷、有效磷、有效钾、有效氮、总氮、有机质对沼泽中泥炭藓的持水能力具有抑制作用。